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<document id="A7535FDA97E0D5142301A074F80436EC" ID-DOI="10.11646/zootaxa.5383.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350971" ID-ZooBank="1D2AD0FF-CE98-462C-A238-CC3DBF2906C9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702283818271" checkinUser="plazi" docAuthor="Lin, Chen, Zhang, Chunmin &amp; Yang, Ding" docDate="2023" docId="03F03213FFA4196D9AD6FF0264D546ED" docLanguage="en" docName="zootaxa.5383.2.3.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.3/52446" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus latimaculatus Lin &amp; Zhang &amp; Yang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="163" masterDocId="FFC94A6BFFAE19679A41FF95672F4308" masterDocTitle="The genus Systenus Meigen, 1803 (Diptera: Dolichopodidae) from Yunnan, China, with descriptions of six new species" masterLastPageNumber="166" masterPageNumber="153" pageNumber="163" updateTime="1702284527099" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2D56AC166A13D0FE8E4ECF4BB71C76E4">The genus Systenus Meigen, 1803 (Diptera: Dolichopodidae) from Yunnan, China, with descriptions of six new species</mods:title>
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<mods:namePart id="F935C28525DDB8921F10C17C122B3F62">Lin, Chen</mods:namePart>
<mods:affiliation id="E18F2C6E1956DA9E7BC254E50F9A6709">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China &amp; Institute of Life Science and Technology, Inner Mongolia Normal University, Huhhot 010022, China</mods:affiliation>
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<mods:namePart id="BE32CDA7DFD4AFFE0213AA55A97F92CD">Zhang, Chunmin</mods:namePart>
<mods:affiliation id="C658C872457F4329E092F75597CF9648">Tongliao Agricultural Technology Extension Center, Tongliao 028000, China</mods:affiliation>
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<mods:namePart id="34EC842E861E75C99802D1D8E4C4CEC8">Yang, Ding</mods:namePart>
<mods:affiliation id="FC2307E75B409AF0855B40BA8B531BE4">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China</mods:affiliation>
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<subSubSection id="C343D08EFFA4196D9AD6FF02656D43B9" box="[151,578,151,178]" pageId="10" pageNumber="163" type="nomenclature">
<paragraph id="8BE68305FFA4196D9AD6FF02656D43B9" blockId="10.[151,578,151,213]" box="[151,578,151,178]" pageId="10" pageNumber="163">
<heading id="D0AE3469FFA4196D9AD6FF02656D43B9" bold="true" box="[151,578,151,178]" fontSize="11" level="1" pageId="10" pageNumber="163" reason="1">
<emphasis id="B92D5F17FFA4196D9AD6FF02656D43B9" bold="true" box="[151,578,151,178]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9AD6FF0266F243BA" bold="true" box="[151,477,151,178]" italics="true" pageId="10" pageNumber="163">
7.
<taxonomicName id="4C59F886FFA4196D9A86FF0266F243BA" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[199,477,151,178]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="163" phylum="Arthropoda" rank="species" species="latimaculatus" status="sp. nov.">Systenus latimaculatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A21EE26CFFA4196D9BA5FF02656D43B9" box="[484,578,151,177]" pageId="10" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9AD6FF2E666C43DD" box="[151,323,187,213]" pageId="10" pageNumber="163" type="description">
<paragraph id="8BE68305FFA4196D9AD6FF2E666C43DD" blockId="10.[151,578,151,213]" box="[151,323,187,213]" pageId="10" pageNumber="163">
(
<figureCitation id="13629F80FFA4196D9ADEFF2E67C743DD" box="[159,232,187,213]" captionStart="FIGURES 16. 1" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[203,1384,189,1907]" captionTargetId="figure-21@2.[193,1394,179,1917]" captionTargetPageId="2" captionText="FIGURES 16. 1. Systenus apicidigitatus sp. nov.; 2. Systenus basiflavus sp. nov.; 3. Systenus digitatus sp. nov.; 4. Systenus flavilateralis sp. nov.; 5. Systenus huanglianus sp. nov.; 6. Systenus latimaculatus sp. nov. All male. Scale bar =1.0mm." figureDoi="http://doi.org/10.5281/zenodo.10350973" httpUri="https://zenodo.org/record/10350973/files/figure.png" pageId="10" pageNumber="163">Figs 6</figureCitation>
, 1718)
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9AD6FE96634E4281" pageId="10" pageNumber="163" type="diagnosis">
<paragraph id="8BE68305FFA4196D9AD6FE96634E4281" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9AD6FE9666254215" bold="true" box="[151,266,259,285]" pageId="10" pageNumber="163">Diagnosis</emphasis>
. Postpedicel 3.2 times longer than wide. Wing with wide grayish brown apical band. R
<subScript id="17DD8140FFA4196D9E93FE8063C3422B" attach="left" box="[1234,1260,277,291]" fontSize="6" pageId="10" pageNumber="163">4+5</subScript>
and M
<subScript id="17DD8140FFA4196D9F7BFE83626D422C" attach="left" box="[1338,1346,278,292]" fontSize="6" pageId="10" pageNumber="163">1</subScript>
parallel apically. Fore coxa yellow, mid and hind coxae dark brown. Hind femur narrowly brown near extreme tip. Mid tibia brown at narrow tip, hind tibia dark brown at apical 1/3. Abdominal segment 7 petiolate. Surstylus large and broad, apically distinctly widened, and with a long finger-like dorsal process directed inward.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9A86FE06668D47E1" pageId="10" pageNumber="163" type="description">
<paragraph id="8BE68305FFA4196D9A86FE0664D342A6" blockId="10.[151,1437,259,1510]" box="[199,1020,403,430]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9A86FE06667642A5" bold="true" box="[199,345,403,429]" pageId="10" pageNumber="163">Description.</emphasis>
Male (
<figureCitation id="13629F80FFA4196D9BE9FE0666C242A5" box="[424,493,403,429]" captionStart="FIGURES 16. 1" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[203,1384,189,1907]" captionTargetId="figure-21@2.[193,1394,179,1917]" captionTargetPageId="2" captionText="FIGURES 16. 1. Systenus apicidigitatus sp. nov.; 2. Systenus basiflavus sp. nov.; 3. Systenus digitatus sp. nov.; 4. Systenus flavilateralis sp. nov.; 5. Systenus huanglianus sp. nov.; 6. Systenus latimaculatus sp. nov. All male. Scale bar =1.0mm." figureDoi="http://doi.org/10.5281/zenodo.10350973" httpUri="https://zenodo.org/record/10350973/files/figure.png" pageId="10" pageNumber="163">Fig. 6</figureCitation>
). Body length
<quantity id="4CA12EE0FFA4196D98D2FE0665D642A6" box="[659,761,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="10" pageNumber="163" unit="mm" value="3.25">3.25 mm</quantity>
, wing length
<quantity id="4CA12EE0FFA4196D99D1FE0664D842A6" box="[912,1015,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.13" pageId="10" pageNumber="163" unit="mm" value="3.13">3.13 mm</quantity>
.
</paragraph>
<paragraph id="8BE68305FFA4196D9A86FE2263DC418D" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">Head metallic green with pale gray pollinosity. Frons metallic green with white gray pollinosity; face dark metallic green with dense silvery white pollinosity, slightly wider than width of postpedicel. Hairs and bristles on head black, but lower postocular bristles yellowish. 2 oc, 2 vt, 2 pvt. Antenna [apical segment of arista broken] black; pedicel with blackish apical bristles; postpedicel 3.2 times longer than wide, basally rather thick, apically long finger-like, and with dense dark brown pubescence; arista apical (Fig. 17). Proboscis short, dark brown with short blackish hairs; palpus small, blackish, with black hairs and 1 slightly long black apical bristle.</paragraph>
<paragraph id="8BE68305FFA4196D9A86FD1A668841F9" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">Thorax metallic green with pale gray pollinosity. Hairs and bristles on thorax black; 6 dc, 1314 irregularly paired acr; 2 strong npl, 1 strong sa, 2 strong pa; sc long and strong, lsc short and hair-like. Propleuron with 1 brown bristle on lower portion.</paragraph>
<paragraph id="8BE68305FFA4196D9A86FD69665740AD" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">
Legs [mid and hind tarsi broken] mainly yellow; fore coxa yellow, mid and hind coxae dark brown; hind femur narrowly brown near extreme tip; mid tibia brown at narrow tip, hind tibia dark brown at apical 1/3; fore tarsomere 5 brown. Hairs and bristles on legs black except those on coxae yellow. Mid tibia with
<specimenCount id="9D5F488CFFA4196D9E17FCD663A44055" box="[1110,1163,835,861]" count="1" pageId="10" pageNumber="163" type="generic">1 ad.</specimenCount>
Relative length of tibiae and five tarsomeres of legs LI: 4.2: 1.9: 1.7: 1.0: 0.6: 0.4; LII: 5.7: distal podomeres missing; LIII: 6.9: distal podomeres missing.
</paragraph>
<paragraph id="8BE68305FFA4196D9A86FC3A622140E5" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">
Wing nearly hyaline with wide grayish brown apical band; veins dark brown, R
<subScript id="17DD8140FFA4196D9E08FC54634C40C7" attach="left" box="[1097,1123,961,975]" fontSize="6" pageId="10" pageNumber="163">4+5</subScript>
and M
<subScript id="17DD8140FFA4196D9EF2FC57639440D8" attach="left" box="[1203,1211,962,976]" fontSize="6" pageId="10" pageNumber="163">1</subScript>
distinctly separated and parallel apically; CuAx ratio 0.5. Squama yellow, with fan of pale white hairs. Halter pale yellow.
</paragraph>
<paragraph id="8BE68305FFA4196D9A86FC626656473D" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">Abdomen dark metallic green with pale gray pollinosity. Hairs and bristles on abdomen black. Abdominal segment 7 petiolate.</paragraph>
<paragraph id="8BE68305FFA4196D9A86FBAA649A47CD" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">Male genitalia (Fig. 18): epandrium distinctly longer than wide; epandrial lobe basally thick, distinctly narrowed toward tip, short hook-like at extreme tip; surstylus large and broad, apically distinctly widened with nearly truncate apical margin, and with long finger-like dorsal process directed inward. Cercus basally rather wide, apically finger-like. Hypandrium long spine-like, slightly curved. Pallus long and thin.</paragraph>
<paragraph id="8BE68305FFA4196D9A86FB5A668D47E1" blockId="10.[151,1437,259,1510]" box="[199,418,1231,1257]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9A86FB5A660A47E1" bold="true" box="[199,293,1231,1257]" pageId="10" pageNumber="163">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9A86FB6663734639" pageId="10" pageNumber="163" type="materials_examined">
<paragraph id="8BE68305FFA4196D9A86FB6663734639" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9A86FB66665E4605" bold="true" box="[199,369,1267,1293]" pageId="10" pageNumber="163">Type material.</emphasis>
<materialsCitation id="3B318958FFA4196D9B32FB6663734639" collectingDate="2019-04-19" collectingDateMax="2019-05-19" collectingDateMin="2019-04-19" collectionCode="CAU" collectorName="Malaise &amp; Liang Wang &amp; Xin Li" country="China" county="Lvchun" elevation="1779" latitude="23.057" location="Shuikuxiafang" longLatPrecision="1" longitude="102.777145" municipality="Yakou" pageId="10" pageNumber="163" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B92D5F17FFA4196D9B32FB6665364605" bold="true" box="[371,537,1267,1293]" pageId="10" pageNumber="163">
<typeStatus id="54E23DA7FFA4196D9B32FB6665364605" box="[371,537,1267,1293]" pageId="10" pageNumber="163" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="9D5F488CFFA4196D985DFB61651C4606" box="[540,563,1268,1294]" count="1" pageId="10" pageNumber="163" type="male"></specimenCount>
,
<collectingCountry id="F34EC395FFA4196D987AFB6665AD4605" box="[571,642,1267,1293]" name="China" pageId="10" pageNumber="163">China</collectingCountry>
:
<collectingRegion id="499D4DE7FFA4196D98C6FB6165CF4606" box="[647,736,1268,1294]" country="China" name="Yunnan" pageId="10" pageNumber="163">Yunnan</collectingRegion>
,
<collectingCounty id="6287FB89FFA4196D98A7FB6664114605" box="[742,830,1267,1293]" pageId="10" pageNumber="163">Lvchun</collectingCounty>
,
<collectingMunicipality id="6B82197FFFA4196D9905FB6664A04605" box="[836,911,1267,1293]" pageId="10" pageNumber="163">Yakou</collectingMunicipality>
,
<location id="8E86D5DEFFA4196D99D4FB6663174605" LSID="urn:lsid:plazi:treatment:03F03213FFA4196D9AD6FF0264D546ED:8E86D5DEFFA4196D99D4FB6663174605" box="[917,1080,1267,1293]" country="China" county="Lvchun" latitude="23.057" longLatPrecision="1" longitude="102.777145" municipality="Yakou" name="Shuikuxiafang" pageId="10" pageNumber="163" stateProvince="Yunnan">Shuikuxiafang</location>
[
<geoCoordinate id="EE6DE5C2FFA4196D9E03FB6663F84606" box="[1090,1239,1267,1294]" degrees="23" direction="north" minutes="3" orientation="latitude" pageId="10" pageNumber="163" precision="1" seconds="25.2" value="23.057">23°325.2”N</geoCoordinate>
,
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,
<quantity id="4CA12EE0FFA4196D9AD6FA8267DF463A" box="[151,240,1303,1330]" metricMagnitude="3" metricUnit="m" metricValue="1.779" pageId="10" pageNumber="163" unit="m" value="1779.0">
<elevation id="00746436FFA4196D9AD6FA8267DF463A" box="[151,240,1303,1330]" metricMagnitude="3" metricUnit="m" metricValue="1.779" pageId="10" pageNumber="163" unit="m" value="1779.0">1779 m</elevation>
</quantity>
], collected by
<collectorName id="26ACE6D3FFA4196D9BD3FA8266C44639" box="[402,491,1303,1329]" pageId="10" pageNumber="163">Malaise</collectorName>
trap,
<date id="FFE7A5C5FFA4196D986DFA8265C34639" box="[556,748,1303,1329]" pageId="10" pageNumber="163" value="2019-04-19" valueMax="2019-05-19" valueMin="2019-04-19">
<collectingDate id="EFA35C2DFFA4196D986DFA8265C34639" box="[556,748,1303,1329]" pageId="10" pageNumber="163" value="2019-04-19" valueMax="2019-05-19" valueMin="2019-04-19">2019.IV.19V.19</collectingDate>
</date>
,
<collectorName id="26ACE6D3FFA4196D98B6FA8264AF463A" box="[759,896,1303,1330]" pageId="10" pageNumber="163">Liang Wang</collectorName>
and
<collectorName id="26ACE6D3FFA4196D99F7FA82632F4639" box="[950,1024,1303,1329]" pageId="10" pageNumber="163">Xin Li</collectorName>
(
<collectionCode id="ED481BC0FFA4196D9E4EFA82637F4639" box="[1039,1104,1303,1329]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="10" pageNumber="163">CAU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9A86FAAE650E465D" box="[199,545,1339,1365]" pageId="10" pageNumber="163" type="distribution">
<paragraph id="8BE68305FFA4196D9A86FAAE650E465D" blockId="10.[151,1437,259,1510]" box="[199,545,1339,1365]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9A86FAAE664E465D" bold="true" box="[199,353,1339,1365]" pageId="10" pageNumber="163">Distribution.</emphasis>
<collectingCountry id="F34EC395FFA4196D9B26FAAE6685465D" box="[359,426,1339,1365]" name="China" pageId="10" pageNumber="163">China</collectingCountry>
(
<collectingRegion id="499D4DE7FFA4196D9BF8FAAE653A465D" box="[441,533,1339,1365]" country="China" name="Yunnan" pageId="10" pageNumber="163">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9A86FACA634E46C9" pageId="10" pageNumber="163" type="discussion">
<paragraph id="8BE68305FFA4196D9A86FACA634E46C9" blockId="10.[151,1437,259,1510]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9A86FACA66144671" bold="true" box="[199,315,1375,1401]" pageId="10" pageNumber="163">Remarks.</emphasis>
<taxonomicName id="4C59F886FFA4196D9B04FACA65674671" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[325,584,1375,1401]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="163" phylum="Arthropoda" rank="species" species="latimaculatus" status="sp. nov.">
<emphasis id="B92D5F17FFA4196D9B04FACA65674671" box="[325,584,1375,1401]" italics="true" pageId="10" pageNumber="163">Systenus latimaculatus</emphasis>
</taxonomicName>
<emphasis id="B92D5F17FFA4196D9813FACA65804671" bold="true" box="[594,687,1375,1401]" pageId="10" pageNumber="163">
<taxonomicNameLabel id="A21EE26CFFA4196D9813FACA65804671" box="[594,687,1375,1401]" pageId="10" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is peculiar and can be easily separated from other Asian known species of the genus by the following features: pregenital segments petiolate; wing with wide grayish brown apical band; R
<subScript id="17DD8140FFA4196D9AAEFA2C662646CF" attach="left" box="[239,265,1465,1479]" fontSize="6" pageId="10" pageNumber="163">4+5</subScript>
and M
<subScript id="17DD8140FFA4196D9B19FA2F664F46C0" attach="left" box="[344,352,1466,1480]" fontSize="6" pageId="10" pageNumber="163">1</subScript>
parallel apically (
<bibRefCitation id="EFC8FEF4FFA4196D9868FA32645446C9" author="Olejnicek, J. &amp; Kozanek, M." box="[553,891,1447,1473]" pageId="10" pageNumber="163" pagination="127 - 129" refId="ref8730" refString="Olejnicek, J. &amp; Kozanek, M. (1997) Systenus slovaki sp. nov. (Diptera, Dolichopodidae) from Korea. Entomological Problems, 28 (2), 127 - 129." type="journal article" year="1997">Olejnícek and Kozánek, 1997</bibRefCitation>
;
<bibRefCitation id="EFC8FEF4FFA4196D99C6FA32637A46C9" author="Lin, C. &amp; Yang, D." box="[903,1109,1447,1474]" pageId="10" pageNumber="163" pagination="293 - 295" refId="ref8282" refString="Lin, C. &amp; Yang, D. (2022) One new species of Systenus Loew found in the Oriental region (Diptera: Empidoidea: Dolichopodidae). Journal of Asia-Pacific Biodiversity, 15, 293 - 295. https: // doi. org / 10.1016 / j. japb. 2021.12.004" type="journal article" year="2022">Lin &amp; Yang, 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA4196D9A86FA5E64D546ED" box="[199,1018,1483,1510]" pageId="10" pageNumber="163" type="etymology">
<paragraph id="8BE68305FFA4196D9A86FA5E64D546ED" blockId="10.[151,1437,259,1510]" box="[199,1018,1483,1510]" pageId="10" pageNumber="163">
<emphasis id="B92D5F17FFA4196D9A86FA5E666146ED" bold="true" box="[199,334,1483,1509]" pageId="10" pageNumber="163">Etymology.</emphasis>
The specific name refers to the wide apical spot of the wing.
</paragraph>
</subSubSection>
</treatment>
</document>