treatments-xml/data/03/F0/96/03F09661FFF48065FED5E393791C519F.xml

<document ID-DOI="10.1080/00222933.2022.2115950" ID-ISSN="1464-5262" ID-Zenodo-Dep="7156474" checkinTime="1665136984522" checkinUser="plazi" docAuthor="Medeiros, Rone A. F., Seidel, Matthias &amp; Grossi, Paschoal C." docDate="2022" docId="03F09661FFF48065FED5E393791C519F" docLanguage="en" docName="JNATHIST.56.29-32.1315-1364.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 56 (29 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2022.2115950" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Byrsopolis quadraticeps Blanchard 1851" docType="treatment" docVersion="3" lastPageNumber="1329" masterDocId="FFC9EE19FFFF806BFFD0E116782B521F" masterDocTitle="Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay" masterLastPageNumber="1364" masterPageNumber="1315" pageNumber="1326" updateTime="1665173136910" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title>Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Medeiros, Rone A. F.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grossi, Paschoal C.</mods:namePart>
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<mods:title>Journal of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="series">
<mods:title>J. Nat. Hist.</mods:title>
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<mods:detail type="pubDate">
<mods:number>2022-09-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>29 - 32</mods:number>
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<mods:start>1315</mods:start>
<mods:end>1364</mods:end>
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<mods:identifier type="DOI">10.1080/00222933.2022.2115950</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">7156474</mods:identifier>
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<subSubSection box="[261,1054,645,671]" pageId="11" pageNumber="1326" type="nomenclature">
<paragraph blockId="11.[261,1054,645,671]" box="[261,1054,645,671]" pageId="11" pageNumber="1326">
<heading bold="true" box="[261,1054,645,671]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="1326" reason="3">
<emphasis bold="true" box="[261,1054,645,671]" pageId="11" pageNumber="1326">
<taxonomicName authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[261,766,645,671]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis bold="true" box="[261,554,645,671]" italics="true" pageId="11" pageNumber="1326">Byrsopolis quadraticeps</emphasis>
<bibRefCitation author="Blanchard E" box="[562,766,645,671]" pageId="11" pageNumber="1326" pagination="241" refId="ref25750" refString="Blanchard E. 1851. Gide et Baudry. In: Milne-Edwards H, Blanchard E, Lucas H, editors. Catalogue des collections Entomologiques du Museum d'Histoire naturelle de Paris, Classe des Insectes. Ordre des Coleopteres. Paris, France: Museum d'histoire naturelle de Paris. (1850) 1. (1850) 1128, (1851) 129240; p. 129240; p. 241." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[775,889,645,671]" captionStart="Figure 3" captionStartId="12.[160,225,945,968]" captionTargetBox="[163,1153,146,913]" captionTargetId="figure-212@12.[162,1154,142,914]" captionTargetPageId="12" captionText="Figure 3. Byrsopolis quadraticeps Blanchard, holotype (MNHN-EC-EC1804), dorsal and lateral habitus (a–b); labels attached (c); aedeagus caudal and lateral view (d–e). Images from Antoine Mantilleri (MNHN). Scale bars: a, b = 5 mm; d, e = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7156482" httpUri="https://zenodo.org/record/7156482/files/figure.png" pageId="11" pageNumber="1326">Figures 3</figureCitation>
(a–e), 7(a–g))
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="1326" type="reference_group">
<paragraph blockId="11.[160,1156,698,931]" pageId="11" pageNumber="1326">
<treatmentCitationGroup pageId="11" pageNumber="1326">
<treatmentCitation author="Blanchard E" box="[160,656,698,723]" page="219" pageId="11" pageNumber="1326" year="1851">
<taxonomicName authority="Blanchard 1851: 219" authorityName="Blanchard" authorityPageNumber="219" authorityYear="1851" box="[160,656,698,723]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis box="[160,417,698,723]" italics="true" pageId="11" pageNumber="1326">Byrsopolis quadraticeps</emphasis>
<bibRefCitation author="Blanchard E" box="[426,656,698,723]" pageId="11" pageNumber="1326" pagination="241" refId="ref25750" refString="Blanchard E. 1851. Gide et Baudry. In: Milne-Edwards H, Blanchard E, Lucas H, editors. Catalogue des collections Entomologiques du Museum d'Histoire naturelle de Paris, Classe des Insectes. Ordre des Coleopteres. Paris, France: Museum d'histoire naturelle de Paris. (1850) 1. (1850) 1128, (1851) 129240; p. 129240; p. 241." type="journal article" year="1851">Blanchard 1851: 219</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description); 
<treatmentCitation author="Lacordaire JT" box="[914,1153,698,723]" page="364" pageId="11" pageNumber="1326" year="1856">
<bibRefCitation author="Lacordaire JT" box="[914,1153,698,723]" pageId="11" pageNumber="1326" pagination="594" refId="ref26568" refString="Lacordaire JT. 1856. Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes. Paris: Librairie Encyclopedique de Roret; p. 594." type="book chapter" year="1856">Lacordaire 1856: 364</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Castelnau FL &amp; Lucas MH" box="[160,523,732,757]" page="130" pageId="11" pageNumber="1326" year="1857">
<bibRefCitation author="Castelnau FL &amp; Lucas MH" box="[160,523,732,757]" pageId="11" pageNumber="1326" pagination="473" refId="ref26028" refString="Castelnau FL, Lucas MH. 1857. Animaux nouveaux ou rares recueilles pendant l'Expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima et de Lima au Para sous la direction du Comte Francis de Castelnau. Chez P. Bertrand. Paris: Librareie-Editeur; p. 473." type="book chapter" year="1857">Castelnau and Lucas 1857: 130</bibRefCitation>
</treatmentCitation>
(distribution and illustration); 
<treatmentCitation author="Gemminger M &amp; Harold B" page="1226" pageId="11" pageNumber="1326" year="1869">
<bibRefCitation author="Gemminger M &amp; Harold B" pageId="11" pageNumber="1326" pagination="1240" refId="ref26271" refString="Gemminger M, Harold B. 1869. Catalogus Coleopterum: Scarabaeidae, Tom. IV. 4: 1240." type="journal article" year="1869">Gemminger and Harold 1869: 1226</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation author="Machatschke JW" box="[432,679,767,792]" page="3" pageId="11" pageNumber="1326" year="1972">
<bibRefCitation author="Machatschke JW" box="[432,679,767,792]" pageId="11" pageNumber="1326" pagination="361" refId="ref26699" refString="Machatschke JW. 1972. Scarabaeoidea: Melolonthinae, Rutelinae. Coleopterorum Catalogus. Supplementa. 66: 361." type="journal article" year="1972">Machatschke 1972: 3</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation author="Krajcik M" box="[837,1024,767,792]" page="58" pageId="11" pageNumber="1326" year="2008">
<bibRefCitation author="Krajcik M" box="[837,1024,767,792]" pageId="11" pageNumber="1326" pagination="142" refId="ref26488" refString="Krajcik M. 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). 2 nd ed. Plzen, Czech Republic: Milan Krajcik; p. 142." type="book chapter" year="2008">Krajcik 2008: 58</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation author="Carvalho TG &amp; Grossi PC" box="[160,510,802,827]" page="372" pageId="11" pageNumber="1326" year="2018">
<bibRefCitation author="Carvalho TG &amp; Grossi PC" box="[160,510,802,827]" pageId="11" pageNumber="1326" pagination="369 - 372" refId="ref25906" refString="Carvalho TG, Grossi PC. 2018. Description of a second species of the genus Moronius Grossi &amp; Vaz-deMello, 2015 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). Zootaxa. 4434 (2): 369 - 372. doi: 10. 11646 / zootaxa. 4434.2.7." type="journal article" year="2018">Carvalho and Grossi 2018: 372</bibRefCitation>
</treatmentCitation>
(checklist, distribution); 
<treatmentCitation author="Menis FT &amp; Rodrigues SR" box="[792,1129,802,827]" page="3" pageId="11" pageNumber="1326" year="2021">
<bibRefCitation author="Menis FT &amp; Rodrigues SR" box="[792,1129,802,827]" pageId="11" pageNumber="1326" pagination="928" refId="ref26719" refString="Menis FT, Rodrigues SR. 2021. Occurrence of Phytophagous Scarabaeidae (Coleoptera) in a pasture area at the Balsamo municipality, Sao Paulo, Brazil. EntomoBrasilis, 14: e 928. doi: 10.12741 / ebrasi lis. v 14. e 928" type="journal article" year="2021">Menis and Rodrigues 2021: 3</bibRefCitation>
</treatmentCitation>
, 5 (citation); 
<bibRefCitation author="Vaz-de-Mello FZ &amp; Grossi PC" box="[273,607,837,862]" pageId="11" pageNumber="1326" refId="ref27519" refString="Vaz-de-Mello FZ, Grossi PC. 2022. Melolonthidae in Catalogo Taxonomico da Fauna do Brasil. PNUD. http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 127169." type="url" year="2022">Vaz-de-Mello and Grossi 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="11.[160,1156,698,931]" pageId="11" pageNumber="1326">
<treatmentCitationGroup pageId="11" pageNumber="1326">
<taxonomicName authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[192,770,871,896]" class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="species" species="quadraticeps" subGenus="Byrsopolis">
<emphasis box="[192,277,871,896]" italics="true" pageId="11" pageNumber="1326">Cotalpa</emphasis>
(
<emphasis box="[295,403,871,896]" italics="true" pageId="11" pageNumber="1326">Byrsopolis</emphasis>
) 
<emphasis box="[421,561,871,896]" italics="true" pageId="11" pageNumber="1326">quadraticeps</emphasis>
(
<bibRefCitation author="Blanchard E" box="[577,762,871,896]" pageId="11" pageNumber="1326" pagination="241" refId="ref25750" refString="Blanchard E. 1851. Gide et Baudry. In: Milne-Edwards H, Blanchard E, Lucas H, editors. Catalogue des collections Entomologiques du Museum d'Histoire naturelle de Paris, Classe des Insectes. Ordre des Coleopteres. Paris, France: Museum d'histoire naturelle de Paris. (1850) 1. (1850) 1128, (1851) 129240; p. 129240; p. 241." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
: 219; 
<treatmentCitation author="Ohaus F" box="[842,1021,871,896]" page="10" pageId="11" pageNumber="1326" year="1918">
<bibRefCitation author="Ohaus F" box="[842,1021,871,896]" pageId="11" pageNumber="1326" pagination="241" refId="ref27050" refString="Ohaus F. 1918. Coleopterorum Catalogus, pars. 66. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelianae. Vol. XX. Berlin, Germany: W. Junk; p. 241." type="book chapter" year="1918">Ohaus 1918: 10</bibRefCitation>
</treatmentCitation>
(catalogue, distribution); 
<treatmentCitation author="Ohaus F" box="[311,484,906,931]" page="38" pageId="11" pageNumber="1326" year="1934">
<bibRefCitation author="Ohaus F" box="[311,484,906,931]" pageId="11" pageNumber="1326" pagination="172" refId="ref27151" refString="Ohaus F. 1934. Genera Insectorum, Coleoptera, fam. Scarabaidae: subfam. Rutelinae, Erster Teil, fasc. 199. Tervuren, Belgique; p. 172." type="book chapter" year="1934">Ohaus 1934: 38</bibRefCitation>
</treatmentCitation>
(citation, checklist); 
<treatmentCitation author="Blackwelder RE" box="[714,964,906,931]" page="235" pageId="11" pageNumber="1326" year="1944">
<bibRefCitation author="Blackwelder RE" box="[714,964,906,931]" pageId="11" pageNumber="1326" pagination="189 - 306" refId="ref25704" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, The West Indies, and South America. Washington: Smithsonian Institution, United States National Museum. Bulletin 185, Part 2; p. 189 - 306." type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="1328" pageId="11" pageNumber="1326" type="materials_examined">
<paragraph blockId="11.[160,554,974,1000]" box="[160,554,974,1000]" pageId="11" pageNumber="1326">
<emphasis bold="true" box="[160,317,974,1000]" italics="true" pageId="11" pageNumber="1326">
<typeStatus box="[160,216,974,1000]" pageId="11" pageNumber="1326">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[332,395,975,1000]" name="Brazil" pageId="11" pageNumber="1326">Brazil</collectingCountry>
, 
<collectingRegion box="[405,551,975,1000]" country="Brazil" name="Minas Gerais" pageId="11" pageNumber="1326">Minas Gerais</collectingRegion>
.
</paragraph>
<paragraph blockId="11.[160,1156,1039,1208]" pageId="11" pageNumber="1326">
<emphasis bold="true" box="[160,334,1044,1070]" italics="true" pageId="11" pageNumber="1326">Type material.</emphasis>
<materialsCitation collectionCode="MNHN" pageId="11" pageNumber="1326" specimenCode="MNHN-EC-EC1804" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis box="[350,707,1045,1070]" italics="true" pageId="11" pageNumber="1326">
<typeStatus box="[350,447,1045,1070]" pageId="11" pageNumber="1326">Holotype</typeStatus>
<specimenCount box="[459,514,1045,1070]" pageId="11" pageNumber="1326" type="male">male</specimenCount>
, here designated
</emphasis>
. (a) ‘MUSEUM PARIS/OEST CAPIT e/DES MINES’, (b) ‘OUEST DE/LA CAPITE DES MINES [handwritten]’, (c) ‘ 
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[878,995,1079,1104]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/ 
<taxonomicName authority=", Bl." authorityName="Bl." class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="species" species="quadraticeps">quadraticeps, Bl.</taxonomicName>
[handwritten]’, (d) 
<taxonomicName box="[399,573,1114,1139]" class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="species" species="quadraticeps">B. quadraticeps</taxonomicName>
/Cat. Mus/Brésil./M. Aug. Saint-Hilaire [handwritten]’, (e) HOLOTYPE [red label]’, (f) ‘MNHN/EC1804’ (
<specimenCount box="[696,727,1149,1174]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[737,814,1149,1174]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="1326">MNHN</collectionCode>
, specimen examined through images by A. Mantilleri; 
<specimenCode box="[430,635,1183,1208]" pageId="11" pageNumber="1326">MNHN-EC-EC1804</specimenCode>
, 
<bibRefCitation author="MNHN" box="[645,782,1183,1208]" pageId="11" pageNumber="1326" refId="ref26767" refString="MNHN. 2016. Museum National d'Histoire Naturelle: the Coleoptera collection (EC) of the Museum national d'Histoire naturelle (MNHN - Paris). www. gbif. org / occurrence / 583421279." type="book" year="2016">MNHN 2016</bibRefCitation>
) (
<figureCitation box="[804,895,1183,1208]" captionStart="Figure 3" captionStartId="12.[160,225,945,968]" captionTargetBox="[163,1153,146,913]" captionTargetId="figure-212@12.[162,1154,142,914]" captionTargetPageId="12" captionText="Figure 3. Byrsopolis quadraticeps Blanchard, holotype (MNHN-EC-EC1804), dorsal and lateral habitus (a–b); labels attached (c); aedeagus caudal and lateral view (d–e). Images from Antoine Mantilleri (MNHN). Scale bars: a, b = 5 mm; d, e = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7156482" httpUri="https://zenodo.org/record/7156482/files/figure.png" pageId="11" pageNumber="1326">Figure 3</figureCitation>
(a-e)).
</materialsCitation>
</paragraph>
<paragraph blockId="11.[160,1157,1252,1728]" lastBlockId="13.[160,1156,142,237]" lastPageId="13" lastPageNumber="1328" pageId="11" pageNumber="1326">
<emphasis bold="true" box="[160,399,1252,1278]" italics="true" pageId="11" pageNumber="1326">Additional material.</emphasis>
[
<specimenCount box="[420,579,1253,1278]" pageId="11" pageNumber="1326" type="generic">59 specimens</specimenCount>
] 
<emphasis box="[593,807,1253,1278]" italics="true" pageId="11" pageNumber="1326">
<collectorName box="[593,654,1253,1278]" pageId="11" pageNumber="1326">Brazil</collectorName>
, 
<collectingRegion box="[666,807,1253,1278]" inferredBySuffix="true" pageId="11" pageNumber="1326">Minas Gerais</collectingRegion>
</emphasis>
. ‘ 
<materialsCitation collectingDate="2012-12-03" collectingMethod="Light" collectionCode="CEMT, CERPE" collectorName="C. M Oliveira'" country="Brazil" location="Farm Querencia" municipality="Buritis" pageId="11" pageNumber="1326" specimenCount="17" specimenCount-male="17" stateProvince="Minas Gerais">
<collectingCountry box="[824,907,1253,1278]" name="Brazil" pageId="11" pageNumber="1326">BRASIL</collectingCountry>
: 
<collectingRegion box="[919,1065,1253,1278]" country="Brazil" name="Minas Gerais" pageId="11" pageNumber="1326">Minas Gerais</collectingRegion>
. 
<collectingMunicipality box="[1076,1147,1253,1278]" pageId="11" pageNumber="1326">Buritis</collectingMunicipality>
/ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FF70E411797E573F" box="[160,341,1287,1312]" country="Brazil" municipality="Buritis" name="Farm Querencia" pageId="11" pageNumber="1326" stateProvince="Minas Gerais">Farm Querência</location>
. 
<date box="[352,481,1287,1312]" pageId="11" pageNumber="1326" value="2012-12-03">
<collectingDate box="[352,481,1287,1312]" pageId="11" pageNumber="1326" value="2012-12-03">03-xii-2012</collectingDate>
</date>
./ 
<collectingMethod box="[492,550,1287,1312]" pageId="11" pageNumber="1326">Light</collectingMethod>
. 
<collectorName box="[560,708,1287,1312]" pageId="11" pageNumber="1326">C. M Oliveira’</collectorName>
(
<specimenCount box="[723,757,1287,1312]" pageId="11" pageNumber="1326" type="male">9♂</specimenCount>
<collectionCode box="[765,831,1287,1312]" pageId="11" pageNumber="1326">CEMT</collectionCode>
, 
<specimenCount box="[841,875,1287,1312]" pageId="11" pageNumber="1326" type="male">8♂</specimenCount>
<collectionCode box="[882,958,1287,1312]" collectionName="CERPE" pageId="11" pageNumber="1326">CERPE</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="2003-10-15" collectionCode="CEMT" collectorName="T. BERNARDO" location="IJACI" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1" stateProvince="MG">
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FC00E4117C27573F" box="[976,1036,1287,1312]" name="IJACI" pageId="11" pageNumber="1326" stateProvince="MG">IJACI</location>
, 
<collectingRegion box="[1046,1087,1287,1312]" pageId="11" pageNumber="1326">MG</collectingRegion>
/ 
<date pageId="11" pageNumber="1326" value="2003-10-15">
<collectingDate pageId="11" pageNumber="1326" value="2003-10-15">15/X/ 03</collectingDate>
</date>
/ 
<collectorName box="[196,352,1322,1347]" pageId="11" pageNumber="1326">T. BERNARDO</collectorName>
Col’ (
<specimenCount box="[426,457,1322,1347]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[468,536,1322,1347]" pageId="11" pageNumber="1326">CEMT</collectionCode>
).
</materialsCitation>
<materialsCitation collectionCode="CMN" location="Rondonopolis" municipality="Rondonopolis" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso">
<emphasis box="[556,696,1322,1347]" italics="true" pageId="11" pageNumber="1326">
<collectingRegion box="[556,696,1322,1347]" country="Brazil" name="Mato Grosso" pageId="11" pageNumber="1326">Mato Grosso</collectingRegion>
</emphasis>
. ‘ 
<collectingMunicipality box="[715,875,1322,1347]" pageId="11" pageNumber="1326">Rondonópolis</collectingMunicipality>
/ 
<collectingRegion box="[884,1032,1322,1347]" country="Brazil" name="Mato Grosso" pageId="11" pageNumber="1326">Mato Grosso</collectingRegion>
/10.62’ (
<specimenCount box="[1124,1155,1322,1347]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[160,219,1357,1382]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="11" pageNumber="1326" type="Museum">CMN</collectionCode>
); ‘
</materialsCitation>
<materialsCitation box="[235,806,1357,1382]" collectingDate="2012-10-13" collectionCode="CEMT" collectorName="/ Ana'" location="UNIVAG" municipality="Cuiaba" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1" stateProvince="MT">
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FF3BE45B79625779" box="[235,329,1357,1382]" municipality="Cuiaba" name="UNIVAG" pageId="11" pageNumber="1326" stateProvince="MT">UNIVAG</location>
/ 
<collectingMunicipality box="[336,414,1357,1382]" pageId="11" pageNumber="1326">Cuiabá</collectingMunicipality>
– 
<collectingRegion box="[438,475,1357,1382]" pageId="11" pageNumber="1326">MT</collectingRegion>
/ 
<date box="[483,609,1357,1382]" pageId="11" pageNumber="1326" value="2012-10-13">
<collectingDate box="[483,609,1357,1382]" pageId="11" pageNumber="1326" value="2012-10-13">13/10/2012</collectingDate>
</date>
<collectorName box="[609,669,1357,1382]" pageId="11" pageNumber="1326">/Ana’</collectorName>
(
<specimenCount box="[684,715,1357,1382]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[721,789,1357,1382]" pageId="11" pageNumber="1326">CEMT</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="2004-10-23" collectionCode="CERPE" collectorName="Pereira &amp; Lucimar'" location="UNIVAG" municipality="UNIVAG" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1" stateProvince="MT">
<collectingMunicipality box="[805,901,1357,1382]" pageId="11" pageNumber="1326">UNIVAG</collectingMunicipality>
/ 
<collectingMunicipality box="[908,1014,1357,1382]" pageId="11" pageNumber="1326">V. Grande</collectingMunicipality>
<collectingRegion box="[1020,1055,1357,1382]" pageId="11" pageNumber="1326">MT</collectingRegion>
–/ 
<date pageId="11" pageNumber="1326" value="2004-10-23">
<collectingDate pageId="11" pageNumber="1326" value="2004-10-23">23/10/ 2004</collectingDate>
</date>
/ 
<collectorName box="[224,304,1391,1416]" pageId="11" pageNumber="1326">Pereira</collectorName>
, 
<collectorName box="[319,414,1391,1416]" pageId="11" pageNumber="1326">Lucimar’</collectorName>
(
<specimenCount box="[436,467,1391,1416]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[479,555,1391,1416]" collectionName="CERPE" pageId="11" pageNumber="1326">CERPE</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="15 / 10.2004" collectionCode="CEMT" collectorName="Vilma'" location="UNIVAG" municipality="Varzea Grande" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1" stateProvince="MT">
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FD90E4797AB75797" box="[576,668,1391,1416]" municipality="Varzea Grande" name="UNIVAG" pageId="11" pageNumber="1326" stateProvince="MT">UNIVAG</location>
/ 
<collectingMunicipality box="[676,850,1391,1416]" pageId="11" pageNumber="1326">Várzea Grande</collectingMunicipality>
/ 
<collectingRegion box="[857,893,1391,1416]" pageId="11" pageNumber="1326">MT</collectingRegion>
/ 
<collectingDate box="[901,1025,1391,1416]" pageId="11" pageNumber="1326">15/10.2004</collectingDate>
/ 
<collectorName box="[1034,1103,1391,1416]" pageId="11" pageNumber="1326">Vilma’</collectorName>
(
<specimenCount box="[1124,1155,1391,1416]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[160,229,1426,1451]" pageId="11" pageNumber="1326">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation box="[249,1040,1426,1451]" collectingDate="2004-10-16" collectionCode="CERPE" collectorName="Danny Rotilli'" location="V. Grande" municipality="V. Grande" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1" stateProvince="MT">
same, except ‘ 
<collectingRegion box="[416,454,1426,1451]" pageId="11" pageNumber="1326">MT</collectingRegion>
– 
<collectingMunicipality box="[491,610,1426,1451]" pageId="11" pageNumber="1326">V. Grande</collectingMunicipality>
/ 
<date box="[618,726,1426,1451]" pageId="11" pageNumber="1326" value="2004-10-16">
<collectingDate box="[618,726,1426,1451]" pageId="11" pageNumber="1326" value="2004-10-16">16.X.2004</collectingDate>
</date>
/ 
<collectorName box="[734,888,1426,1451]" pageId="11" pageNumber="1326">Danny Rotilli’</collectorName>
(
<specimenCount box="[911,942,1426,1451]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[954,1031,1426,1451]" collectionName="CERPE" pageId="11" pageNumber="1326">CERPE</collectionCode>
).
</materialsCitation>
<materialsCitation collectingDate="1935-11-10" collectionCode="ZMHB" collectorName="R. Spitz S. &amp; Ohaus" location="L. Bulhoes" pageId="11" pageNumber="1326" specimenCount="2" specimenCount-male="2" stateProvince="Goias">
<emphasis box="[1050,1111,1426,1451]" italics="true" pageId="11" pageNumber="1326">
<collectingRegion box="[1050,1111,1426,1451]" country="Brazil" name="Goias" pageId="11" pageNumber="1326">Goiás</collectingRegion>
</emphasis>
. ‘C. GOYAZ/ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FF2FE4A3795F57D1" box="[255,372,1461,1486]" name="L. Bulhoes" pageId="11" pageNumber="1326" stateProvince="Goias">L. Bulhões</location>
/ 
<collectorName box="[380,498,1461,1486]" pageId="11" pageNumber="1326">R. Spitz S.</collectorName>
\ 
<date box="[505,589,1461,1486]" pageId="11" pageNumber="1326" value="1935-11-10">
<collectingDate box="[505,589,1461,1486]" pageId="11" pageNumber="1326" value="1935-11-10">10.xi.35</collectingDate>
</date>
/a 
<specimenCount box="[623,643,1461,1484]" pageId="11" pageNumber="1326" type="male">♂</specimenCount>
’ // ‘ 
<collectorName box="[693,768,1461,1486]" pageId="11" pageNumber="1326">Ohaus</collectorName>
determ’ (
<specimenCount box="[888,919,1461,1486]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[931,1004,1461,1486]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="11" pageNumber="1326">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="1937-12" collectionCode="MNRJ" collectorName="Dr. Nick'" location="LEOPOLDO BULHOES" municipality="Est. Goyaz" pageId="11" pageNumber="1326" specimenCount="2" specimenCount-male="2">
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FBD2E4A3793857EF" municipality="Est. Goyaz" name="LEOPOLDO BULHOES" pageId="11" pageNumber="1326">LEOPOLDO BULHOES</location>
/ 
<collectingMunicipality box="[283,402,1495,1520]" pageId="11" pageNumber="1326">Est. Goyaz</collectingMunicipality>
<date box="[416,509,1495,1520]" pageId="11" pageNumber="1326" value="1937-12">
<collectingDate box="[416,509,1495,1520]" pageId="11" pageNumber="1326" value="1937-12">Dez. 37</collectingDate>
</date>
/ 
<collectorName box="[518,617,1495,1520]" pageId="11" pageNumber="1326">Dr. Nick’</collectorName>
(
<specimenCount box="[642,673,1495,1520]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[687,757,1495,1520]" collectionName="Brazil, Rio de Janeiro, Sao Cristovao, Universidade do Rio Janeiro, Museu Nacional" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="1326">MNRJ</collectionCode>
); same (
<specimenCount box="[863,894,1495,1520]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[908,978,1495,1520]" collectionName="Brazil, Rio de Janeiro, Sao Cristovao, Universidade do Rio Janeiro, Museu Nacional" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="1326">MNRJ</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="2016-11-03" collectingDateMax="2016-11-08" collectingDateMin="2016-11-03" collectionCode="CERPE, MZUSP" collectorName="Luz, N. C. M. &amp; Souza &amp; Dr. Nick'" country="Brazil" location="Fazenda Larga Grande" municipality="Cristalina" pageId="11" pageNumber="1326" specimenCount="4" specimenCount-male="4" stateProvince="Goias">
<collectingCountry box="[1003,1070,1495,1520]" name="Brazil" pageId="11" pageNumber="1326">Brasil</collectingCountry>
, 
<collectingRegion box="[1086,1152,1495,1520]" country="Brazil" name="Goias" pageId="11" pageNumber="1326">Goiás</collectingRegion>
, 
<collectingMunicipality box="[160,265,1530,1555]" pageId="11" pageNumber="1326">Cristalina</collectingMunicipality>
,/ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FEC4E4EC7A0B540C" box="[276,544,1530,1555]" country="Brazil" county="Brasil" municipality="Cristalina" name="Fazenda Larga Grande" pageId="11" pageNumber="1326" stateProvince="Goias">Fazenda Larga Grande</location>
/(
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FDFEE4EC7ABA540C" box="[558,657,1530,1555]" country="Brazil" county="Brasil" municipality="Cristalina" name="Sorgatto" pageId="11" pageNumber="1326" stateProvince="Goias">Sorgatto</location>
), 
<date box="[677,832,1530,1555]" pageId="11" pageNumber="1326" value="2016-11-03" valueMax="2016-11-08" valueMin="2016-11-03">
<collectingDate box="[677,832,1530,1555]" pageId="11" pageNumber="1326" value="2016-11-03" valueMax="2016-11-08" valueMin="2016-11-03">03–08.xi.2016</collectingDate>
</date>
,/ 
<collectorName box="[844,973,1530,1555]" pageId="11" pageNumber="1326">Luz, N.C.M.</collectorName>
<collectorName box="[984,1051,1530,1555]" pageId="11" pageNumber="1326">Souza</collectorName>
leg’ (
<specimenCount box="[1120,1155,1530,1555]" pageId="11" pageNumber="1326" type="male">2♂</specimenCount>
<collectionCode box="[160,236,1565,1590]" collectionName="CERPE" pageId="11" pageNumber="1326">CERPE</collectionCode>
); ‘ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FED2E70B79D15429" box="[258,506,1565,1590]" country="Brazil" municipality="Cristalina" name="LEOPOLDO / BULHOES" pageId="11" pageNumber="1326" stateProvince="Goias">LEOPOLDO/BULHOES</location>
/Est de 
<collectingRegion box="[596,660,1565,1590]" country="Brazil" name="Goias" pageId="11" pageNumber="1326">Goias</collectingRegion>
/(Brasil)/DIRINGS\XII. 
<collectorName box="[899,996,1565,1590]" pageId="11" pageNumber="1326">Dr. Nick’</collectorName>
(
<specimenCount box="[1015,1049,1565,1590]" pageId="11" pageNumber="1326" type="male">2♂</specimenCount>
<collectionCode box="[1061,1147,1565,1590]" collectionName="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="1326">MZUSP</collectionCode>
); ‘
</materialsCitation>
<materialsCitation box="[163,886,1599,1624]" collectionCode="IBSP" collectorName="J. Guerin'" county="Leopoldo Bulhoes" location="Goiaz" municipality="Goiaz" pageId="11" pageNumber="1326" specimenCount="1" specimenCount-male="1">
<collectingCounty box="[163,373,1599,1624]" pageId="11" pageNumber="1326">Leopoldo Bulhões</collectingCounty>
/ 
<collectingMunicipality box="[381,446,1599,1624]" pageId="11" pageNumber="1326">Goiaz</collectingMunicipality>
/12.937’ // ‘Coll. 
<collectorName box="[638,746,1599,1624]" pageId="11" pageNumber="1326">J. Guerin’</collectorName>
(
<specimenCount box="[767,798,1599,1624]" pageId="11" pageNumber="1326" type="male">1♂</specimenCount>
<collectionCode box="[810,863,1599,1624]" collectionName="Brazil,Sao Paulo, Sao Paulo, Instituto Butantan" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="11" pageNumber="1326">IBSP</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="1989-11" collectionCode="MSPC" location="Cristalina" municipality="Brazil" pageId="11" pageNumber="1326" specimenCode="WSD00347498, WSD00347499" specimenCount="2" specimenCount-male="2" stateProvince="Goias">
<collectingMunicipality box="[885,950,1599,1624]" pageId="11" pageNumber="1326">Brazil</collectingMunicipality>
, 
<collectingRegion box="[965,1029,1599,1624]" country="Brazil" name="Goias" pageId="11" pageNumber="1326">Goias</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF48060FBC3E7297C505447" box="[1043,1147,1599,1624]" country="Brazil" municipality="Brazil" name="Cristalina" pageId="11" pageNumber="1326" stateProvince="Goias">Cristalina</location>
/ 
<date box="[160,239,1634,1659]" pageId="11" pageNumber="1326" value="1989-11">
<collectingDate box="[160,239,1634,1659]" pageId="11" pageNumber="1326" value="1989-11">xi.1989</collectingDate>
</date>
/unknown collector’ // ‘Coll. Matthias/Seidel 2016’ // ‘ 
<taxonomicName authorityName="Blanchard" authorityYear="1851" box="[870,1147,1634,1659]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1326" phylum="Arthropoda" rank="species" species="quadraticeps">Byrsopolis quadraticeps</taxonomicName>
/ 
<bibRefCitation author="Blanchard E" box="[160,348,1669,1694]" pageId="11" pageNumber="1326" pagination="241" refId="ref25750" refString="Blanchard E. 1851. Gide et Baudry. In: Milne-Edwards H, Blanchard E, Lucas H, editors. Catalogue des collections Entomologiques du Museum d'Histoire naturelle de Paris, Classe des Insectes. Ordre des Coleopteres. Paris, France: Museum d'histoire naturelle de Paris. (1850) 1. (1850) 1128, (1851) 129240; p. 129240; p. 241." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
/det. M. Seidel 2017’ // ‘WORLD/SCARAB./DATABASE/[
<specimenCode box="[985,1150,1669,1694]" pageId="11" pageNumber="1326">WSD00347498</specimenCode>
- 
<specimenCode box="[160,329,1703,1728]" pageId="11" pageNumber="1326">WSD00347499</specimenCode>
]’ (
<specimenCount box="[354,389,1703,1728]" pageId="11" pageNumber="1326" type="male">2♂</specimenCount>
<collectionCode box="[400,469,1703,1728]" collectionName="Italy, Varallo, Museo di Storia Naturale &quot;Pietro Calderini&quot;" country="Italy" httpUri="http://grbio.org/cool/t9g4-v7fn" name="Museo di Storia Naturale &amp;quot;Pietro Calderini&amp;quot;" pageId="11" pageNumber="1326">MSPC</collectionCode>
);
</materialsCitation>
<materialsCitation collectionCode="ZMH" lastPageId="12" lastPageNumber="1327" pageId="11" pageNumber="1326" specimenCode="ZMH 833674" specimenCount="1" specimenCount-male="1">
same, except ‘WORLD/SCARAB./DATABASE/WSD00347500’ // ‘ 
<specimenCode box="[188,336,1114,1139]" pageId="12" pageNumber="1327">ZMH 833674</specimenCode>
’ (
<specimenCount box="[355,387,1114,1139]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[393,451,1114,1139]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="12" pageNumber="1327" type="Museum">ZMH</collectionCode>
).
</materialsCitation>
<materialsCitation collectionCode="ZMHB" collectorName="Alwine Braatz V'" location="Campinas" municipality="Campinas" pageId="12" pageNumber="1327" specimenCount="3" specimenCount-male="3" stateProvince="Sao Paulo">
<emphasis box="[467,575,1114,1139]" italics="true" pageId="12" pageNumber="1327">
<collectingRegion box="[467,575,1114,1139]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="1327">São Paulo</collectingRegion>
</emphasis>
. ‘Bresilien/Prov. S. Paulo/ 
<collectingMunicipality box="[855,970,1114,1139]" pageId="12" pageNumber="1327">Campinas</collectingMunicipality>
/ 
<collectorName box="[977,1155,1114,1139]" pageId="12" pageNumber="1327">Alwine Braatz V’</collectorName>
(
<specimenCount box="[167,201,1149,1174]" pageId="12" pageNumber="1327" type="male">3♂</specimenCount>
<collectionCode box="[207,280,1149,1174]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="12" pageNumber="1327">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="1911-07-05" collectionCode="ZMHB" location="Pr. S. Paulo" municipality="Pr. S. Paulo" pageId="12" pageNumber="1327" specimenCount="2" specimenCount-male="2">
<collectingMunicipality box="[296,420,1149,1174]" pageId="12" pageNumber="1327">Pr. S. Paulo</collectingMunicipality>
/Bras’. // ‘ 
<specimenCount box="[521,541,1149,1172]" pageId="12" pageNumber="1327" type="male">♂</specimenCount>
’ // ‘ 
<taxonomicName authority="Bl." authorityName="Bl." box="[579,726,1149,1174]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="genus">Byrsopolis Bl.</taxonomicName>
/quadraticeps/m.d. Type vergt./Paris 
<date pageId="12" pageNumber="1327" value="1911-07-05">
<collectingDate pageId="12" pageNumber="1327" value="1911-07-05">5. VII.1911</collectingDate>
</date>
[handwritten by Ohaus]’ (
<specimenCount box="[544,575,1183,1208]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[582,655,1183,1208]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="12" pageNumber="1327">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectionCode="ZMHB" location="Campinas" municipality="Campinas" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1" stateProvince="San Paulo">
<collectingMunicipality box="[673,786,1183,1208]" pageId="12" pageNumber="1327">Campinas</collectingMunicipality>
/ 
<collectingRegion box="[794,906,1183,1208]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="1327">San Paulo</collectingRegion>
/Bresilien’ // ‘quadraticeps Blanch’ (
<specimenCount box="[328,359,1218,1243]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[371,445,1218,1243]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="12" pageNumber="1327">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectionCode="ZMHB" location="Bresilien" municipality="Bresilien" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[468,564,1218,1243]" pageId="12" pageNumber="1327">Bresilien</collectingMunicipality>
/SanPaulo’ // ‘ 
<taxonomicName box="[732,850,1218,1243]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/quadraticeps Blanch’ (
<specimenCount box="[1124,1155,1218,1243]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[160,233,1253,1278]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="12" pageNumber="1327">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation box="[250,982,1253,1278]" collectionCode="ZMHB" location="Pr. S. Paulo" municipality="Pr. S. Paulo" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[250,373,1253,1278]" pageId="12" pageNumber="1327">Pr. S. Paulo</collectingMunicipality>
/Brasil’ // ‘ 
<taxonomicName authorityName="Blanchard" authorityYear="1851" box="[480,750,1253,1278]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="species" species="quadraticeps">Byrsopolis quadraticeps</taxonomicName>
/Blanch’ (
<specimenCount box="[854,885,1253,1278]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[891,964,1253,1278]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="12" pageNumber="1327">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectionCode="ZMHB" location="Campin" municipality="Campin" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[981,1073,1253,1278]" pageId="12" pageNumber="1327">Campin</collectingMunicipality>
. [handwritten]’ // ‘ 
<taxonomicName box="[301,417,1287,1312]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/ 
<taxonomicName authority="Blch" authorityName="Blanchard" authorityYear="1851" box="[425,559,1287,1312]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="species" species="crassa">crassa Blch</taxonomicName>
/?? Mey. [handwritten]’ // ‘ 
<taxonomicName box="[877,996,1287,1312]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/quadraticeps Blanch’ (
<specimenCount box="[267,298,1322,1347]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[312,386,1322,1347]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="12" pageNumber="1327">ZMHB</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectionCode="CMN" collectorName="Martinez'" location="Sao Paulo" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
Oct. 943/ 
<collectingRegion box="[522,647,1322,1347]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="1327">Sao Paulo</collectingRegion>
/P. Epitacio/Zellibor-leg/Coll. 
<collectorName box="[994,1100,1322,1347]" pageId="12" pageNumber="1327">Martinez’</collectorName>
(
<specimenCount box="[1124,1155,1322,1347]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[160,220,1357,1382]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="12" pageNumber="1327" type="Museum">CMN</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectionCode="CMN" collectorName="Martinez" location="Campinas" municipality="Campinas" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
Dic. 937/ 
<collectingRegion box="[355,477,1357,1382]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="1327">Sao Paulo</collectingRegion>
/ 
<collectingMunicipality box="[484,602,1357,1382]" pageId="12" pageNumber="1327">Campinas</collectingMunicipality>
/R. Spitz-leg./Coll. 
<collectorName box="[822,922,1357,1382]" pageId="12" pageNumber="1327">Martinez</collectorName>
[handwritten]’ (
<specimenCount box="[1124,1155,1357,1382]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[160,220,1391,1416]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="12" pageNumber="1327" type="Museum">CMN</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="1981-09-19" collectionCode="CEMT" collectorName="Luiz Evandro" country="Brazil" location="Faz Sta Clara" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1" stateProvince="SP">
<collectingCountry box="[244,308,1391,1416]" name="Brazil" pageId="12" pageNumber="1327">Brasil</collectingCountry>
_ 
<collectingRegion box="[346,374,1391,1416]" pageId="12" pageNumber="1327">SP</collectingRegion>
Lins/ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FE66E4797A715797" box="[438,602,1391,1416]" country="Brazil" name="Faz Sta Clara" pageId="12" pageNumber="1327" stateProvince="SP">Faz Sta Clara</location>
/ 
<date box="[610,729,1391,1416]" pageId="12" pageNumber="1327" value="1981-09-19">
<collectingDate box="[610,729,1391,1416]" pageId="12" pageNumber="1327" value="1981-09-19">19-IX-1981</collectingDate>
</date>
/ 
<collectorName box="[737,895,1391,1416]" pageId="12" pageNumber="1327">Luiz Evandro</collectorName>
/Col: Solo’ (
<specimenCount box="[1033,1065,1391,1416]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[1077,1147,1391,1416]" pageId="12" pageNumber="1327">CEMT</collectionCode>
). 
<emphasis box="[160,326,1426,1451]" italics="true" pageId="12" pageNumber="1327">Distrito Federal</emphasis>
. ‘
</materialsCitation>
<materialsCitation collectingDate="1993-12" collectionCode="EPGC" collectorName="E. &amp; P. Grossi" country="Brazil" location="Planaltina" municipality="Brasilia" pageId="12" pageNumber="1327" specimenCount="2" specimenCount-male="2" stateProvince="Distrito Federal">
<collectingMunicipality box="[343,424,1426,1451]" pageId="12" pageNumber="1327">Brasília</collectingMunicipality>
DF/ 
<collectingCountry box="[472,532,1426,1451]" name="Brazil" pageId="12" pageNumber="1327">Brasil</collectingCountry>
<date box="[541,608,1426,1451]" pageId="12" pageNumber="1327" value="1993-12">
<collectingDate box="[541,608,1426,1451]" pageId="12" pageNumber="1327" value="1993-12">XII 93</collectingDate>
</date>
’ // ‘ 
<collectorName box="[650,672,1426,1451]" pageId="12" pageNumber="1327">E.</collectorName>
&amp; 
<collectorName box="[705,805,1426,1451]" pageId="12" pageNumber="1327">P. Grossi</collectorName>
/legs [handwritten]’ (
<specimenCount box="[1039,1073,1426,1451]" pageId="12" pageNumber="1327" type="male">2♂</specimenCount>
<collectionCode box="[1082,1147,1426,1451]" pageId="12" pageNumber="1327">EPGC</collectionCode>
); ‘ 
<collectingCountry box="[163,230,1461,1486]" name="Brazil" pageId="12" pageNumber="1327">Brasil</collectingCountry>
: 
<collectingRegion box="[247,434,1461,1486]" country="Brazil" name="Distrito Federal" pageId="12" pageNumber="1327">Distrito Federal</collectingRegion>
/ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FE6AE4A37A0557D1" box="[442,558,1461,1486]" country="Brazil" municipality="Brasilia" name="Planaltina" pageId="12" pageNumber="1327" stateProvince="Distrito Federal">Planaltina</location>
.
</materialsCitation>
<materialsCitation collectingDate="2005-11-18" collectingMethod="Light" collectionCode="CERPE" collectorName="C. Oliveira'" location="Embrapa Cerrados" longitude="-47.71417" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FDEEE4A37B3657D1" box="[574,797,1461,1486]" longitude="-47.71417" name="Embrapa Cerrados" pageId="12" pageNumber="1327">Embrapa Cerrados</location>
./Área Agrícola. 
<geoCoordinate box="[990,1109,1460,1486]" pageId="12" pageNumber="1327">
15°36 
<emphasis bold="true" box="[1055,1061,1460,1484]" pageId="12" pageNumber="1327">ʹ</emphasis>
54’S
</geoCoordinate>
/ 
<geoCoordinate degrees="47" direction="west" minutes="42" orientation="longitude" pageId="12" pageNumber="1327" precision="15" seconds="51" value="-47.71417">
47° 42 
<emphasis bold="true" box="[188,194,1495,1519]" pageId="12" pageNumber="1327">ʹ</emphasis>
51”W
</geoCoordinate>
. 
<date box="[271,398,1495,1520]" pageId="12" pageNumber="1327" value="2005-11-18">
<collectingDate box="[271,398,1495,1520]" pageId="12" pageNumber="1327" value="2005-11-18">18-XI-2005</collectingDate>
</date>
./ 
<collectingMethod box="[410,467,1495,1520]" pageId="12" pageNumber="1327">Light</collectingMethod>
/ 
<collectorName box="[475,598,1495,1520]" pageId="12" pageNumber="1327">C. Oliveira’</collectorName>
(
<specimenCount box="[621,652,1495,1520]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[664,741,1495,1520]" collectionName="CERPE" pageId="12" pageNumber="1327">CERPE</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="2005-11-18" collectingMethod="Light" collectionCode="CEMT" collectorName="C. Oliveira'" location="Fragmento Cerrado" municipality="Fragmento Cerrado" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
same, except ‘ 
<collectingMunicipality box="[926,1156,1495,1520]" pageId="12" pageNumber="1327">Fragmento Cerrado</collectingMunicipality>
<geoCoordinate box="[160,288,1530,1555]" pageId="12" pageNumber="1327">
15°36 
<emphasis bold="true" box="[225,231,1530,1554]" pageId="12" pageNumber="1327">ʹ</emphasis>
20’/S
</geoCoordinate>
<geoCoordinate box="[307,446,1530,1555]" pageId="12" pageNumber="1327">
47°42 
<emphasis bold="true" box="[372,378,1530,1554]" pageId="12" pageNumber="1327">ʹ</emphasis>
26”W.
</geoCoordinate>
<date box="[464,591,1530,1555]" pageId="12" pageNumber="1327" value="2005-11-18">
<collectingDate box="[464,591,1530,1555]" pageId="12" pageNumber="1327" value="2005-11-18">18-xi-2005</collectingDate>
</date>
./ 
<collectingMethod box="[602,662,1530,1555]" pageId="12" pageNumber="1327">Light</collectingMethod>
. 
<collectorName box="[684,818,1530,1555]" pageId="12" pageNumber="1327">C. Oliveira’</collectorName>
(
<specimenCount box="[846,878,1530,1555]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[897,967,1530,1555]" pageId="12" pageNumber="1327">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation collectionCode="CEMT" location="Embrapa Cerrados" municipality="Planaltina" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
same, except ‘ 
<collectingMunicipality box="[163,276,1565,1590]" pageId="12" pageNumber="1327">Planaltina</collectingMunicipality>
./ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FECEE70B79DE5429" box="[286,501,1565,1590]" municipality="Planaltina" name="Embrapa Cerrados" pageId="12" pageNumber="1327">Embrapa Cerrados</location>
. 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FDD0E70B7AB15429" box="[512,666,1565,1590]" municipality="Planaltina" name="Area Agricola" pageId="12" pageNumber="1327">Área Agrícola</location>
. 15°36’/20’S 47°42 
<emphasis bold="true" box="[872,878,1565,1589]" pageId="12" pageNumber="1327">ʹ</emphasis>
26”W’. (
<specimenCount box="[966,998,1565,1590]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[1006,1074,1565,1590]" pageId="12" pageNumber="1327">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="2005-11-18" collectingMethod="Light" collectionCode="CEMT" collectorName="C. Oliveira'" location="Planaltina" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
same, except ‘ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FF29E72979415447" box="[249,362,1599,1624]" name="Planaltina" pageId="12" pageNumber="1327">Planaltina</location>
./ 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FEA4E7297A655447" box="[372,590,1599,1624]" name="Embrapa Cerrados" pageId="12" pageNumber="1327">Embrapa Cerrados</location>
. 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FD8CE7297AD25447" box="[604,761,1599,1624]" name="Area Agricola" pageId="12" pageNumber="1327">Área Agrícola</location>
. 15°35 
<emphasis bold="true" box="[838,844,1599,1623]" pageId="12" pageNumber="1327">ʹ</emphasis>
54’S/47°42 
<emphasis bold="true" box="[962,968,1599,1623]" pageId="12" pageNumber="1327">ʹ</emphasis>
26”W. 
<date box="[1032,1153,1599,1624]" pageId="12" pageNumber="1327" value="2005-11-18">
<collectingDate box="[1032,1153,1599,1624]" pageId="12" pageNumber="1327" value="2005-11-18">18-xi-2005</collectingDate>
</date>
. 
<collectingMethod box="[160,220,1634,1659]" pageId="12" pageNumber="1327">Light</collectingMethod>
./ 
<collectorName box="[232,350,1634,1659]" pageId="12" pageNumber="1327">C. Oliveira’</collectorName>
(
<specimenCount box="[368,399,1634,1659]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[406,474,1634,1659]" pageId="12" pageNumber="1327">CEMT</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="2015-02-11" collectingMethod="pitfall" collectionCode="CERPE" collectorName="Marina Regina Frizzas'" location="Cerrado" municipality="Cerrado" pageId="12" pageNumber="1327" specimenCount="1" specimenCount-male="1">
DF. PNB/ 
<collectingMunicipality box="[608,705,1634,1659]" pageId="12" pageNumber="1327">Cerrado</collectingMunicipality>
/ 
<date box="[713,777,1634,1659]" pageId="12" pageNumber="1327" value="2015-02-11">
<collectingDate box="[713,777,1634,1659]" pageId="12" pageNumber="1327" value="2015-02-11">11.ii.15</collectingDate>
</date>
/ 
<collectingMethod box="[785,846,1634,1659]" pageId="12" pageNumber="1327">pitfall</collectingMethod>
/ 
<collectorName box="[854,1107,1634,1659]" pageId="12" pageNumber="1327">
<taxonomicName authority="Regina" authorityName="Regina" box="[854,1020,1634,1659]" class="Magnoliopsida" family="Fabaceae" genus="Marina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="1327" phylum="Tracheophyta" rank="genus">Marina Regina</taxonomicName>
Frizzas’
</collectorName>
(
<specimenCount box="[1124,1155,1634,1659]" pageId="12" pageNumber="1327" type="male">1♂</specimenCount>
<collectionCode box="[160,238,1669,1694]" collectionName="CERPE" pageId="12" pageNumber="1327">CERPE</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="1992-11" collectionCode="MSPC" collectorName="Matthias / Seidel" country="Brazil" lastPageId="13" lastPageNumber="1328" location="Brasilia D. F." pageId="12" pageNumber="1327" specimenCode="WSD00347501, WSD00347502" specimenCount="2" specimenCount-male="2">
<collectingCountry box="[261,329,1669,1694]" name="Brazil" pageId="12" pageNumber="1327">Brazil</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:03F09661FFF48065FED5E393791C519F:8E8671ACFFF38067FE88E79379C05481" box="[344,491,1669,1694]" country="Brazil" name="Brasilia D. F." pageId="12" pageNumber="1327">Brasilia D. F.</location>
/ 
<date box="[498,581,1669,1694]" pageId="12" pageNumber="1327" value="1992-11">
<collectingDate box="[498,581,1669,1694]" pageId="12" pageNumber="1327" value="1992-11">xi.1992</collectingDate>
</date>
/local collector’ // ‘Coll. 
<collectorName box="[873,1051,1669,1694]" pageId="12" pageNumber="1327">Matthias/Seidel</collectorName>
2017’ // ‘ 
<taxonomicName box="[163,282,1703,1728]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="genus">Byrsopolis</taxonomicName>
/quadraticeps/Bl, 1851/det. M. Seidel 2017’ // ‘WORLD/SCARAB./DATABASE/ [
<specimenCode box="[165,332,142,167]" pageId="13" pageNumber="1328">WSD00347501</specimenCode>
- 
<specimenCode box="[348,517,142,167]" pageId="13" pageNumber="1328">WSD00347502</specimenCode>
]’ (
<specimenCount box="[541,575,142,167]" pageId="13" pageNumber="1328" type="male">2♂</specimenCount>
<collectionCode box="[585,654,142,167]" collectionName="Italy, Varallo, Museo di Storia Naturale &quot;Pietro Calderini&quot;" country="Italy" httpUri="http://grbio.org/cool/t9g4-v7fn" name="Museo di Storia Naturale &amp;quot;Pietro Calderini&amp;quot;" pageId="13" pageNumber="1328">MSPC</collectionCode>
).
</materialsCitation>
<materialsCitation collectionCode="IBSP" collectorName="J. Guerin'" location="Curitiba" municipality="Curitiba" pageId="13" pageNumber="1328" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
<emphasis box="[672,750,142,167]" italics="true" pageId="13" pageNumber="1328">
<collectingRegion box="[672,750,142,167]" country="Brazil" name="Parana" pageId="13" pageNumber="1328">Paraná</collectingRegion>
</emphasis>
. ‘ 
<collectingMunicipality box="[768,858,142,167]" pageId="13" pageNumber="1328">Curitiba</collectingMunicipality>
./ 
<collectingRegion box="[868,950,142,167]" country="Brazil" name="Parana" pageId="13" pageNumber="1328">Paraná</collectingRegion>
/12.936’ // ‘Coll. 
<collectorName pageId="13" pageNumber="1328">J. Guerin’</collectorName>
(
<specimenCount box="[261,293,177,202]" pageId="13" pageNumber="1328" type="male">1♂</specimenCount>
<collectionCode box="[303,356,177,202]" collectionName="Brazil,Sao Paulo, Sao Paulo, Instituto Butantan" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="13" pageNumber="1328">IBSP</collectionCode>
); ‘
</materialsCitation>
<materialsCitation collectingDate="2002-04-15" collectionCode="CEMT" collectorName="Andre Manfio Maia'" country="Brazil" location="Londrina" municipality="Londrina" pageId="13" pageNumber="1328" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
<collectingCountry box="[378,440,177,202]" name="Brazil" pageId="13" pageNumber="1328">Brasil</collectingCountry>
– 
<collectingRegion box="[475,551,177,202]" country="Brazil" name="Parana" pageId="13" pageNumber="1328">Paraná</collectingRegion>
– 
<collectingMunicipality box="[586,688,177,202]" pageId="13" pageNumber="1328">Londrina</collectingMunicipality>
/ 
<date box="[696,819,177,202]" pageId="13" pageNumber="1328" value="2002-04-15">
<collectingDate box="[696,819,177,202]" pageId="13" pageNumber="1328" value="2002-04-15">15-IV-2002</collectingDate>
</date>
/Col. 
<collectorName box="[877,1103,177,202]" pageId="13" pageNumber="1328">André Manfio Maia’</collectorName>
(
<specimenCount box="[1124,1155,177,202]" pageId="13" pageNumber="1328" type="male">1♂</specimenCount>
<collectionCode box="[160,228,212,237]" pageId="13" pageNumber="1328">CEMT</collectionCode>
).
</materialsCitation>
<materialsCitation box="[244,709,212,237]" collectionCode="CERPE, CEMT" pageId="13" pageNumber="1328" specimenCount="3" specimenCount-male="3">
<emphasis box="[244,443,212,237]" italics="true" pageId="13" pageNumber="1328">No collection data</emphasis>
(
<specimenCount box="[457,492,212,237]" pageId="13" pageNumber="1328" type="male">2♂</specimenCount>
<collectionCode box="[499,572,212,237]" collectionName="CERPE" pageId="13" pageNumber="1328">CERPE</collectionCode>
, 
<specimenCount box="[582,602,212,235]" pageId="13" pageNumber="1328" type="male">♂</specimenCount>
2 
<collectionCode box="[631,699,212,237]" pageId="13" pageNumber="1328">CEMT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7156482" ID-Zenodo-Dep="7156482" httpUri="https://zenodo.org/record/7156482/files/figure.png" pageId="12" pageNumber="1327" startId="12.[160,225,945,968]" targetBox="[163,1153,146,913]" targetPageId="12">
<paragraph blockId="12.[160,1156,945,1027]" pageId="12" pageNumber="1327">
<emphasis bold="true" box="[160,251,945,968]" pageId="12" pageNumber="1327">Figure 3.</emphasis>
<taxonomicName authority="Blanchard" authorityName="Blanchard" authorityYear="1851" box="[259,593,945,968]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1327" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis box="[259,485,945,968]" italics="true" pageId="12" pageNumber="1327">Byrsopolis quadraticeps</emphasis>
Blanchard
</taxonomicName>
, holotype (MNHN-EC-EC1804), dorsal and lateral habitus (a–b); labels attached (c); aedeagus caudal and lateral view (d–e). Images from Antoine Mantilleri (MNHN). Scale bars: a, b = 5 mm; d, e = 2 mm.
</paragraph>
</caption>
<subSubSection pageId="13" pageNumber="1328" type="diagnosis">
<paragraph blockId="13.[160,1155,292,491]" pageId="13" pageNumber="1328">
<emphasis bold="true" box="[160,282,292,318]" italics="true" pageId="13" pageNumber="1328">Diagnosis.</emphasis>
Body yellowish to yellowish brown, without metallic green reflections; clypeus subrectangular with rounded angles, apex moderately bilobed; antennal club about 2x longer than antennomeres II–VII combined; scutellar plate subtriangular, base subacute; elytral apex with right angles; elytra with indistinct striae, punctate, humeri weakly punctate; protibial spur about 4x longer than wide; parameres symmetrical, weakly convergent, moderately elongated, weakly angled internally, proximal margin bisinuate.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="1329" pageId="13" pageNumber="1328" type="description">
<paragraph blockId="13.[160,1156,546,1647]" pageId="13" pageNumber="1328">
<emphasis bold="true" box="[160,326,546,572]" italics="true" pageId="13" pageNumber="1328">Redescription.</emphasis>
<emphasis box="[342,461,547,572]" italics="true" pageId="13" pageNumber="1328">Male. Body</emphasis>
. Oval, elongated, sides rounded; yellowish to yellowish brown, without green reflections, marginal areas orange-brown to dark-brown (
<figureCitation box="[974,1077,582,607]" captionStart="Figure 3" captionStartId="12.[160,225,945,968]" captionTargetBox="[163,1153,146,913]" captionTargetId="figure-212@12.[162,1154,142,914]" captionTargetPageId="12" captionText="Figure 3. Byrsopolis quadraticeps Blanchard, holotype (MNHN-EC-EC1804), dorsal and lateral habitus (a–b); labels attached (c); aedeagus caudal and lateral view (d–e). Images from Antoine Mantilleri (MNHN). Scale bars: a, b = 5 mm; d, e = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7156482" httpUri="https://zenodo.org/record/7156482/files/figure.png" pageId="13" pageNumber="1328">Figures 3</figureCitation>
(a-b), 7 (a-b)); clypeal border darker, reddish to dark red; frontoclypeal suture darker than frons, reddish. Length 
<quantity box="[350,541,651,676]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" metricValueMax="2.54" metricValueMin="2.04" pageId="13" pageNumber="1328" unit="mm" value="22.9" valueMax="25.4" valueMin="20.4">20.40–25.40 mm</quantity>
. Humeral width 
<quantity box="[736,914,651,676]" metricMagnitude="-2" metricUnit="m" metricValue="1.1164999999999998" metricValueMax="1.283" metricValueMin="0.95" pageId="13" pageNumber="1328" unit="mm" value="11.165" valueMax="12.83" valueMin="9.5">9.50–12.83 mm</quantity>
. Elytral width 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.267" metricValueMax="1.4289999999999998" metricValueMin="1.105" pageId="13" pageNumber="1328" unit="mm" value="12.67" valueMax="14.29" valueMin="11.05">11.05– 14.29 mm</quantity>
. 
<emphasis box="[287,344,686,711]" italics="true" pageId="13" pageNumber="1328">Head</emphasis>
. Clypeus about 1.60x wider than long, subrectangular, rounded angles, weakly to moderately bilobed (
<figureCitation box="[524,619,721,746]" captionStart="Figure 7" captionStartId="18.[160,225,943,966]" captionTargetBox="[163,1152,143,911]" captionTargetId="figure-147@18.[162,1154,142,911]" captionTargetPageId="18" captionText="Figure 7. Byrsopolis quadraticeps Blanchard, Minas Gerais, Buritis. Dorsal and lateral habitus (a–b); caudal view of aedeagus (c); dorsal view of left mandible (d); ventral view of left maxilla (e); ventral view of mentum (f); dorsal view of labrum (g). Scale bars: a, b = 5 mm; c–g = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156496" httpUri="https://zenodo.org/record/7156496/files/figure.png" pageId="13" pageNumber="1328">Figure 7</figureCitation>
(a)); surface moderately concave; frontoclypeal suture almost straight between curvilinear extremities, rounded apical corners; punctures small to moderate, moderately to densely punctate; setae moderate to large, moderate to densely distributed; length of frons subequal to clypeus (0.95x shorter than clypeus); punctures small to moderately large, moderate to densely punctate; microsetae in moderate punctures; mandibles with small and sparse punctures and setae near scissorial region (
<figureCitation box="[249,344,929,954]" captionStart="Figure 7" captionStartId="18.[160,225,943,966]" captionTargetBox="[163,1152,143,911]" captionTargetId="figure-147@18.[162,1154,142,911]" captionTargetPageId="18" captionText="Figure 7. Byrsopolis quadraticeps Blanchard, Minas Gerais, Buritis. Dorsal and lateral habitus (a–b); caudal view of aedeagus (c); dorsal view of left mandible (d); ventral view of left maxilla (e); ventral view of mentum (f); dorsal view of labrum (g). Scale bars: a, b = 5 mm; c–g = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156496" httpUri="https://zenodo.org/record/7156496/files/figure.png" pageId="13" pageNumber="1328">Figure 7</figureCitation>
(d)); galea with three equidistant teeth, equidistant, apical tooth longer and thicker than basal tooth, basal tooth longer and thicker than medial tooth, basal tooth with two long setae (
<figureCitation box="[409,504,998,1023]" captionStart="Figure 7" captionStartId="18.[160,225,943,966]" captionTargetBox="[163,1152,143,911]" captionTargetId="figure-147@18.[162,1154,142,911]" captionTargetPageId="18" captionText="Figure 7. Byrsopolis quadraticeps Blanchard, Minas Gerais, Buritis. Dorsal and lateral habitus (a–b); caudal view of aedeagus (c); dorsal view of left mandible (d); ventral view of left maxilla (e); ventral view of mentum (f); dorsal view of labrum (g). Scale bars: a, b = 5 mm; c–g = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156496" httpUri="https://zenodo.org/record/7156496/files/figure.png" pageId="13" pageNumber="1328">Figure 7</figureCitation>
(e)); last maxillary palpomere with oblong sensorial area; apex of prementum weakly emarginate; last labial palpomere about 2x longer than palpomere II (
<figureCitation box="[327,426,1067,1092]" captionStart="Figure 7" captionStartId="18.[160,225,943,966]" captionTargetBox="[163,1152,143,911]" captionTargetId="figure-147@18.[162,1154,142,911]" captionTargetPageId="18" captionText="Figure 7. Byrsopolis quadraticeps Blanchard, Minas Gerais, Buritis. Dorsal and lateral habitus (a–b); caudal view of aedeagus (c); dorsal view of left mandible (d); ventral view of left maxilla (e); ventral view of mentum (f); dorsal view of labrum (g). Scale bars: a, b = 5 mm; c–g = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156496" httpUri="https://zenodo.org/record/7156496/files/figure.png" pageId="13" pageNumber="1328">Figure 7</figureCitation>
(f)); labrum strongly emarginate (
<figureCitation box="[818,917,1067,1092]" captionStart="Figure 7" captionStartId="18.[160,225,943,966]" captionTargetBox="[163,1152,143,911]" captionTargetId="figure-147@18.[162,1154,142,911]" captionTargetPageId="18" captionText="Figure 7. Byrsopolis quadraticeps Blanchard, Minas Gerais, Buritis. Dorsal and lateral habitus (a–b); caudal view of aedeagus (c); dorsal view of left mandible (d); ventral view of left maxilla (e); ventral view of mentum (f); dorsal view of labrum (g). Scale bars: a, b = 5 mm; c–g = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156496" httpUri="https://zenodo.org/record/7156496/files/figure.png" pageId="13" pageNumber="1328">Figure 7</figureCitation>
(g)); antennomere III subcylindrical, antennomere IV subquadrate, antennomeres V–VII subconical; antennal club about 2x longer than antennomeres II–VII combined. 
<emphasis box="[825,900,1137,1162]" italics="true" pageId="13" pageNumber="1328">Thorax</emphasis>
. Pronotum transverse, about 1.5x wider than long (
<figureCitation box="[475,566,1171,1196]" captionStart="Figure 7" captionStartId="18.[160,225,943,966]" captionTargetBox="[163,1152,143,911]" captionTargetId="figure-147@18.[162,1154,142,911]" captionTargetPageId="18" captionText="Figure 7. Byrsopolis quadraticeps Blanchard, Minas Gerais, Buritis. Dorsal and lateral habitus (a–b); caudal view of aedeagus (c); dorsal view of left mandible (d); ventral view of left maxilla (e); ventral view of mentum (f); dorsal view of labrum (g). Scale bars: a, b = 5 mm; c–g = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7156496" httpUri="https://zenodo.org/record/7156496/files/figure.png" pageId="13" pageNumber="1328">Figure 7</figureCitation>
(a)); punctures small to moderately large, moderately to densely punctate, glabrous; marginal areas sparsely to moderately punctate, with sparse setae; longitudinal midline formed by absence of punctures; scutellar plate subtriangular, subacute, sides almost right, base moderately rounded, small to moderate setae, punctures small to moderately large, moderately to densely punctate. 
<emphasis box="[1013,1074,1310,1335]" italics="true" pageId="13" pageNumber="1328">Elytra</emphasis>
. About 1.10x longer than wide; elytral surface smooth, without striae; punctures small to moderately large, moderately to densely punctate (
<figureCitation box="[674,780,1379,1404]" captionStart="Figure 3" captionStartId="12.[160,225,945,968]" captionTargetBox="[163,1153,146,913]" captionTargetId="figure-212@12.[162,1154,142,914]" captionTargetPageId="12" captionText="Figure 3. Byrsopolis quadraticeps Blanchard, holotype (MNHN-EC-EC1804), dorsal and lateral habitus (a–b); labels attached (c); aedeagus caudal and lateral view (d–e). Images from Antoine Mantilleri (MNHN). Scale bars: a, b = 5 mm; d, e = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7156482" httpUri="https://zenodo.org/record/7156482/files/figure.png" pageId="13" pageNumber="1328">Figures 3</figureCitation>
(a), 3(a)); microsetae in moderate punctures; humeri weakly punctate; interstriae without coalescent punctures, moderately distributed. 
<emphasis box="[296,344,1449,1474]" italics="true" pageId="13" pageNumber="1328">Legs</emphasis>
. Protibial spur about 4x longer than wide; mesotibiae with two carinae; metatibiae with one carina. 
<emphasis box="[488,593,1483,1508]" italics="true" pageId="13" pageNumber="1328">Aedeagus</emphasis>
. Parameres moderately elongated; basal margin bisinuate; apex rounded, symmetrical, subparallel, weakly convergent, separated by a Ushaped gap; apex with weakly sinuous internal angles; paramere with lateral margins longitudinally excavated forming a moderate carina; weak longitudinal concavity near base (
<figureCitation box="[227,332,1622,1647]" captionStart="Figure 3" captionStartId="12.[160,225,945,968]" captionTargetBox="[163,1153,146,913]" captionTargetId="figure-212@12.[162,1154,142,914]" captionTargetPageId="12" captionText="Figure 3. Byrsopolis quadraticeps Blanchard, holotype (MNHN-EC-EC1804), dorsal and lateral habitus (a–b); labels attached (c); aedeagus caudal and lateral view (d–e). Images from Antoine Mantilleri (MNHN). Scale bars: a, b = 5 mm; d, e = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7156482" httpUri="https://zenodo.org/record/7156482/files/figure.png" pageId="13" pageNumber="1328">Figures 3</figureCitation>
(d,e), 3(c)).
</paragraph>
<paragraph blockId="13.[160,383,1702,1728]" box="[160,383,1702,1728]" pageId="13" pageNumber="1328">
<emphasis bold="true" box="[160,253,1702,1728]" italics="true" pageId="13" pageNumber="1328">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="14.[160,1155,142,237]" pageId="14" pageNumber="1329">
<emphasis bold="true" box="[160,277,142,168]" italics="true" pageId="14" pageNumber="1329">Variation.</emphasis>
Some specimens exhibit asymmetrical maxillae, lacking apical and basal teeth in one or both galea; labial palps with different colour, one darker than another; some specimens have dark brown or black spots on elytral surface.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="1329" type="etymology">
<paragraph blockId="14.[160,1155,281,341]" pageId="14" pageNumber="1329">
<emphasis bold="true" box="[160,291,281,307]" italics="true" pageId="14" pageNumber="1329">Etymology.</emphasis>
The species name 
<taxonomicName authorityName="Blanchard" authorityYear="1851" box="[526,666,282,307]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1329" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis box="[526,666,282,307]" italics="true" pageId="14" pageNumber="1329">quadraticeps</emphasis>
</taxonomicName>
is derived from Latin words 
<emphasis box="[1010,1121,282,307]" italics="true" pageId="14" pageNumber="1329">quadrātus</emphasis>
(= quadrate) and 
<emphasis box="[327,374,316,341]" italics="true" pageId="14" pageNumber="1329">ceps</emphasis>
(= head) reffering to the quadrate shape of the head.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="1329" type="distribution">
<paragraph blockId="14.[160,1156,385,480]" pageId="14" pageNumber="1329">
<emphasis bold="true" box="[160,307,385,411]" italics="true" pageId="14" pageNumber="1329">Distribution.</emphasis>
<collectingRegion box="[323,465,386,411]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="1329">Minas Gerais</collectingRegion>
(Buritis and Ijaci), 
<collectingRegion box="[669,731,386,411]" country="Brazil" name="Goias" pageId="14" pageNumber="1329">Goiás</collectingRegion>
(Leopoldo de Bulhões and Cristalina), 
<collectingRegion box="[160,336,420,445]" country="Mexico" name="Distrito Federal" pageId="14" pageNumber="1329">Distrito Federal</collectingRegion>
(Brasília and Planaltina), 
<collectingRegion box="[632,747,420,445]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="1329">São Paulo</collectingRegion>
(
<collectingRegion box="[763,882,420,445]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="1329">São Paulo</collectingRegion>
, Campinas) and 
<collectingRegion box="[1079,1155,420,445]" country="Brazil" name="Parana" pageId="14" pageNumber="1329">Paraná</collectingRegion>
(Curitiba, Londrina) (
<figureCitation box="[391,496,455,480]" captionStart="Figure 19" captionStartId="41.[160,225,961,984]" captionTargetBox="[162,1154,142,929]" captionTargetId="figure-144@41.[162,1154,142,929]" captionTargetPageId="41" captionText="Figure 19. Distribution map of Byrsopolis species." figureDoi="http://doi.org/10.5281/zenodo.7156532" httpUri="https://zenodo.org/record/7156532/files/figure.png" pageId="14" pageNumber="1329">Figure 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="1329" type="discussion">
<paragraph blockId="14.[160,1156,524,896]" pageId="14" pageNumber="1329">
<emphasis bold="true" box="[160,268,524,550]" italics="true" pageId="14" pageNumber="1329">Remarks.</emphasis>
This species was described from 
<collectingRegion box="[668,813,524,549]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="1329">Minas Gerais</collectingRegion>
in the original description of 
<bibRefCitation author="Blanchard E" box="[160,352,559,584]" pageId="14" pageNumber="1329" pagination="241" refId="ref25750" refString="Blanchard E. 1851. Gide et Baudry. In: Milne-Edwards H, Blanchard E, Lucas H, editors. Catalogue des collections Entomologiques du Museum d'Histoire naturelle de Paris, Classe des Insectes. Ordre des Coleopteres. Paris, France: Museum d'histoire naturelle de Paris. (1850) 1. (1850) 1128, (1851) 129240; p. 129240; p. 241." type="journal article" year="1851">Blanchard (1851)</bibRefCitation>
; however, he did not specify the city where Saint-Hilaire collected the specimens used for the description. The 
<typeStatus box="[605,655,594,619]" pageId="14" pageNumber="1329">type</typeStatus>
was possibly collected in Serra do Espinhaço, occurring in sympatry with 
<taxonomicName authorityName="Burmeister" authorityYear="1855" box="[471,674,628,653]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1329" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis box="[471,674,628,653]" italics="true" pageId="14" pageNumber="1329">Byrsopolis laticollis</emphasis>
</taxonomicName>
. 
<taxonomicName authorityName="Blanchard" authorityYear="1851" box="[687,943,628,653]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1329" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis box="[687,943,628,653]" italics="true" pageId="14" pageNumber="1329">Byrsopolis quadraticeps</emphasis>
</taxonomicName>
was at the time of its description the only species within the genus with a bilobed and size-reduced clypeus compared to the remaining species. In the present study a second species similar to 
<taxonomicName authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1329" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis italics="true" pageId="14" pageNumber="1329">B. quadraticeps</emphasis>
</taxonomicName>
is described as 
<taxonomicName authority="Medeiros &amp; Seidel &amp; Grossi, 2022" authorityName="Medeiros &amp; Seidel &amp; Grossi" authorityYear="2022" box="[480,726,732,757]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1329" phylum="Arthropoda" rank="species" species="angelootatii" status="sp. nov.">
<emphasis box="[480,726,732,757]" italics="true" pageId="14" pageNumber="1329">Byrsopolis angelootatii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[734,821,732,757]" pageId="14" pageNumber="1329" rank="species">sp. nov.</taxonomicNameLabel>
(see below). Additionally, the records for the states of 
<collectingRegion box="[456,520,767,792]" country="Brazil" name="Goias" pageId="14" pageNumber="1329">Goiás</collectingRegion>
, 
<collectingRegion box="[534,713,767,792]" country="Mexico" name="Distrito Federal" pageId="14" pageNumber="1329">Distrito Federal</collectingRegion>
, 
<collectingRegion box="[727,842,767,792]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="1329">São Paulo</collectingRegion>
and 
<collectingRegion box="[907,983,767,792]" country="Brazil" name="Parana" pageId="14" pageNumber="1329">Paraná</collectingRegion>
are new state records. According to label data, this species is found in the Midwest, Southeast and South regions. It occurs predominantly in the Cerrado biome and can be collected with the aid of light traps.
</paragraph>
</subSubSection>
</treatment>
</document>