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<document id="39F420FA4C32A55D9591D819D0A3952B" ID-DOI="10.11646/zootaxa.3985.3.4" ID-GBIF-Dataset="ed70e8d1-c948-4fcb-8f55-313589a2125b" ID-ISSN="1175-5326" ID-Zenodo-Dep="243242" ID-ZooBank="1A2D2123-FC7B-4490-B23F-ABE88838F0E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461258546781" checkinUser="plazi" docAuthor="Valdesalici, Stefano &amp; Voort, Stefan Van Der" docDate="2015" docId="03F05146FFEEBD11FF50FF289E4D283F" docLanguage="en" docName="zt03985p408.pdf" docOrigin="Zootaxa 3985 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Badis autumnum Valdesalici &amp; Voort, 2015, new species" docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="FFC9293EFFEBBD19FFC7FFBF9A632C67" masterDocTitle="Four new species of the Indo-Burmese genus Badis from West Bengal, India (Actinopterygii: Perciformes: Badidae)" masterLastPageNumber="408" masterPageNumber="391" pageNumber="396" updateTime="1698608819460" updateUser="plazi">
<mods:mods id="6815DCF4821D2903E7987952EE5ED191" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EA1306F0C4D1EB5C286729E1C340E647">
<mods:title id="709CC3B42D8FDEDA9F9D24C1B1058291">Four new species of the Indo-Burmese genus Badis from West Bengal, India (Actinopterygii: Perciformes: Badidae)</mods:title>
</mods:titleInfo>
<mods:name id="16072EE8C96D6740D0F20FEFA10EF5FA" type="personal">
<mods:role id="89FEDB973178401AE0FFECEEF3D02126">
<mods:roleTerm id="E82C855758C0CEBEC15174B743ADC8A4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA5DAB32EFB322C910D3C85A986689C0">Valdesalici, Stefano</mods:namePart>
</mods:name>
<mods:name id="09A50FC1718D944474DD85D5716BE1B6" type="personal">
<mods:role id="AD99640D0DD7E2EAAE72FEDCBEB0AFD8">
<mods:roleTerm id="BBA1B51E252BF63ED66E859678DF5B72">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3EB8E5CFDB2E25FFAC081980AFD4E7E">Voort, Stefan Van Der</mods:namePart>
</mods:name>
<mods:typeOfResource id="0C93251FA928A1D7CDDF459828DCC683">text</mods:typeOfResource>
<mods:relatedItem id="A6FA4C15D34D08CB39266C99FEFE1526" type="host">
<mods:titleInfo id="0E665F5A5503D24469BDDB9DE1B04A49">
<mods:title id="CA65D46EF5964C855F3A630FC28EE9C0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="32959B5AB8ED15951A2C6178BCB9BEF5">
<mods:date id="7CB808BE768B8D844823B731F959190D">2015</mods:date>
<mods:detail id="314792903AAF2001B205D1210755BA5D" type="volume">
<mods:number id="8DE243C320054B52C0AB8EC1824A8E3F">3985</mods:number>
</mods:detail>
<mods:detail id="20556D2BE59956C023224DED8E1EC8A1" type="issue">
<mods:number id="D012C4498217FF060109753B3DD96AD8">3</mods:number>
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<mods:start id="A81454378FF8223D695C0C9F382597CD">391</mods:start>
<mods:end id="3D50E04B0CF620AB63DD46659C91ABB3">408</mods:end>
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<mods:classification id="A21A0D05988B2C1B95DCCDE2D3E728F3">journal article</mods:classification>
<mods:identifier id="7E1A02ED3A41D1FD71D07681C767FE6E" type="DOI">10.11646/zootaxa.3985.3.4</mods:identifier>
<mods:identifier id="0FFA5A8AA7760D4E1FBE0A0AD335ABA6" type="GBIF-Dataset">ed70e8d1-c948-4fcb-8f55-313589a2125b</mods:identifier>
<mods:identifier id="22BDFB1847D216323CB93B0A0D90137A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F5E39D399BCCF4169FCD9CE05B1AB9BE" type="Zenodo-Dep">243242</mods:identifier>
<mods:identifier id="9B7886777580B70D702F625F49FCA972" type="ZooBank">1A2D2123-FC7B-4490-B23F-ABE88838F0E4</mods:identifier>
</mods:mods>
<treatment id="03F05146FFEEBD11FF50FF289E4D283F" ID-DOI="http://doi.org/10.5281/zenodo.6096986" ID-GBIF-Taxon="119636590" ID-Zenodo-Dep="6096986" LSID="urn:lsid:plazi:treatment:03F05146FFEEBD11FF50FF289E4D283F" httpUri="http://treatment.plazi.org/id/03F05146FFEEBD11FF50FF289E4D283F" lastPageId="8" lastPageNumber="399" pageId="5" pageNumber="396">
<subSubSection id="C343B3DBFFEEBD1CFF50FF289B712CB4" pageId="5" pageNumber="396" type="nomenclature">
<paragraph id="8BE6E050FFEEBD1CFF50FF2898652CD5" blockId="5.[151,518,151,211]" box="[151,518,151,178]" pageId="5" pageNumber="396">
<heading id="D0AE573CFFEEBD1CFF50FF2898652CD5" bold="true" box="[151,518,151,178]" fontSize="11" level="1" pageId="5" pageNumber="396" reason="1">
<emphasis id="B92D3C42FFEEBD1CFF50FF2898652CD5" bold="true" box="[151,518,151,178]" pageId="5" pageNumber="396">
<taxonomicName id="4C599BD3FFEEBD1CFF50FF289B0B2CD6" ID-CoL="KBQC" box="[151,360,151,177]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="autumnum" status="sp. nov.">
<emphasis id="B92D3C42FFEEBD1CFF50FF289B0B2CD6" bold="true" box="[151,360,151,177]" italics="true" pageId="5" pageNumber="396">Badis autumnum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21E8139FFEEBD1CFEB0FF2798652CD5" box="[375,518,152,178]" pageId="5" pageNumber="396" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE6E050FFEEBD1CFF50FF059B712CB4" blockId="5.[151,518,151,211]" box="[151,274,186,211]" pageId="5" pageNumber="396">
(
<figureCitation id="1362FCD5FFEEBD1CFF58FF059B6A2CB4" box="[159,265,186,211]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,1950,1972]" captionStartId-1="6.[151,250,750,772]" captionStartId-2="6.[151,250,1730,1752]" captionTargetBox-0="[151,1435,1415,1929]" captionTargetBox-1="[244,1342,289,665]" captionTargetBox-2="[151,1435,852,1708]" captionTargetId-0="figure@5.[151,1435,1415,1929]" captionTargetId-1="figure@6.[150,1436,193,729]" captionTargetId-2="figure@6.[151,1436,852,1708]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="6" captionText-0="FIGURE 4. Badis autumnum, MSNG 57568, paratype, male, 35.9 mm SL; India: West Bengal, Singimari River drainage, Rathbari stream at Petla. Note the oblique stripe on operculum." captionText-1="FIGURE 5. Badis autumnum, MSNG 57570, paratype, female, 29.4 mm SL; India: West Bengal, Singimari River drainage, Rathbari stream at Petla. Note the blotch above base of opercular spine." captionText-2="FIGURE 6. Live male individual of Badis autumnum, ca 35 mm SL (not included in type series); India: West Bengal, Singimari River drainage, Rathbari stream at Petla." httpUri-0="https://zenodo.org/record/243246/files/figure.png" httpUri-1="https://zenodo.org/record/243247/files/figure.png" httpUri-2="https://zenodo.org/record/243248/files/figure.png" pageId="5" pageNumber="396">Figs. 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C343B3DBFFEEBD1CFF50FEBD98E52DA8" pageId="5" pageNumber="396" type="materials_examined">
<paragraph id="8BE6E050FFEEBD1CFF50FEBD9AAF2D05" blockId="5.[151,1437,258,1363]" pageId="5" pageNumber="396">
<emphasis id="B92D3C42FFEEBD1CFF50FEBD9B682D7C" bold="true" box="[151,267,258,283]" pageId="5" pageNumber="396">
<typeStatus id="54E25EF2FFEEBD1CFF50FEBD9B652D7C" box="[151,262,258,283]" pageId="5" pageNumber="396" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED487895FFEEBD1CFED5FEBC9B002D7C" box="[274,355,259,283]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="5" pageNumber="396">MSNG</collectionCode>
57567,
<quantity id="4CA14DB5FFEEBD1CFE7AFEBD98422D7D" box="[445,545,258,283]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="5" pageNumber="396" unit="mm" value="35.8">35.8 mm</quantity>
SL, male,
<date id="FFE7C690FFEEBD1CFD5DFEBC99142D7C" box="[666,887,258,283]" pageId="5" pageNumber="396" value="2013-09-20">20 September 2013</date>
;
<collectingCountry id="F34EA0C0FFEEBD1CFC45FEBD99A32D7C" box="[898,960,258,283]" name="India" pageId="5" pageNumber="396">India</collectingCountry>
: West Bengal: Cooch Behar district: Petla: near Dinhata town: Singimari River drainage, Rathbari stream,
<geoCoordinate id="EE6D8697FFEEBD1CFCBBFE999E7A2D58" box="[892,1049,294,319]" direction="north" orientation="latitude" pageId="5" pageNumber="396" precision="1" value="26.223528">26°12’84.7”N</geoCoordinate>
<geoCoordinate id="EE6D8697FFEEBD1CFBE2FE999EB22D58" box="[1061,1233,294,319]" direction="east" orientation="longitude" pageId="5" pageNumber="396" precision="1" value="89.734764">89°43’65.15”E</geoCoordinate>
; coll. Andrew A. Rao.
</paragraph>
<paragraph id="8BE6E050FFEEBD1CFF00FED198E52DA8" blockId="5.[151,1437,258,1363]" pageId="5" pageNumber="396">
<emphasis id="B92D3C42FFEEBD1CFF00FED19B252DE0" bold="true" box="[199,326,366,391]" pageId="5" pageNumber="396">
<typeStatus id="54E25EF2FFEEBD1CFF00FED19B222DE0" box="[199,321,366,391]" pageId="5" pageNumber="396" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED487895FFEEBD1CFE97FED09BC22DE0" box="[336,417,367,391]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="5" pageNumber="396">MSNG</collectionCode>
57568,
<specimenCount id="9D5F2BD9FFEEBD1CFDC5FED098302DE0" box="[514,595,366,391]" pageId="5" pageNumber="396" type="male">1 male</specimenCount>
,
<quantity id="4CA14DB5FFEEBD1CFDA5FED198AA2DE1" box="[610,713,366,391]" metricMagnitude="-2" metricUnit="m" metricValue="3.59" pageId="5" pageNumber="396" unit="mm" value="35.9">35.9 mm</quantity>
SL,
<date id="FFE7C690FFEEBD1CFCC3FED099F22DE0" box="[772,913,366,391]" pageId="5" pageNumber="396" value="2013-07-08">8 July 2013</date>
: same location as
<typeStatus id="54E25EF2FFEEBD1CFBAEFED19EAC2DE0" box="[1129,1231,366,391]" pageId="5" pageNumber="396" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED487895FFEEBD1CFB1BFED09F4E2DE0" box="[1244,1325,367,391]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="5" pageNumber="396">MSNG</collectionCode>
57569,
<specimenCount id="9D5F2BD9FFEEBD1CFA49FED09AB32DCC" pageId="5" pageNumber="396" type="male">1 male</specimenCount>
,
<quantity id="4CA14DB5FFEEBD1CFF26FE2C9B282DCD" box="[225,331,403,427]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="5" pageNumber="396" unit="mm" value="29.4">29.4 mm</quantity>
SL C&amp;S,
<date id="FFE7C690FFEEBD1CFE16FE2C98072DCC" box="[465,612,402,427]" pageId="5" pageNumber="396" value="2013-07-08">8 July 2013</date>
: same location as
<typeStatus id="54E25EF2FFEEBD1CFC80FE2D99CE2DCC" box="[839,941,402,427]" pageId="5" pageNumber="396" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED487895FFEEBD1CFC7BFE2C9E6E2DCC" box="[956,1037,403,427]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="5" pageNumber="396">MSNG</collectionCode>
57570, 5,
<quantity id="4CA14DB5FFEEBD1CFB51FE2D9F5C2DCD" box="[1174,1343,402,427]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" metricValueMax="3.27" metricValueMin="2.75" pageId="5" pageNumber="396" unit="mm" value="30.1" valueMax="32.7" valueMin="27.5">27.5–32.7 mm</quantity>
SL,
<date id="FFE7C690FFEEBD1CFA47FE2C9B322DA8" pageId="5" pageNumber="396" value="2013-09-20">20 September 2013</date>
: same location as
<typeStatus id="54E25EF2FFEEBD1CFDDAFE0998E02DA8" box="[541,643,438,463]" pageId="5" pageNumber="396" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C343B3DBFFEEBD1CFF00FE65983828F8" pageId="5" pageNumber="396" type="diagnosis">
<paragraph id="8BE6E050FFEEBD1CFF00FE65983828F8" blockId="5.[151,1437,258,1363]" pageId="5" pageNumber="396">
<emphasis id="B92D3C42FFEEBD1CFF00FE659B222D94" bold="true" box="[199,321,474,499]" pageId="5" pageNumber="396">Diagnosis.</emphasis>
<taxonomicName id="4C599BD3FFEEBD1CFE8EFE64986B2D95" box="[329,520,475,499]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEEBD1CFE8EFE64986B2D95" box="[329,520,475,499]" italics="true" pageId="5" pageNumber="396">Badis autumnum</emphasis>
</taxonomicName>
differs from all remaining
<taxonomicName id="4C599BD3FFEEBD1CFCF9FE64991D2D94" box="[830,894,475,499]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="genus">
<emphasis id="B92D3C42FFEEBD1CFCF9FE64991D2D94" box="[830,894,475,499]" italics="true" pageId="5" pageNumber="396">Badis</emphasis>
</taxonomicName>
species by a colour pattern which includes two autapomorphies.
<taxonomicName id="4C599BD3FFEEBD1CFE9AFE4198432E71" box="[349,544,510,534]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEEBD1CFE9AFE4198432E71" box="[349,544,510,534]" italics="true" pageId="5" pageNumber="396">Badis autumnum</emphasis>
</taxonomicName>
can be distinguished from every member of the
<taxonomicName id="4C599BD3FFEEBD1CFBB2FE409EB42E71" box="[1141,1239,510,534]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="badis">
<emphasis id="B92D3C42FFEEBD1CFBB2FE409EB42E71" box="[1141,1239,510,534]" italics="true" pageId="5" pageNumber="396">B. badis</emphasis>
</taxonomicName>
group (included species:
<taxonomicName id="4C599BD3FFEEBD1CFF31FD9C9B322E5C" box="[246,337,547,571]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="badis">
<emphasis id="B92D3C42FFEEBD1CFF31FD9C9B322E5C" box="[246,337,547,571]" italics="true" pageId="5" pageNumber="396">B. badis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C599BD3FFEEBD1CFE98FD9C986B2E5C" box="[351,520,547,571]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="chittagongis">
<emphasis id="B92D3C42FFEEBD1CFE98FD9C986B2E5C" box="[351,520,547,571]" italics="true" pageId="5" pageNumber="396">B. chittagongis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C599BD3FFEEBD1CFDD1FD9C98C92E5C" box="[534,682,547,571]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="dibruensis">
<emphasis id="B92D3C42FFEEBD1CFDD1FD9C98C92E5C" box="[534,682,547,571]" italics="true" pageId="5" pageNumber="396">B. dibruensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C599BD3FFEEBD1CFD7FFD9C99502E5C" box="[696,819,547,571]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="ferrarisi">
<emphasis id="B92D3C42FFEEBD1CFD7FFD9C99502E5C" box="[696,819,547,571]" italics="true" pageId="5" pageNumber="396">B. ferrarisi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C599BD3FFEEBD1CFC85FD9C99DE2E5C" box="[834,957,547,571]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="kanabos">
<emphasis id="B92D3C42FFEEBD1CFC85FD9C99DE2E5C" box="[834,957,547,571]" italics="true" pageId="5" pageNumber="396">B. kanabos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C599BD3FFEEBD1CFC0BFD9C9E552E5D" box="[972,1078,547,570]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="soraya">
<emphasis id="B92D3C42FFEEBD1CFC0BFD9C9E552E5D" box="[972,1078,547,570]" italics="true" pageId="5" pageNumber="396">B. soraya</emphasis>
</taxonomicName>
and
<taxonomicName id="4C599BD3FFEEBD1CFBABFD9C9EBC2E5D" box="[1132,1247,547,570]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="tuivaiei">
<emphasis id="B92D3C42FFEEBD1CFBABFD9C9EBC2E5D" box="[1132,1247,547,570]" italics="true" pageId="5" pageNumber="396">B. tuivaiei</emphasis>
</taxonomicName>
) by absence of a blotch on the superficial part of the cleithrum above pectoral-fin base, from all members of the
<taxonomicName id="4C599BD3FFEEBD1CFB7FFDF89F372E39" box="[1208,1364,583,606]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="B92D3C42FFEEBD1CFB7FFDF89F372E39" box="[1208,1364,583,606]" italics="true" pageId="5" pageNumber="396">B. assamensis</emphasis>
</taxonomicName>
group (included species:
<taxonomicName id="4C599BD3FFEEBD1CFEACFDD4986A2EE5" box="[363,521,619,642]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="B92D3C42FFEEBD1CFEACFDD4986A2EE5" box="[363,521,619,642]" italics="true" pageId="5" pageNumber="396">B. assamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C599BD3FFEEBD1CFD83FDD498A02EE4" box="[580,707,619,643]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="blosyrus">
<emphasis id="B92D3C42FFEEBD1CFD83FDD498A02EE4" box="[580,707,619,643]" italics="true" pageId="5" pageNumber="396">B. blosyrus</emphasis>
</taxonomicName>
) by absence of a posterodorsal opercular blotch and absence of two parallel rows of dark spots and alternating light and dark stripes along its physique, from every member of the
<taxonomicName id="4C599BD3FFEEBD1CFF50FD0C9A902EAC" box="[151,243,691,715]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B92D3C42FFEEBD1CFF50FD0C9A902EAC" box="[151,243,691,715]" italics="true" pageId="5" pageNumber="396">B. ruber</emphasis>
</taxonomicName>
group (included species:
<taxonomicName id="4C599BD3FFEEBD1CFDD5FD0C981B2EAC" box="[530,632,691,715]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="khwae">
<emphasis id="B92D3C42FFEEBD1CFDD5FD0C981B2EAC" box="[530,632,691,715]" italics="true" pageId="5" pageNumber="396">B. khwae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C599BD3FFEEBD1CFD42FD0C98812EAC" box="[645,738,691,715]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="ruber">
<emphasis id="B92D3C42FFEEBD1CFD42FD0C98812EAC" box="[645,738,691,715]" italics="true" pageId="5" pageNumber="396">B. ruber</emphasis>
</taxonomicName>
and
<taxonomicName id="4C599BD3FFEEBD1CFCDFFD0C99C02EAD" box="[792,931,691,714]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="B92D3C42FFEEBD1CFCDFFD0C99C02EAD" box="[792,931,691,714]" italics="true" pageId="5" pageNumber="396">B. siamensis</emphasis>
</taxonomicName>
) by absence of a cleithral blotch and absence of a blotch on the dorsolateral aspect of the caudal peduncle, from all members of the
<taxonomicName id="4C599BD3FFEEBD1CFB93FD689E802E89" box="[1108,1251,727,750]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="corycaeus">
<emphasis id="B92D3C42FFEEBD1CFB93FD689E802E89" box="[1108,1251,727,750]" italics="true" pageId="5" pageNumber="396">B. corycaeus</emphasis>
</taxonomicName>
group (included species:
<taxonomicName id="4C599BD3FFEEBD1CFF31FD449BE42F75" box="[246,391,763,786]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="corycaeus">
<emphasis id="B92D3C42FFEEBD1CFF31FD449BE42F75" box="[246,391,763,786]" italics="true" pageId="5" pageNumber="396">B. corycaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C599BD3FFEEBD1CFE7AFD4498462F75" box="[445,549,763,786]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="pyema">
<emphasis id="B92D3C42FFEEBD1CFE7AFD4498462F75" box="[445,549,763,786]" italics="true" pageId="5" pageNumber="396">B. pyema</emphasis>
</taxonomicName>
) by absence of an ocellus on the caudal-fin base, from
<taxonomicName id="4C599BD3FFEEBD1CFB49FD449EBC2F74" box="[1166,1247,763,787]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="kyar">
<emphasis id="B92D3C42FFEEBD1CFB49FD449EBC2F74" box="[1166,1247,763,787]" italics="true" pageId="5" pageNumber="396">B. kyar</emphasis>
</taxonomicName>
by presence of a conspicuous median black blotch on the caudal peduncle, from
<taxonomicName id="4C599BD3FFEEBD1CFC4BFCA09E532F51" box="[908,1072,799,822]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="singenensis">
<emphasis id="B92D3C42FFEEBD1CFC4BFCA09E532F51" box="[908,1072,799,822]" italics="true" pageId="5" pageNumber="396">B. singenensis</emphasis>
</taxonomicName>
by absence of a posterodorsal opercle blotch and absence of three dorsal-fin blotches and a single round blotch on the anal-fin base, from
<taxonomicName id="4C599BD3FFEEBD1CFA43FCFC9B292F19" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="juergenschmidti">
<emphasis id="B92D3C42FFEEBD1CFA43FCFC9B292F19" italics="true" pageId="5" pageNumber="396">B. juergenschmidti</emphasis>
</taxonomicName>
by absence of white margins on dorsal and ventral aspects of the caudal fin in males and presence of a strongly curved caudal-fin base bar, and from
<taxonomicName id="4C599BD3FFEEBD1CFD1EFC3499562FC4" box="[729,821,907,931]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="britzi">
<emphasis id="B92D3C42FFEEBD1CFD1EFC3499562FC4" box="[729,821,907,931]" italics="true" pageId="5" pageNumber="396">B. britzi</emphasis>
</taxonomicName>
by presence of a conspicuous median black blotch on the caudal peduncle. The new species most closely resembles
<taxonomicName id="4C599BD3FFEEBD1CFCAAFC109E712FA1" box="[877,1042,942,966]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="andrewraoi">
<emphasis id="B92D3C42FFEEBD1CFCAAFC109E712FA1" box="[877,1042,942,966]" italics="true" pageId="5" pageNumber="396">B. andrewraoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C599BD3FFEEBD1CFB88FC109EDC2FA1" box="[1103,1215,942,966]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="kyanos">
<emphasis id="B92D3C42FFEEBD1CFB88FC109EDC2FA1" box="[1103,1215,942,966]" italics="true" pageId="5" pageNumber="396">B. kyanos</emphasis>
</taxonomicName>
in terms of colour pattern, from which it can be distinguished by two exclusive characters that also distinguish it from all other species of the genus: presence of a blotch above the base of the opercular spine, and of a conspicuous dark blotch on pectoral-fin base. It also differs from
<taxonomicName id="4C599BD3FFEEBD1CFDAAFBA498BE2854" box="[621,733,1051,1075]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="kyanos">
<emphasis id="B92D3C42FFEEBD1CFDAAFBA498BE2854" box="[621,733,1051,1075]" italics="true" pageId="5" pageNumber="396">B. kyanos</emphasis>
</taxonomicName>
in having fewer circumpeduncular scales (16–18 vs. 18–19). Furthermore,
<taxonomicName id="4C599BD3FFEEBD1CFEF5FB809BAB2831" box="[306,456,1087,1110]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEEBD1CFEF5FB809BAB2831" box="[306,456,1087,1110]" italics="true" pageId="5" pageNumber="396">B. autumnum</emphasis>
</taxonomicName>
differs from all remaining species by presence of a black caudal-fin margin, outlining entire fin. Additionally,
<taxonomicName id="4C599BD3FFEEBD1CFE7DFBDC9835281D" box="[442,598,1123,1146]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEEBD1CFE7DFBDC9835281D" box="[442,598,1123,1146]" italics="true" pageId="5" pageNumber="396">B. autumnum</emphasis>
</taxonomicName>
can be biometrically separated from
<taxonomicName id="4C599BD3FFEEBD1CFBE5FBDC9EA9281C" box="[1058,1226,1123,1147]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="andrewraoi">
<emphasis id="B92D3C42FFEEBD1CFBE5FBDC9EA9281C" box="[1058,1226,1123,1147]" italics="true" pageId="5" pageNumber="396">B. andrewraoi</emphasis>
</taxonomicName>
by having fewer circumpeduncular scales (16–18 vs. 18).
</paragraph>
</subSubSection>
<subSubSection id="C343B3DBFFEEBD11FF00FB159E6B2DCA" lastPageId="8" lastPageNumber="399" pageId="5" pageNumber="396" type="description">
<paragraph id="8BE6E050FFEEBD1CFF00FB1599F02934" blockId="5.[151,1437,258,1363]" pageId="5" pageNumber="396">
<emphasis id="B92D3C42FFEEBD1CFF00FB159B3A28A4" bold="true" box="[199,345,1194,1219]" pageId="5" pageNumber="396">Description.</emphasis>
General body shape as in
<figureCitation id="1362FCD5FFEEBD1CFD5EFB15994528A4" box="[665,806,1194,1219]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="5.[151,250,1950,1972]" captionStartId-1="6.[151,250,750,772]" captionTargetBox-0="[151,1435,1415,1929]" captionTargetBox-1="[244,1342,289,665]" captionTargetId-0="figure@5.[151,1435,1415,1929]" captionTargetId-1="figure@6.[150,1436,193,729]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 4. Badis autumnum, MSNG 57568, paratype, male, 35.9 mm SL; India: West Bengal, Singimari River drainage, Rathbari stream at Petla. Note the oblique stripe on operculum." captionText-1="FIGURE 5. Badis autumnum, MSNG 57570, paratype, female, 29.4 mm SL; India: West Bengal, Singimari River drainage, Rathbari stream at Petla. Note the blotch above base of opercular spine." httpUri-0="https://zenodo.org/record/243246/files/figure.png" httpUri-1="https://zenodo.org/record/243247/files/figure.png" pageId="5" pageNumber="396">Figures 4–5</figureCitation>
. Morphometric characters are listed in
<tableCitation id="C6DBD5EBFFEEBD1CFB31FB159F3328A4" box="[1270,1360,1194,1219]" captionStart="TABLE 2" captionStartId="7.[151,239,328,351]" captionTargetBox="[159,1320,410,1094]" captionTargetPageId="7" captionText="TABLE 2. Morphometric characters of holotype (MSNG 57567) and seven paratypes (MSNG 57568, MSNG 57569, MSNG 57570) of Badis autumnum, new species, expressed as a percentage of standard length (SL) or head length (HL)." httpUri="http://table.plazi.org/id/DF26B0D8FFECBD1EFF50FEF79FF82D18" pageId="5" pageNumber="396" tableUuid="DF26B0D8FFECBD1EFF50FEF79FF82D18">Table 2</tableCitation>
. Body moderately elongate and laterally compressed. Snout and head rounded in lateral aspect. Orbit situated in anterior half of head and upper mid-axis of body. Opercular spine triangular, with a single tip. Predorsal contour concave, in smaller specimens straighter. Dorsal-fin base slightly concave. Caudal peduncle nearly straight, longer than deep. Abdominal contour concave in males, somewhat convex in females.
</paragraph>
<caption id="DF26B0D8FFEEBD1CFF50F82199742BB4" httpUri="https://zenodo.org/record/243246/files/figure.png" pageId="5" pageNumber="396" targetBox="[151,1435,1415,1929]" targetPageId="5">
<paragraph id="8BE6E050FFEEBD1CFF50F82199742BB4" blockId="5.[151,1436,1950,2003]" pageId="5" pageNumber="396">
<emphasis id="B92D3C42FFEEBD1CFF50F8219B762BD3" bold="true" box="[151,277,1950,1972]" pageId="5" pageNumber="396">FIGURE 4.</emphasis>
<taxonomicName id="4C599BD3FFEEBD1CFED8F8219BAE2BD2" box="[287,461,1950,1973]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="396" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEEBD1CFED8F8219BAE2BD2" box="[287,461,1950,1973]" italics="true" pageId="5" pageNumber="396">Badis autumnum</emphasis>
</taxonomicName>
, MSNG 57568, paratype, male, 35.9 mm SL; India: West Bengal, Singimari River drainage, Rathbari stream at Petla. Note the oblique stripe on operculum.
</paragraph>
</caption>
<caption id="DF26B0D8FFEDBD1FFF50FD51990F2F45" httpUri="https://zenodo.org/record/243247/files/figure.png" pageId="6" pageNumber="397" targetBox="[244,1342,289,665]" targetPageId="6">
<paragraph id="8BE6E050FFEDBD1FFF50FD51990F2F45" blockId="6.[151,1436,750,802]" pageId="6" pageNumber="397">
<emphasis id="B92D3C42FFEDBD1FFF50FD519B772F64" bold="true" box="[151,276,750,772]" pageId="6" pageNumber="397">FIGURE 5.</emphasis>
<taxonomicName id="4C599BD3FFEDBD1FFEDBFD519BAA2F63" box="[284,457,750,772]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="397" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEDBD1FFEDBFD519BAA2F63" box="[284,457,750,772]" italics="true" pageId="6" pageNumber="397">Badis autumnum</emphasis>
</taxonomicName>
, MSNG 57570, paratype, female, 29.4 mm SL; India: West Bengal, Singimari River drainage, Rathbari stream at Petla. Note the blotch above base of opercular spine.
</paragraph>
</caption>
<caption id="DF26B0D8FFEDBD1FFF50F97D98FE2A90" httpUri="https://zenodo.org/record/243248/files/figure.png" pageId="6" pageNumber="397" targetBox="[151,1435,852,1708]" targetPageId="6">
<paragraph id="8BE6E050FFEDBD1FFF50F97D98FE2A90" blockId="6.[151,1436,1730,1783]" pageId="6" pageNumber="397">
<emphasis id="B92D3C42FFEDBD1FFF50F97D9B742AB0" bold="true" box="[151,279,1730,1752]" pageId="6" pageNumber="397">FIGURE 6.</emphasis>
Live male individual of
<taxonomicName id="4C599BD3FFEDBD1FFDEDF97D98BA2ABF" box="[554,729,1730,1752]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="397" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFEDBD1FFDEDF97D98BA2ABF" box="[554,729,1730,1752]" italics="true" pageId="6" pageNumber="397">Badis autumnum</emphasis>
</taxonomicName>
, ca 35 mm SL (not included in type series); India: West Bengal, Singimari River drainage, Rathbari stream at Petla.
</paragraph>
</caption>
<paragraph id="8BE6E050FFEDBD1FFF00F89F99FC2BE7" blockId="6.[151,1436,1823,2028]" pageId="6" pageNumber="397">Dentary pores 1–3, anguloarticular pores 1, preopercular pores 1–6, nasal pores 2, frontal pores 2–3–4, extrascapular pores 1–2–3, posttemporal opening pores 1–2, coronalis pore 1, lachrymal pores 1–3, infraorbital pores 1–4. Predorsal scales 4 anterior to coronalis pore, 7 posteriorly.</paragraph>
<paragraph id="8BE6E050FFEDBD1EFF00F8349E3F2CB3" blockId="6.[151,1436,1823,2028]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="398" pageId="6" pageNumber="397">
Scales on side strongly ctenoid, on top of head cycloid, scales on side of chest cycloid. Opercular scales ctenoid, preopercular and subopercular scales cycloid. Circumpeduncular scales 7–8 above and 7–8 below lateral line, totalling 16–18 scales. Scales in vertical row 1½–2 above and 6–7, below lateral line. Scales in lateral row 25– 27, modally 25, 26. Tubed scales in lateral line
<date id="FFE7C690FFECBD1EFD71FF28994C2CD7" box="[694,815,151,176]" pageId="7" pageNumber="398" value="1920-06-04" valueMax="1920-06-25" valueMin="1920-06-04">20/4–25/6.</date>
Dorsal-fin base scaled with 1 scale row anteriorly and 1½–2 posteriorly, anal-fin base with 1 scale row. Caudal-fin base with 3–5 scale rows.
</paragraph>
<paragraph id="8BE6E050FFECBD1EFF00FF6098202D7A" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="398">
Dorsal-fin rays XV+10–XVII+11, modally XVI+10. Anal-fin rays III+
<date id="FFE7C690FFECBD1EFC38FF5F9E5A2C9F" box="[1023,1081,224,248]" bridgedPair="-" pageId="7" pageNumber="398">7–III</date>
+9, modally III+8. Pectoral-fin rays 12–14, modally 13. Vertebrae 26.
</paragraph>
<caption id="DF26B0D8FFECBD1EFF50FEF79FF82D18" ID-Table-UUID="DF26B0D8FFECBD1EFF50FEF79FF82D18" httpUri="http://table.plazi.org/id/DF26B0D8FFECBD1EFF50FEF79FF82D18" pageId="7" pageNumber="398" targetBox="[159,1320,410,1094]" targetIsTable="true" targetPageId="7">
<paragraph id="8BE6E050FFECBD1EFF50FEF79FF82D18" blockId="7.[151,1435,328,383]" pageId="7" pageNumber="398">
<emphasis id="B92D3C42FFECBD1EFF50FEF79B682D38" bold="true" box="[151,267,328,351]" pageId="7" pageNumber="398">TABLE 2.</emphasis>
Morphometric characters of holotype (MSNG 57567) and seven paratypes (MSNG 57568, MSNG 57569, MSNG 57570) of
<taxonomicName id="4C599BD3FFECBD1EFE9EFED8986E2D18" box="[345,525,359,383]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="398" phylum="Chordata" rank="species" species="autumnum" status="sp. nov.">
<emphasis id="B92D3C42FFECBD1EFE9EFED8986E2D18" box="[345,525,359,383]" italics="true" pageId="7" pageNumber="398">Badis autumnum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21E8139FFECBD1EFDDCFED798FD2D18" box="[539,670,360,383]" pageId="7" pageNumber="398" rank="species">new species</taxonomicNameLabel>
, expressed as a percentage of standard length (SL) or head length (HL).
</paragraph>
</caption>
<paragraph id="8BE6E050FFECBD1EFD64FE259F6D2821" pageId="7" pageNumber="398">
<table id="F95912F0FFEC42E6FF58FE259F4B2821" box="[159,1320,410,1094]" gridcols="3" gridrows="15" pageId="7" pageNumber="398">
<tr id="3569E212FFEC42E6FF58FE259F4B2DD7" box="[159,1320,410,432]" gridrow="0" pageId="7" pageNumber="398" rowspan-0="1">
<th id="76B88B6EFFEC42E6FD64FE2599622DD7" box="[675,769,410,432]" gridcol="1" gridrow="0" pageId="7" pageNumber="398">Holotype</th>
<th id="76B88B6EFFEC42E6FC62FE259F4B2DD7" box="[933,1320,410,432]" gridcol="2" gridrow="0" pageId="7" pageNumber="398">Paratype range Mean ± S.D.</th>
</tr>
<tr id="3569E212FFEC42E6FF58FE7B9F4B2DBD" box="[159,1320,452,474]" gridrow="1" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FE7B9BD92DBD" box="[159,442,452,474]" gridcol="0" gridrow="1" pageId="7" pageNumber="398">Standard length (SL) in mm</th>
<td id="76B88B6EFFEC42E6FD64FE7B99622DBD" box="[675,769,452,474]" gridcol="1" gridrow="1" pageId="7" pageNumber="398">35.8</td>
<td id="76B88B6EFFEC42E6FC62FE7B9F4B2DBD" box="[933,1320,452,474]" gridcol="2" gridrow="1" pageId="7" pageNumber="398">27.5–35.9 31.6 ± 3.0</td>
</tr>
<tr id="3569E212FFEC42E6FF58FDA99F4B2E4B" box="[159,1320,534,556]" gridrow="2" pageId="7" pageNumber="398" rowspan-1="1" rowspan-2="1">
<th id="76B88B6EFFEC42E6FF58FDA99BD92E4B" box="[159,442,534,556]" gridcol="0" gridrow="2" pageId="7" pageNumber="398">As percentage of SL</th>
</tr>
<tr id="3569E212FFEC42E6FF58FDFF9F4B2E31" box="[159,1320,576,598]" gridrow="3" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FDFF9BD92E31" box="[159,442,576,598]" gridcol="0" gridrow="3" pageId="7" pageNumber="398">Head length</th>
<td id="76B88B6EFFEC42E6FD64FDFF99622E31" box="[675,769,576,598]" gridcol="1" gridrow="3" pageId="7" pageNumber="398">29.8</td>
<td id="76B88B6EFFEC42E6FC62FDFF9F4B2E31" box="[933,1320,576,598]" gridcol="2" gridrow="3" pageId="7" pageNumber="398">27.5–34.6 30.5 ± 2.0</td>
</tr>
<tr id="3569E212FFEC42E6FF58FDD69F4B2E18" box="[159,1320,617,639]" gridrow="4" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FDD69BD92E18" box="[159,442,617,639]" gridcol="0" gridrow="4" pageId="7" pageNumber="398">Snout length</th>
<td id="76B88B6EFFEC42E6FD64FDD699622E18" box="[675,769,617,639]" gridcol="1" gridrow="4" pageId="7" pageNumber="398">7.2</td>
<td id="76B88B6EFFEC42E6FC62FDD69F4B2E18" box="[933,1320,617,639]" gridcol="2" gridrow="4" pageId="7" pageNumber="398">5.5–8.5 6.4 ± 0.9</td>
</tr>
<tr id="3569E212FFEC42E6FF58FD2D9F4B2ECF" box="[159,1320,658,680]" gridrow="5" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FD2D9BD92ECF" box="[159,442,658,680]" gridcol="0" gridrow="5" pageId="7" pageNumber="398">Orbital diameter</th>
<td id="76B88B6EFFEC42E6FD64FD2D99622ECF" box="[675,769,658,680]" gridcol="1" gridrow="5" pageId="7" pageNumber="398">10.0</td>
<td id="76B88B6EFFEC42E6FC62FD2D9F4B2ECF" box="[933,1320,658,680]" gridcol="2" gridrow="5" pageId="7" pageNumber="398">7.2–10.0 9.0 ± 0.9</td>
</tr>
<tr id="3569E212FFEC42E6FF58FD039F4B2EB5" box="[159,1320,700,722]" gridrow="6" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FD039BD92EB5" box="[159,442,700,722]" gridcol="0" gridrow="6" pageId="7" pageNumber="398">Interorbital width</th>
<td id="76B88B6EFFEC42E6FD64FD0399622EB5" box="[675,769,700,722]" gridcol="1" gridrow="6" pageId="7" pageNumber="398">6.9</td>
<td id="76B88B6EFFEC42E6FC62FD039F4B2EB5" box="[933,1320,700,722]" gridcol="2" gridrow="6" pageId="7" pageNumber="398">4.3–8.1 6.3 ± 1.3</td>
</tr>
<tr id="3569E212FFEC42E6FF58FD5A9F4B2E9C" box="[159,1320,741,763]" gridrow="7" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FD5A9BD92E9C" box="[159,442,741,763]" gridcol="0" gridrow="7" pageId="7" pageNumber="398">Upper jaw length</th>
<td id="76B88B6EFFEC42E6FD64FD5A99622E9C" box="[675,769,741,763]" gridcol="1" gridrow="7" pageId="7" pageNumber="398">10.6</td>
<td id="76B88B6EFFEC42E6FC62FD5A9F4B2E9C" box="[933,1320,741,763]" gridcol="2" gridrow="7" pageId="7" pageNumber="398">7.5–10.8 10.0 ± 1.0</td>
</tr>
<tr id="3569E212FFEC42E6FF58FCB19F4B2F43" box="[159,1320,782,804]" gridrow="8" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FCB19BD92F43" box="[159,442,782,804]" gridcol="0" gridrow="8" pageId="7" pageNumber="398">Lower jaw length</th>
<td id="76B88B6EFFEC42E6FD64FCB199622F43" box="[675,769,782,804]" gridcol="1" gridrow="8" pageId="7" pageNumber="398">13.1</td>
<td id="76B88B6EFFEC42E6FC62FCB19F4B2F43" box="[933,1320,782,804]" gridcol="2" gridrow="8" pageId="7" pageNumber="398">10.8–13.6 12.3 ± 0.9</td>
</tr>
<tr id="3569E212FFEC42E6FF58FC879F4B2F29" box="[159,1320,824,846]" gridrow="9" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FC879BD92F29" box="[159,442,824,846]" gridcol="0" gridrow="9" pageId="7" pageNumber="398">Body depth</th>
<td id="76B88B6EFFEC42E6FD64FC8799622F29" box="[675,769,824,846]" gridcol="1" gridrow="9" pageId="7" pageNumber="398">29.8</td>
<td id="76B88B6EFFEC42E6FC62FC879F4B2F29" box="[933,1320,824,846]" gridcol="2" gridrow="9" pageId="7" pageNumber="398">27.6–33.6 29.6 ± 1.8</td>
</tr>
<tr id="3569E212FFEC42E6FF58FCDE9F4B2F10" box="[159,1320,865,887]" gridrow="10" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FCDE9BD92F10" box="[159,442,865,887]" gridcol="0" gridrow="10" pageId="7" pageNumber="398">Pelvic fin length</th>
<td id="76B88B6EFFEC42E6FD64FCDE99622F10" box="[675,769,865,887]" gridcol="1" gridrow="10" pageId="7" pageNumber="398">24.5</td>
<td id="76B88B6EFFEC42E6FC62FCDE9F4B2F10" box="[933,1320,865,887]" gridcol="2" gridrow="10" pageId="7" pageNumber="398">22.5–28.3 24.8 ± 2.3</td>
</tr>
<tr id="3569E212FFEC42E6FF58FC359F4B2FC7" box="[159,1320,906,928]" gridrow="11" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FC359BD92FC7" box="[159,442,906,928]" gridcol="0" gridrow="11" pageId="7" pageNumber="398">Pelvic to anal distance</th>
<td id="76B88B6EFFEC42E6FD64FC3599622FC7" box="[675,769,906,928]" gridcol="1" gridrow="11" pageId="7" pageNumber="398">30.7</td>
<td id="76B88B6EFFEC42E6FC62FC359F4B2FC7" box="[933,1320,906,928]" gridcol="2" gridrow="11" pageId="7" pageNumber="398">25.7–35.0 29.7 ± 2.7</td>
</tr>
<tr id="3569E212FFEC42E6FF58FC629F4B2F94" box="[159,1320,989,1011]" gridrow="12" pageId="7" pageNumber="398" rowspan-1="1" rowspan-2="1">
<th id="76B88B6EFFEC42E6FF58FC629BD92F94" box="[159,442,989,1011]" gridcol="0" gridrow="12" pageId="7" pageNumber="398">As percentage of HL</th>
</tr>
<tr id="3569E212FFEC42E6FF58FBB99F4B287B" box="[159,1320,1030,1052]" gridrow="13" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FBB99BD9287B" box="[159,442,1030,1052]" gridcol="0" gridrow="13" pageId="7" pageNumber="398">Snout length</th>
<td id="76B88B6EFFEC42E6FD64FBB99962287B" box="[675,769,1030,1052]" gridcol="1" gridrow="13" pageId="7" pageNumber="398">21.6</td>
<td id="76B88B6EFFEC42E6FC62FBB99F4B287B" box="[933,1320,1030,1052]" gridcol="2" gridrow="13" pageId="7" pageNumber="398">18.1–24.5 20.6 ± 2.3</td>
</tr>
<tr id="3569E212FFEC42E6FF58FB8F9F4B2821" box="[159,1320,1072,1094]" gridrow="14" pageId="7" pageNumber="398">
<th id="76B88B6EFFEC42E6FF58FB8F9BD92821" box="[159,442,1072,1094]" gridcol="0" gridrow="14" pageId="7" pageNumber="398">Orbital diameter</th>
<td id="76B88B6EFFEC42E6FD64FB8F99622821" box="[675,769,1072,1094]" gridcol="1" gridrow="14" pageId="7" pageNumber="398">30.0</td>
<td id="76B88B6EFFEC42E6FC62FB8F9F4B2821" box="[933,1320,1072,1094]" gridcol="2" gridrow="14" pageId="7" pageNumber="398">26.2–33.6 29.3 ± 2.7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE6E050FFECBD1EFF00FB359BF92A48" blockId="7.[151,1437,1162,2015]" pageId="7" pageNumber="398">
<emphasis id="B92D3C42FFECBD1EFF00FB359B3C28C4" bold="true" box="[199,351,1162,1187]" pageId="7" pageNumber="398">Colouration.</emphasis>
In preservative, base colour brown to dark brown. Side of head grey to brown-grey. Postorbital stripe, when discernible, dark grey-brown and crossing eye to lower jaw. Suborbital stripe grey-brown. A small, distinct dark blotch posterodorsally on opercle, above base of opercular spine, forming a short, downward and slightly oblique stripe in a single specimen. Oblique dark brown bar on opercle, more or less parallel to suborbital stripe. A conspicuous dark blotch on pectoral-fin base in subadult specimens, dark brown pigment in smaller specimens, no pigment in adult individuals. Five to six dark grey-brown bars on sides, more distinct posteriorly. Dark grey-brown caudal peduncle blotch. Dorsal, anal and caudal fins grey to grey-brown, darker between fin rays. Dorsal fin darker proximally with a narrow white margin and red-brown submarginal band, and dark blotches between fin spines along middle of fin, frequently forming a complete black band, distinct up to posterior soft part of fin, indistinct onwards. Soft portion of dorsal fin with a dark, arched band. Anal fin with a narrow white perimeter. Pectoral fin pale-hyaline, pelvic fin pale-hyaline, dusky medially. Caudal fin with a black margin, outlining the entire fin.
</paragraph>
<paragraph id="8BE6E050FFECBD11FF00F98499122DEF" blockId="7.[151,1437,1162,2015]" lastBlockId="8.[151,1437,151,1112]" lastPageId="8" lastPageNumber="399" pageId="7" pageNumber="398">
In life, male (
<figureCitation id="1362FCD5FFECBD1EFEB5F9859BDF2A34" box="[370,444,1594,1619]" captionStart="FIGURE 6" captionStartId="6.[151,250,1730,1752]" captionTargetBox="[151,1435,852,1708]" captionTargetId="figure@6.[151,1436,852,1708]" captionTargetPageId="6" captionText="FIGURE 6. Live male individual of Badis autumnum, ca 35 mm SL (not included in type series); India: West Bengal, Singimari River drainage, Rathbari stream at Petla." httpUri="https://zenodo.org/record/243248/files/figure.png" pageId="7" pageNumber="398">Fig. 6</figureCitation>
) colouration mood-dependent. Base colour yellow-brown. Dorsum of head yellow. Preorbital, postorbital and suborbital stripes black and prominent. Oblique dark brown bar on opercle, more or less parallel to suborbital stripe. A small distinct dark blotch posterodorsally on opercle, above the opercular spine base, intermittently forming a short downward and slightly oblique stripe in some individuals. A conspicuous dark blotch on pectoral-fin base in subadult specimens, dark brown pigment in smaller specimens, no pigment in adult individuals. Twelve vertical black bars on sides, six of which displaying as undulated lines on flanks, most prominent centrally and ventrally, two bars on nape. All bars complete less than infrequently. Often bars 1 to 4 prominent and complete, remaining bars fainter and strongly reduced, present only dorsolaterally. Bars 3–10 each corresponding with a conspicuous black blotch on scaly basal cover of dorsal fin. Bars 3 and 4 individually corresponding with a faint black blotch on scaly basal cover of anal fin. Distinct median black blotch on caudal peduncle, often connecting with bar 1. Bar 1 broader than remaining bars, often broader medially, displaying as a second, partially absorbed blotch. A number of scales, especially between bars, with orange spots, 1 scale wide and often an orange spot on the bar. All fins dark, dorsal, anal and caudal fins with orange tinge. Dorsal, anal and pelvic fins with a narrow, bright white marginal band. Dorsal fin with a narrow black submarginal band and dark blotches between fin spines along middle of fin, often forming a complete black band, distinct up to posterior soft part of fin, indistinct thereon. Soft portion with an additional dark, arched band. Anal fin with a broad, black submarginal band, often absorbing most of the orange colouration and a faint but dark band, approximately in the middle of the fin. Caudal fin with a black margin, outlining the entire fin and an additional dark, arched vertical band proximally. Female comparable to male specimens, however duller coloured.
</paragraph>
<paragraph id="8BE6E050FFE3BD11FF00FE2B9E6B2DCA" blockId="8.[151,1437,151,1112]" box="[199,1032,404,429]" pageId="8" pageNumber="399">
When stressed,
<taxonomicName id="4C599BD3FFE3BD11FEB0FE2B98572DCB" box="[375,564,404,428]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFE3BD11FEB0FE2B98572DCB" box="[375,564,404,428]" italics="true" pageId="8" pageNumber="399">Badis autumnum</emphasis>
</taxonomicName>
becomes very pale and nearly colourless.
</paragraph>
</subSubSection>
<subSubSection id="C343B3DBFFE3BD11FF00FE089E162ECF" pageId="8" pageNumber="399" type="distribution">
<paragraph id="8BE6E050FFE3BD11FF00FE089ECF2D93" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="399">
<emphasis id="B92D3C42FFE3BD11FF00FE089B032DB7" bold="true" box="[199,352,439,464]" pageId="8" pageNumber="399">Distribution.</emphasis>
<taxonomicName id="4C599BD3FFE3BD11FEADFE0798492DB7" box="[362,554,440,464]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFE3BD11FEADFE0798492DB7" box="[362,554,440,464]" italics="true" pageId="8" pageNumber="399">Badis autumnum</emphasis>
</taxonomicName>
is presently only known from the
<typeStatus id="54E25EF2FFE3BD11FC07FE0799932DB7" box="[960,1008,440,464]" pageId="8" pageNumber="399">type</typeStatus>
locality at Petla, near Dinhata town, Cooch Behar district, Singimari River drainage, Rathbari stream, West Bengal,
<collectingCountry id="F34EA0C0FFE3BD11FBD6FE639E282D92" box="[1041,1099,476,501]" name="India" pageId="8" pageNumber="399">India</collectingCountry>
(
<figureCitation id="1362FCD5FFE3BD11FB9DFE639EFC2D93" box="[1114,1183,476,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1969,1991]" captionTargetBox="[151,1436,1322,1953]" captionTargetId="figure@4.[151,1436,1322,1954]" captionTargetPageId="4" captionText="FIGURE 3. Type localities of Badis andrewraoi (inverted triangle), B. autumnum (square), B. kyanos (rhomboid) and B. soraya (star) in West Bengal, India." httpUri="https://zenodo.org/record/243245/files/figure.png" pageId="8" pageNumber="399">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BE6E050FFE3BD11FF00FE409F482E7F" blockId="8.[151,1437,151,1112]" box="[199,1323,511,536]" pageId="8" pageNumber="399">
<emphasis id="B92D3C42FFE3BD11FF00FE409BED2E7F" bold="true" box="[199,398,511,536]" pageId="8" pageNumber="399">Ecological notes.</emphasis>
<taxonomicName id="4C599BD3FFE3BD11FE52FDBF98312E7F" box="[405,594,512,536]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFE3BD11FE52FDBF98312E7F" box="[405,594,512,536]" italics="true" pageId="8" pageNumber="399">Badis autumnum</emphasis>
</taxonomicName>
was collected from a swamp-like area (A. A. Rao, pers. comm.).
</paragraph>
<paragraph id="8BE6E050FFE3BD11FF00FD9B9E162ECF" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="399">
<emphasis id="B92D3C42FFE3BD11FF00FD9B9BD82E5A" bold="true" box="[199,443,548,573]" pageId="8" pageNumber="399">Reproductive notes.</emphasis>
<taxonomicName id="4C599BD3FFE3BD11FE0EFD9B98EF2E5B" box="[457,652,548,572]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFE3BD11FE0EFD9B98EF2E5B" box="[457,652,548,572]" italics="true" pageId="8" pageNumber="399">Badis autumnum</emphasis>
</taxonomicName>
represents a cave (e.g. coconut shells, photo canisters) brooding species. Two pairs have been observed for fifteen months. The male individual guards and fans the eggs and guards a small area surrounding the cave. The eggs are almost completely transparent and adhere to the floor, sides and roof of the cave. Paternal care ends as soon as the larvae reach the free-swimming stage.
</paragraph>
</subSubSection>
<subSubSection id="C343B3DBFFE3BD11FF00FD0B9A882F72" pageId="8" pageNumber="399" type="etymology">
<paragraph id="8BE6E050FFE3BD11FF00FD0B9A882F72" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="399">
<emphasis id="B92D3C42FFE3BD11FF00FD0B9B2D2EAA" bold="true" box="[199,334,692,717]" pageId="8" pageNumber="399">Etymology.</emphasis>
The Latin species epithet
<taxonomicName id="4C599BD3FFE3BD11FDBFFD0A988D2EAB" box="[632,750,693,716]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFE3BD11FDBFFD0A988D2EAB" box="[632,750,693,716]" italics="true" pageId="8" pageNumber="399">autumnum</emphasis>
</taxonomicName>
, a noun meaning autumn, is in allusion to numerous colours of autumn which the new species can display during various moods; combinations of brown, black, yellow and orange.
</paragraph>
</subSubSection>
<subSubSection id="C343B3DBFFE3BD11FF00FCA09E4D283F" pageId="8" pageNumber="399" type="discussion">
<paragraph id="8BE6E050FFE3BD11FF00FCA09B0E2F3B" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="399">
<emphasis id="B92D3C42FFE3BD11FF00FCA09B582F5F" bold="true" box="[199,315,799,824]" pageId="8" pageNumber="399">Remarks.</emphasis>
<taxonomicName id="4C599BD3FFE3BD11FE84FC9F98632F5F" box="[323,512,800,824]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="species" species="autumnum">
<emphasis id="B92D3C42FFE3BD11FE84FC9F98632F5F" box="[323,512,800,824]" italics="true" pageId="8" pageNumber="399">Badis autumnum</emphasis>
</taxonomicName>
was first introduced into the aquarium trade as ‘
<taxonomicName id="4C599BD3FFE3BD11FBE0FC9F9E042F5F" box="[1063,1127,800,824]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="399" phylum="Chordata" rank="genus">
<emphasis id="B92D3C42FFE3BD11FBE0FC9F9E042F5F" box="[1063,1127,800,824]" italics="true" pageId="8" pageNumber="399">Badis</emphasis>
</taxonomicName>
sp. (New) Torpedo’ (A. A. Rao, pers. comm.).
</paragraph>
<paragraph id="8BE6E050FFE3BD11FF00FCD89E4D283F" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="399">
The new species is, amongst additional traits, characterised by two autapomorphies; a small conspicuous dark blotch posterodorsally on the opercle above the base of the opercular spine and a conspicuous dark blotch on the pectoral-fin base in subadult specimens. The latter single occurrence apomorphy is only present in individuals ranging
<quantity id="4CA14DB5FFE3BD11FF32FC6B9BFA2F8B" box="[245,409,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.8299999999999996" metricValueMax="3.29" metricValueMin="2.37" pageId="8" pageNumber="399" unit="mm" value="28.299999999999997" valueMax="32.9" valueMin="23.7">23.7–32.9 mm</quantity>
SL, before completely disappearing in adults. Some individuals slightly smaller than
<quantity id="4CA14DB5FFE3BD11FAACFC6B9AA02868" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="8" pageNumber="399" unit="mm" value="32.9">32.9 mm</quantity>
SL had already lost the blotch. Specimens of smaller sizes only depict dark brown pigment, not unlike that on other parts of the body, at the pectoral-fin base. It is presently unknown why the blotch is absent in adult individuals, however obviously its function is no longer relevant in larger entities.
</paragraph>
</subSubSection>
</treatment>
</document>
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