283 lines
32 KiB
XML
283 lines
32 KiB
XML
<document id="B3806B3C55D530C2A8A19785C30353AA" ID-DOI="10.5281/zenodo.184516" ID-GBIF-Dataset="62a45f81-bcff-449c-82d8-fef278006ea8" ID-ISSN="1175-5326" ID-Zenodo-Dep="184516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459985733509" checkinUser="plazi" docAuthor="Artal, Pedro, Guinot, Danièle, Bakel, Barry Van & Castillo, Juan" docDate="2008" docId="03E5D65BEF3C3F6D079CF9C8FD0DD6F7" docLanguage="en" docName="zt01907p027.pdf" docOrigin="Zootaxa 1907" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ibericancridae" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFDCAE23EF3A3F6A070BFF97FFB5D115" masterDocTitle="Ibericancridae, a new dakoticancroid family (Decapoda, Brachyura, Podotremata) from the upper Campanian (Upper Cretaceous) of Spain" masterLastPageNumber="27" masterPageNumber="1" pageNumber="7" updateTime="1698232278591" updateUser="plazi">
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<mods:title id="B4F9130F6111C1D4FC20B7F48CBA1A2C">Ibericancridae, a new dakoticancroid family (Decapoda, Brachyura, Podotremata) from the upper Campanian (Upper Cretaceous) of Spain</mods:title>
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<mods:namePart id="72420D39498664E9D05260C78B99BAF0">Artal, Pedro</mods:namePart>
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<mods:roleTerm id="AB218583ED31008ED4FD5A8BCF828D24">Author</mods:roleTerm>
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<mods:namePart id="CEE1F46F3D6C6FB7A97656B96B59ABB7">Guinot, Danièle</mods:namePart>
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<mods:namePart id="F401478DDA139C306E2CEEDF6F9D7484">Bakel, Barry Van</mods:namePart>
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<mods:namePart id="91C5288724E018822E3F0995E07B063C">Castillo, Juan</mods:namePart>
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<mods:title id="791DBA4029BF3639401D3B6EE582F9EB">Zootaxa</mods:title>
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<mods:part id="7FAFAD6823E3F5928DC24D08CAF46812">
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<mods:date id="1DB510986D95D828C70D9263E8821A25">2008</mods:date>
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<mods:number id="2AA1CF5916865264A352572BADC29902">1907</mods:number>
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<mods:classification id="DA6E6A04536376B12801D8A864DE2E9B">journal article</mods:classification>
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<mods:identifier id="CAB20828A5738D1A0098E0C7CBB9486D" type="DOI">10.5281/zenodo.184516</mods:identifier>
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<mods:identifier id="68FB8CD718433395DB61B0D1750E21BC" type="Zenodo-Dep">184516</mods:identifier>
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<treatment id="03E5D65BEF3C3F6D079CF9C8FD0DD6F7" ID-DOI="http://doi.org/10.5281/zenodo.6233904" ID-GBIF-Taxon="119379215" ID-Zenodo-Dep="6233904" LSID="urn:lsid:plazi:treatment:03E5D65BEF3C3F6D079CF9C8FD0DD6F7" httpUri="http://treatment.plazi.org/id/03E5D65BEF3C3F6D079CF9C8FD0DD6F7" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection id="C35634C6EF3C3F6C079CF9C8FE18D76C" box="[151,429,1631,1657]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="8BF3674DEF3C3F6C079CF9C8FE18D76C" blockId="6.[151,429,1631,1657]" box="[151,429,1631,1657]" pageId="6" pageNumber="7">
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<heading id="D0BBD021EF3C3F6C079CF9C8FE18D76C" bold="true" box="[151,429,1631,1657]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<emphasis id="B938BB5FEF3C3F6C079CF9C8FE18D76C" bold="true" box="[151,429,1631,1657]" pageId="6" pageNumber="7">
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<taxonomicName id="4C4C1CCEEF3C3F6C079CF9C8FEFAD76C" ID-CoL="629869d8-410d-4b3e-8aea-a8f9c635e382" box="[151,335,1631,1657]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
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n. fam.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF3C3F6C079CF93DFD78D7D4" box="[151,717,1706,1730]" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="8BF3674DEF3C3F6C079CF93DFD78D7D4" blockId="6.[151,717,1706,1730]" box="[151,717,1706,1730]" pageId="6" pageNumber="7">
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<emphasis id="B938BB5FEF3C3F6C079CF93DFEA9D7D7" bold="true" box="[151,284,1706,1730]" pageId="6" pageNumber="7">
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<typeStatus id="54F7D9EFEF3C3F6C079CF93DFF78D7D7" box="[151,205,1706,1730]" pageId="6" pageNumber="7">Type</typeStatus>
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genus.
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</emphasis>
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<taxonomicName id="4C4C1CCEEF3C3F6C062FF93DFE15D7D4" box="[292,416,1706,1729]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B938BB5FEF3C3F6C062FF93DFE15D7D4" box="[292,416,1706,1729]" italics="true" pageId="6" pageNumber="7">Ibericancer</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B0624EF3C3F6C06ACF93DFE47D7D7" box="[423,498,1706,1730]" pageId="6" pageNumber="7" rank="genus">
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<emphasis id="B938BB5FEF3C3F6C06ACF93DFE47D7D7" bold="true" box="[423,498,1706,1730]" pageId="6" pageNumber="7">n. gen.</emphasis>
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</taxonomicNameLabel>
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(gender masculine).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF3C3F6D079CF965FD0ED0DF" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
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<paragraph id="8BF3674DEF3C3F6D079CF965FD0ED0DF" blockId="6.[151,1437,1778,2004]" lastBlockId="7.[151,1437,152,2018]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<emphasis id="B938BB5FEF3C3F6C079CF965FEA2D619" bold="true" box="[151,279,1778,1804]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
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Large-sized crabs; carapace suboval, thick, with high flanks; fronto-orbital margin gently salient; front relatively broad, protuberant at inner-orbital angles, somewhat concave from dorsal view, noticeably notched medially; rostrum narrow, strongly downturned, axially grooved, bilobed; orbits small, directed forwards, with deep fissure and supramarginal tooth; dorsal regions relatively inflated, poorly differentiated except for moderately deep branchiocardiac grooves, some depressions, scarcely pronounced swellings; posterior margin of carapace concave, rimmed, axially raised; pterygostomial region large; pleural suture at sides of carapace; thoracic sternum well developed, conspicuously elevated laterally, with strong pointed prominence on each side of the anterior portion of sternite 4; only relative narrow portion of sternum exposed laterally at level of P2–P4; sternum bent, narrowing posteriorly at level of P4, P5; sternite 5 relatively large, sternite 6 as narrow, elongated plate, sternite 7 very reduced, oblique, sternite 8 completely covered by abdomen, not exposed; abdomen weakly dimorphic sexually, relatively broad in males; locking mechanism of press button
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<typeStatus id="54F7D9EFEF3D3F6D063AFEF7FED2D06F" box="[305,359,352,378]" pageId="7" pageNumber="8">type</typeStatus>
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; male gonopore on P5 coxa; female gonopore on P3 coxa; spermathecae at extremities of suture 7/8; chelipeds homochelous, slender, with propodus longer than wide; P2, P3 extremely robust, long; P4, P5 markedly reduced, subequal, subdorsal.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35634C6EF3D3F6D07CDFE40FD0DD6F7" pageId="7" pageNumber="8" type="discussion">
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<paragraph id="8BF3674DEF3D3F6D07CDFE40FC32D24F" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
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<emphasis id="B938BB5FEF3D3F6D07CDFE40FE8AD0E4" bold="true" box="[198,319,471,497]" pageId="7" pageNumber="8">Remarks.</emphasis>
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There are many differences between
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<taxonomicName id="4C4C1CCEEF3D3F6D05F7FE4FFC16D0E7" box="[764,931,472,498]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
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<emphasis id="B938BB5FEF3D3F6D04A7FE40FBB6D0E4" bold="true" box="[940,1027,471,497]" pageId="7" pageNumber="8">n. fam.</emphasis>
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and the
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<taxonomicName id="4C4C1CCEEF3D3F6D0362FE4FFF60D30F" authority="Rathbun, 1917" authorityName="Rathbun" authorityYear="1917" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">
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Dakoticancridae
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<bibRefCitation id="EFDD1ABCEF3D3F6D023AFE4FFF60D30F" author="Rathbun" pageId="7" pageNumber="26" refString="Rathbun, M. J. (1917) New species of South Dakota Cretaceous crabs. Proceedings of the United States National Museum, 52 (2182), 385 - 391." type="journal article" year="1917">Rathbun, 1917</bibRefCitation>
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</taxonomicName>
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. These differences mostly concern ventral characters but there are additional differences in the carapaces. In the
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<taxonomicName id="4C4C1CCEEF3D3F6D063EFDBFFE40D357" box="[309,501,552,578]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
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the deflected rostrum is directly connected to the orbital margins, and the orbitofrontal margin is broadly concave, with advanced outer orbital spines. In
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<taxonomicName id="4C4C1CCEEF3D3F6D0325FDC7FB02D37C" box="[1070,1207,592,617]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B938BB5FEF3D3F6D0325FDC7FB02D37C" box="[1070,1207,592,617]" italics="true" pageId="7" pageNumber="8">Ibericancer</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B0624EF3D3F6D03CAFDD8FAA0D37C" box="[1217,1301,591,617]" pageId="7" pageNumber="8" rank="genus">
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<emphasis id="B938BB5FEF3D3F6D03CAFDD8FAA0D37C" bold="true" box="[1217,1301,591,617]" pageId="7" pageNumber="8">n. gen.</emphasis>
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</taxonomicNameLabel>
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the frontal margin is more salient, somewhat concave in dorsal view, clearly notched medially, and the outer corners are lobed; the rostrum is strongly downturned, with a pronounced axial furrow. In the new genus the orbital margin is small in dorsal view, directed forwards, with the supraorbital tooth bounded by fissures. In contrast, in the
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<taxonomicName id="4C4C1CCEEF3D3F6D07CEFD67FE30D21F" box="[197,389,752,778]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
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the orbits are large and broadly concave, divided into two concave portions in dorsal view.
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<bibRefCitation id="EFDD1ABCEF3D3F6D07D7FC8FFE26D227" author="Bishop" box="[220,403,792,818]" pageId="7" pageNumber="25" refString="Bishop, G. A. (1984 b) Orbital bulla reticularis: A new orbital structure of a Cretaceous crab. Journal of Crustacean Biology, 4, 514 - 517." type="journal article" year="1984" yearSuffix="b">Bishop (1984b)</bibRefCitation>
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and
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<bibRefCitation id="EFDD1ABCEF3D3F6D06C7FC8FFD02D227" box="[460,695,792,818]" pageId="7" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article">
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Bishop
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<emphasis id="B938BB5FEF3D3F6D052CFC8FFDEFD224" box="[551,602,792,817]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
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. (1998)
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</bibRefCitation>
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provided a detailed description of the orbits and of a presumed sensory structure, ‘orbital bulla reticularis’ found in
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<taxonomicName id="4C4C1CCEEF3D3F6D05F7FCD7FC35D24C" box="[764,896,832,858]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="overana">
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<emphasis id="B938BB5FEF3D3F6D05F7FCD7FCA4D24C" box="[764,785,832,857]" italics="true" pageId="7" pageNumber="8">D</emphasis>
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.
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<emphasis id="B938BB5FEF3D3F6D042BFCD7FC35D24C" box="[800,896,832,857]" italics="true" pageId="7" pageNumber="8">overana</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BF3674DEF3D3F6D07CEFCFFFCBDD537" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
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In
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<taxonomicName id="4C4C1CCEEF3D3F6D07EDFCFFFEDAD294" box="[230,367,872,897]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B938BB5FEF3D3F6D07EDFCFFFEDAD294" box="[230,367,872,897]" italics="true" pageId="7" pageNumber="8">Ibericancer</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B0624EF3D3F6D0673FCF0FE7ED294" box="[376,459,871,897]" pageId="7" pageNumber="8" rank="genus">
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<emphasis id="B938BB5FEF3D3F6D0673FCF0FE7ED294" bold="true" box="[376,459,871,897]" pageId="7" pageNumber="8">n. gen.</emphasis>
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</taxonomicNameLabel>
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the posterior margin is concave, slightly convex in dorsal view through the raised posterior portion of the carapace, extremely thick, and medially inflated by the robust conical protuberance of the intestinal region; when seen from posterior view, it shows an unusual V-shape with the apex directed upwards. The posterior margin of the
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<taxonomicName id="4C4C1CCEEF3D3F6D055EFC77FCA0D2EF" box="[597,789,992,1018]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
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is nearly straight in dorsal view, thin, slightly rimmed, and there is no space in the dorso-ventral connection.
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</paragraph>
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<paragraph id="8BF3674DEF3D3F6D07CEFBA7FD63D42F" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
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The thoracic sternum of
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<taxonomicName id="4C4C1CCEEF3D3F6D06EEFBA7FDDBD55C" box="[485,622,1072,1097]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B938BB5FEF3D3F6D06EEFBA7FDDBD55C" box="[485,622,1072,1097]" italics="true" pageId="7" pageNumber="8">Ibericancer</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B0624EF3D3F6D057FFBB8FD73D55C" box="[628,710,1071,1097]" pageId="7" pageNumber="8" rank="genus">
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<emphasis id="B938BB5FEF3D3F6D057FFBB8FD73D55C" bold="true" box="[628,710,1071,1097]" pageId="7" pageNumber="8">n. gen.</emphasis>
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</taxonomicNameLabel>
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is narrower than in the
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<taxonomicName id="4C4C1CCEEF3D3F6D04D0FBA7FB2BD55F" box="[987,1182,1072,1098]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
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, elevated at the sides, stout, and with very thick cuticle; sternites 1–4 are smooth and continuous axially, while in
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<taxonomicName id="4C4C1CCEEF3D3F6D03D7FBCFFA29D567" box="[1244,1436,1112,1138]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
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the sternum is broader, the upper margin of sternite 3 is noticeably rimmed, marking a discontinuity of the anterior sternites 1, 2 (see
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<figureCitation id="13777BC8EF3D3F6D06C7FB3FFD9CD5D7" box="[460,553,1192,1218]" captionStart="FIGURE 3 A – E" captionStartId="5.[151,255,1871,1895]" captionTargetBox="[243,1344,221,1860]" captionTargetId="figure@5.[243,1345,211,1863]" captionTargetPageId="5" captionText="FIGURE 3 A – E. Casts of Dakoticancer overana; A. MNHN B 22902 - 1 (cast of USNM 173529 - 1), ventral view of a male; B. posterior view of the same specimen; C. details of the abdominal holding mechanism, the arrow indicating the press button protuberance on sternite 5; D. MNHM B 22902 - 2 (cast of USNM 173529 - 2), ventral view of a female, without the abdomen; the arrow indicating the aperture of the spermatheca; E. MNHN B 22903 (cast of USNM 173544), ventral view of a female; the arrow indicating the female gonopore on P 3 coxa." httpUri="https://zenodo.org/record/184518/files/figure.png" pageId="7" pageNumber="8">Fig. 3D</figureCitation>
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||
). The abdomen shows a weak sexual dimorphism, in contrast to the
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0241FB3FFEA6D5FF" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
. The supposed propodus of
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0550FB47FD49D5FC" box="[603,764,1232,1257]" class="Malacostraca" family="Dakoticancridae" genus="Tetracarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D0550FB47FD49D5FC" box="[603,764,1232,1257]" italics="true" pageId="7" pageNumber="8">Tetracarcinus</emphasis>
|
||
</taxonomicName>
|
||
figured by
|
||
<bibRefCitation id="EFDD1ABCEF3D3F6D048FFB47FBDCD5FF" author="Bishop" box="[900,1129,1232,1258]" pageId="7" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article" year="1998">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF3D3F6D04D5FB47FBACD5FC" box="[990,1049,1232,1257]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
(1998
|
||
</bibRefCitation>
|
||
: fig. 2.2) was noted as not having been found attached to the crab. This isolated propodus could be perfectly well assigned to a macruran, probably close to
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0663FAB7FD67D42F" authority="Kishinouye, 1926" authorityName="Kishinouye" authorityYear="1926" box="[360,722,1312,1338]" class="Malacostraca" family="Ctenochelidae" genus="Ctenocheles" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D0663FAB7FE42D42C" box="[360,503,1312,1337]" italics="true" pageId="7" pageNumber="8">Ctenocheles</emphasis>
|
||
Kishinouye, 1926
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF3D3F6D07CEFADFFAD8D4A7" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
|
||
The cheliped propodus in the
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D053AFADFFD6CD477" box="[561,729,1352,1378]" class="Malacostraca" family="Ibericancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Ibericancridae</taxonomicName>
|
||
<emphasis id="B938BB5FEF3D3F6D05EFFAD0FC8BD474" bold="true" box="[740,830,1351,1377]" pageId="7" pageNumber="8">n. fam.</emphasis>
|
||
is slender and somewhat dorsoventrally flattened, longer than wide, with very thin and long fingers. In dakoticancrids, the chelipeds are variable among the different species, varying from robust and globose in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D05E1FA0FFCC6D4A4" box="[746,883,1432,1458]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B938BB5FEF3D3F6D05E1FA0FFD4AD4A4" box="[746,767,1432,1457]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D0406FA0FFCC6D4A4" box="[781,883,1432,1457]" italics="true" pageId="7" pageNumber="8">australis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D04A6FA0FFBF4D4A4" box="[941,1089,1432,1457]" class="Malacostraca" family="Dakoticancridae" genus="Avitelmessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D04A6FA0FFBF4D4A4" box="[941,1089,1432,1457]" italics="true" pageId="7" pageNumber="8">Avitelmessus</emphasis>
|
||
</taxonomicName>
|
||
to slender in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D03E9FA0FFAD3D4A4" box="[1250,1382,1432,1458]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF3D3F6D03E9FA0FFB42D4A4" box="[1250,1271,1432,1457]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D020DFA0FFAD3D4A4" box="[1286,1382,1432,1457]" italics="true" pageId="7" pageNumber="8">overana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF3D3F6D07CEFA57FF0AD73F" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
|
||
Both P4 and P5 are subdorsally located, strongly reduced in thickness but weakly reduced in length, subequal, whereas in the presently known
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D056EFA7FFC90D717" box="[613,805,1512,1538]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
only the P5, when preserved, is reduced and subdorsal.
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF3D3F6D07CEF9AFFE49D654" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
|
||
The dorsal regions, somewhat less pronounced in the anterior portion of the carapace, are more uniformly inflated than in the four genera assigned to the family
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0417F9F7FC55D76F" box="[796,992,1632,1658]" class="Malacostraca" family="Dakoticancridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dakoticancridae</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D04E7F9F7FBC0D76C" box="[1004,1141,1632,1657]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF3D3F6D04E7F9F7FBC0D76C" box="[1004,1141,1632,1657]" italics="true" pageId="7" pageNumber="8">Ibericancer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B0624EF3D3F6D0377F9C8FB7AD76C" box="[1148,1231,1631,1657]" pageId="7" pageNumber="8" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D0377F9C8FB7AD76C" bold="true" box="[1148,1231,1631,1657]" pageId="7" pageNumber="8">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shows small and dense granules uniformly distributed on the carapace, thoracic sternum and pereiopods, in contrast to larger granules that tend to be grouped on the more prominent regions in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D04CFF927FBFFD7DC" box="[964,1098,1712,1738]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF3D3F6D04CFF927FC6CD7DC" box="[964,985,1712,1737]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D04E1F927FBFFD7DC" box="[1002,1098,1712,1737]" italics="true" pageId="7" pageNumber="8">overana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0350F927FB4ED7DC" box="[1115,1275,1712,1737]" class="Malacostraca" family="Dakoticancridae" genus="Tetracarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D0350F927FB4ED7DC" box="[1115,1275,1712,1737]" italics="true" pageId="7" pageNumber="8">Tetracarcinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0207F927FAD2D7DC" box="[1292,1383,1712,1737]" class="Malacostraca" family="Dakoticancridae" genus="Seorsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D0207F927FAD2D7DC" box="[1292,1383,1712,1737]" italics="true" pageId="7" pageNumber="8">Seorsus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B938BB5FEF3D3F6D079CF94FFE81D7E4" box="[151,308,1752,1777]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D079CF94FFE84D7E4" box="[151,305,1752,1777]" class="Malacostraca" family="Dakoticancridae" genus="Avitelmessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Avitelmessus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In the latter, there are spines on the carapace, lateral carapace margins and pereiopods (see
|
||
<bibRefCitation id="EFDD1ABCEF3D3F6D079CF897FE73D60F" author="Kesling" box="[151,454,1792,1818]" pageId="7" pageNumber="26" refString="Kesling, R. V. & Reimann, I. G. (1957) An Upper Cretaceous crab, Avitelmessus grapsoideus Rathbun. Contributions from the Museum of Paleontology, University of Michigan, 14, 1 - 15." type="journal article" year="1957">Kesling & Reimann 1957</bibRefCitation>
|
||
). The granulation over the entire surface is more distinct in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0371F897FAB1D60C" box="[1146,1284,1792,1818]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B938BB5FEF3D3F6D0371F897FB3AD60C" box="[1146,1167,1792,1817]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D0396F897FAB1D60C" box="[1181,1284,1792,1817]" italics="true" pageId="7" pageNumber="8">australis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D026AF897FEE4D654" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF3D3F6D026AF897FEE4D654" italics="true" pageId="7" pageNumber="8">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B0624EF3D3F6D0653F8B0FE1FD654" box="[344,426,1831,1857]" pageId="7" pageNumber="8" rank="genus">
|
||
<emphasis id="B938BB5FEF3D3F6D0653F8B0FE1FD654" bold="true" box="[344,426,1831,1857]" pageId="7" pageNumber="8">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B938BB5FEF3D3F6D06B2F8B0FE49D654" bold="true" box="[441,508,1831,1857]" pageId="7" pageNumber="8">n. sp.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF3674DEF3D3F6D07CEF8C7FD0DD6F7" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="8">
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D07CEF8C7FE04D67C" box="[197,433,1872,1897]" class="Malacostraca" family="Ibericancridae" genus="Ibericancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sanchoi" status="gen. nov.">
|
||
<emphasis id="B938BB5FEF3D3F6D07CEF8C7FE04D67C" box="[197,433,1872,1897]" italics="true" pageId="7" pageNumber="8">Ibericancer sanchoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938BB5FEF3D3F6D06B2F8D8FDEBD67C" bold="true" box="[441,606,1871,1897]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="A20B0624EF3D3F6D06B2F8D8FDB9D67C" box="[441,524,1871,1897]" pageId="7" pageNumber="8" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
is closer to
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D05E7F8C7FC5AD67C" box="[748,1007,1872,1897]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="autralis">
|
||
<emphasis id="B938BB5FEF3D3F6D05E7F8C7FC5AD67C" box="[748,1007,1872,1897]" italics="true" pageId="7" pageNumber="8">Dakoticancer autralis</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0344F8C7FB61D67C" box="[1103,1236,1872,1898]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF3D3F6D0344F8C7FBD1D67C" box="[1103,1124,1872,1897]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D037FF8C7FB61D67C" box="[1140,1236,1872,1897]" italics="true" pageId="7" pageNumber="8">overana</emphasis>
|
||
</taxonomicName>
|
||
by having a narrow and more quadrate carapace (more parallel sides in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0427F8EFFC03D684" box="[812,950,1912,1938]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B938BB5FEF3D3F6D0427F8EFFCF4D684" box="[812,833,1912,1937]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D045BF8EFFC03D684" box="[848,950,1912,1937]" italics="true" pageId="7" pageNumber="8">australis</emphasis>
|
||
</taxonomicName>
|
||
), prominent posterior portion (see key in
|
||
<bibRefCitation id="EFDD1ABCEF3D3F6D079CF837FEC2D6AF" author="Bishop" box="[151,375,1952,1978]" pageId="7" pageNumber="25" refString="Bishop, G. A., Feldmann, R. M. & Vega, F. (1998) The Dakoticancridae (Decapoda, Brachyura) from the Late Cretaceous of North America and Mexico. Contributions to Zoology, 67, 237 - 255." type="journal article" year="1998">
|
||
Bishop
|
||
<emphasis id="B938BB5FEF3D3F6D07F8F837FE9DD6AC" box="[243,296,1952,1977]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
: 246), narrower thoracic sternum, and more uniformly distributed dorsal granulation; the propodus is long and slender as in
|
||
<taxonomicName id="4C4C1CCEEF3D3F6D0526F85FFD04D6F4" box="[557,689,1992,2018]" class="Malacostraca" family="Dakoticancridae" genus="Dakoticancer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="overana">
|
||
<emphasis id="B938BB5FEF3D3F6D0526F85FFDF7D6F4" box="[557,578,1992,2017]" italics="true" pageId="7" pageNumber="8">D</emphasis>
|
||
.
|
||
<emphasis id="B938BB5FEF3D3F6D055AF85FFD04D6F4" box="[593,689,1992,2017]" italics="true" pageId="7" pageNumber="8">overana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |