742 lines
86 KiB
XML
742 lines
86 KiB
XML
<document id="9594B31A304B88C55B88BA7A079FB6E6" ID-CLB-Dataset="27352" ID-DOI="10.11646/zootaxa.4577.3.6" ID-GBIF-Dataset="5bf74913-8fc7-40fc-a1a5-40e57058fe24" ID-ISSN="1175-5326" ID-Zenodo-Dep="2632375" ID-ZooBank="806DD869-170C-4E2C-ABCF-5AD8040CC27D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554704699380" checkinUser="plazi" docAuthor="Vogel, Gernot & David, Patrick" docDate="2019" docId="47709B7C0522FFD049D366E2FA6BFBAC" docLanguage="en" docName="zootaxa.4577.3.6.pdf" docOrigin="Zootaxa 4577 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lycodon gibsonae Vogel & David 2019, spec. nov." docType="treatment" docVersion="7" lastPageNumber="522" masterDocId="BB49E3040521FFD74944670CFFF6FFBC" masterDocTitle="A new species of the Lycodon fasciatus complex from the Khorat Plateau, eastern Thailand (Reptiles, Squamata, Colubridae)" masterLastPageNumber="528" masterPageNumber="515" pageNumber="518" updateTime="1698719058041" updateUser="ExternalLinkService">
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<mods:mods id="CEC7C6C7A464E1F503D5E4148B15CB7E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AC7EDCBC05161BA7C6D6916CFE0CF6CC">
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<mods:title id="9152857D3C897F64FEBE7966013C15B4">A new species of the Lycodon fasciatus complex from the Khorat Plateau, eastern Thailand (Reptiles, Squamata, Colubridae)</mods:title>
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<mods:name id="EE4E3C55B4078CF00862AA5E3743821A" type="personal">
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<mods:roleTerm id="F861D1B1730A2B838BBCFCADED715540">Author</mods:roleTerm>
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<mods:namePart id="294CA5AF0F72BF55280B2E3DDB39BF52">Vogel, Gernot</mods:namePart>
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<mods:roleTerm id="FFA1D28EFB0A7B44D680497451A88510">Author</mods:roleTerm>
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<mods:namePart id="0330DC3C5B18C3748488BFA9308BBE0C">David, Patrick</mods:namePart>
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<mods:typeOfResource id="21C6EA2E084FD2741A6FA20AEEFAACA8">text</mods:typeOfResource>
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<mods:title id="9C4E84744E2B53425D9785D2FD4A6B2D">Zootaxa</mods:title>
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<mods:part id="C035D912FC345F6838A85E75436E3EBE">
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<mods:date id="E93AB3A874B851582BA2813AF391BBDE">2019</mods:date>
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<mods:number id="1072F0DED2B6212B5866E414951A40B7">2019-04-08</mods:number>
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<mods:number id="E513F6DDB10510DAED3DC29484E1516C">4577</mods:number>
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<mods:number id="00B559926C2A4EC5CD253537A37AAABB">3</mods:number>
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<treatment id="47709B7C0522FFD049D366E2FA6BFBAC" ID-DOI="http://doi.org/10.5281/zenodo.5930735" ID-GBIF-Taxon="156146835" ID-Zenodo-Dep="5930735" LSID="urn:lsid:plazi:treatment:47709B7C0522FFD049D366E2FA6BFBAC" httpUri="http://treatment.plazi.org/id/47709B7C0522FFD049D366E2FA6BFBAC" lastPageId="7" lastPageNumber="522" pageId="3" pageNumber="518">
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<subSubSection id="87C379E10522FFD449D366E2FE06FDB4" box="[151,496,494,520]" pageId="3" pageNumber="518" type="nomenclature">
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<paragraph id="CF662A6A0522FFD449D366E2FE06FDB4" blockId="3.[151,496,494,554]" box="[151,496,494,520]" pageId="3" pageNumber="518">
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<heading id="942E9D060522FFD449D366E2FE06FDB4" bold="true" box="[151,496,494,520]" fontSize="11" level="1" pageId="3" pageNumber="518" reason="1">
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<emphasis id="FDADF6780522FFD449D366E2FE06FDB4" bold="true" box="[151,496,494,520]" pageId="3" pageNumber="518">
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||
<taxonomicName id="08D951E90522FFD449D366E2FE86FDB4" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[151,368,494,520]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780522FFD449D366E2FE86FDB4" bold="true" box="[151,368,494,520]" italics="true" pageId="3" pageNumber="518">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030522FFD4483C66E2FE06FDB4" box="[376,496,494,520]" pageId="3" pageNumber="518" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10522FFD449D3651DFC3BFDD6" pageId="3" pageNumber="518" type="description">
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<paragraph id="CF662A6A0522FFD449D3651DFEE4FD95" blockId="3.[151,496,494,554]" box="[151,274,529,554]" pageId="3" pageNumber="518">
|
||
(
|
||
<figureCitation id="57E236EF0522FFD449DB651DFEFFFD95" box="[159,265,529,554]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,250,1977,1999]" captionStartId-1="5.[151,250,1913,1935]" captionStartId-2="9.[151,250,1943,1965]" captionStartId-3="10.[151,250,1595,1617]" captionTargetBox-0="[398,1194,194,1960]" captionTargetBox-1="[313,1273,1152,1901]" captionTargetBox-2="[163,1423,193,1922]" captionTargetBox-3="[151,1436,193,1574]" captionTargetId-0="figure@4.[398,1194,193,1960]" captionTargetId-1="figure@5.[313,1273,1152,1901]" captionTargetId-2="figure@9.[163,1423,193,1922]" captionTargetId-3="figure@10.[151,1436,193,1574]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 1. Lycodon gibsonae spec. nov., FMNH 180146, preserved holotype, A: dorsal view; B: ventral view; C: lateral view of head. Photographs by Joshua Mata." captionText-1="FIGURE 2. Lycodon gibsonae spec. nov., FMNH 180145, preserved paratype, dorsal view. Photograph by Joshua Mata." captionText-2="FIGURE 3. Lycodon gibsonae spec. nov., FMNH 180146, preserved holotype, compared with an adult L. fasciatus (FMNH 270716, Nan Province, Thailand). A: dorsal view of body; B: dorsal view of head. Photographs by Joshua Mata." captionText-3="FIGURE 4. Lycodon gibsonae spec. nov., FMNH 180144, preserved paratype, compared with an adult L. fasciatus of the same size (FMNH 270716, Nan Province, Thailand). A: dorsal view of body; B: ventral view of body. Compare the shape of the bands which are well defined posteriorly in L. gibsonae spec. nov. whereas in Lycodon fasciatus posterior bands are much more strongly speckled with dark dots than anterior bands, up to turning nearly entirely brown. Photographs by Joshua Mata." httpUri-0="https://zenodo.org/record/2632377/files/figure.png" httpUri-1="https://zenodo.org/record/2632379/files/figure.png" httpUri-2="https://zenodo.org/record/2632381/files/figure.png" httpUri-3="https://zenodo.org/record/2632385/files/figure.png" pageId="3" pageNumber="518">Figs. 1–4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD449D36558FC3BFDD6" blockId="3.[151,973,596,618]" box="[151,973,596,618]" pageId="3" pageNumber="518">
|
||
<treatmentCitationGroup id="EFC90D440522FFD449D36558FC3BFDD6" box="[151,973,596,618]" pageId="3" pageNumber="518">
|
||
<taxonomicName id="08D951E90522FFD449D36558FEBBFDD6" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[151,333,596,618]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780522FFD449D36558FEBBFDD6" box="[151,333,596,618]" italics="true" pageId="3" pageNumber="518">Lycodon fasciatus</emphasis>
|
||
</taxonomicName>
|
||
(nec
|
||
<taxonomicName id="08D951E90522FFD448C26558FD55FDD6" authority="Anderson" authorityName="Anderson" authorityYear="1879" box="[390,675,596,618]" class="Reptilia" family="Colubridae" genus="Ophites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780522FFD448C26558FDC3FDD6" box="[390,565,596,618]" italics="true" pageId="3" pageNumber="518">Ophites fasciatus</emphasis>
|
||
Anderson
|
||
</taxonomicName>
|
||
):
|
||
<treatmentCitation id="4E780C7B0522FFD44BF76559FC3FFDD6" author="Inger, R. F. & Colwell, R. K." box="[691,969,596,618]" page="235" pageId="3" pageNumber="518" year="1977">
|
||
<bibRefCitation id="AB48579B0522FFD44BF76559FC60FDD6" author="Inger, R. F. & Colwell, R. K." box="[691,918,596,618]" pageId="3" pageNumber="518" pagination="229 - 253" refId="ref6449" refString="Inger, R. F. & Colwell, R. K. (1977) Organization of contiguous communities of amphibians and reptiles in Thailand. Ecological Monographs, 47, 229 - 253. https: // doi. org / 10.2307 / 1942516" type="journal article" year="1977">Inger & Colwell 1977</bibRefCitation>
|
||
: 235
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10522FFD449D3659AFEEEFC33" pageId="3" pageNumber="518" type="materials_examined">
|
||
<paragraph id="CF662A6A0522FFD449D3659AFAAAFD46" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<materialsCitation id="7FB120370522FFD449D3659AFAAAFD46" ID-GBIF-Occurrence="2235579194" collectingDate="1969-05-05" collectionCode="FMNH" collectorName="W. Ronald Heyer" country="Thailand" location="Sakaerat Environmental Research Station" pageId="3" pageNumber="518" specimenCode="FMNH 180146" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="holotype">
|
||
<emphasis id="FDADF6780522FFD449D3659AFEF3FD13" box="[151,261,662,687]" italics="true" pageId="3" pageNumber="518">
|
||
<typeStatus id="106294C80522FFD449D3659AFEF7FD13" box="[151,257,662,687]" pageId="3" pageNumber="518" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="9F7F82110522FFD44848659BFE4AFD13" box="[268,444,662,687]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="518" type="Museum">
|
||
FMNH
|
||
<date id="BB670CAA0522FFD44821659AFE4AFD13" box="[357,444,662,687]" pageId="3" pageNumber="518" value="46-01-18">180146</date>
|
||
</specimenCode>
|
||
, an adult male (tail dissected), from “Sakaerat Exp. Station, Amphoe Pak Thong Chai”, now the
|
||
<location id="CA067CB10522FFD449B265B6FD3FFD6F" LSID="urn:lsid:plazi:treatment:47709B7C0522FFD049D366E2FA6BFBAC:CA067CB10522FFD449B265B6FD3FFD6F" box="[246,713,698,723]" country="Thailand" name="Sakaerat Environmental Research Station" pageId="3" pageNumber="518" stateProvince="Nakhon Ratchasima">Sakaerat Environmental Research Station</location>
|
||
, Amphoe Wang Nam Khiao, at the south-western edge of
|
||
<collectingRegion id="0D1DE4880522FFD44C0A65B6FA6AFD6F" box="[1358,1436,698,723]" country="Thailand" name="Nakhon Ratchasima" pageId="3" pageNumber="518">Khorat</collectingRegion>
|
||
Plateau,
|
||
<collectingRegion id="0D1DE4880522FFD449B365EDFDB0FD46" box="[247,582,737,762]" country="Thailand" name="Nakhon Ratchasima" pageId="3" pageNumber="518">Nakhon Ratchasima Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="B7CE6AFA0522FFD44B1665EDFD40FD46" box="[594,694,737,762]" name="Thailand" pageId="3" pageNumber="518">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="57E236EF0522FFD44B8165EDFCE4FD46" box="[709,786,737,762]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,1999]" captionTargetBox="[398,1194,194,1960]" captionTargetId="figure@4.[398,1194,193,1960]" captionTargetPageId="4" captionText="FIGURE 1. Lycodon gibsonae spec. nov., FMNH 180146, preserved holotype, A: dorsal view; B: ventral view; C: lateral view of head. Photographs by Joshua Mata." httpUri="https://zenodo.org/record/2632377/files/figure.png" pageId="3" pageNumber="518">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="57E236EF0522FFD44A6465EEFCD9FD46" box="[800,815,738,762]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1965]" captionTargetBox="[163,1423,193,1922]" captionTargetId="figure@9.[163,1423,193,1922]" captionTargetPageId="9" captionText="FIGURE 3. Lycodon gibsonae spec. nov., FMNH 180146, preserved holotype, compared with an adult L. fasciatus (FMNH 270716, Nan Province, Thailand). A: dorsal view of body; B: dorsal view of head. Photographs by Joshua Mata." httpUri="https://zenodo.org/record/2632381/files/figure.png" pageId="3" pageNumber="518">3</figureCitation>
|
||
); collected by
|
||
<collectorName id="622C4FBC0522FFD44A9165EEFB61FD46" box="[981,1175,737,762]" pageId="3" pageNumber="518">W. Ronald Heyer</collectorName>
|
||
on
|
||
<collectingDate id="AB23F5420522FFD44D8565EDFAA1FD46" box="[1217,1367,733,762]" pageId="3" pageNumber="518" value="1969-05-05">
|
||
<date id="BB670CAA0522FFD44D8565EDFAA1FD46" box="[1217,1367,733,762]" pageId="3" pageNumber="518" value="1969-05-05">
|
||
5
|
||
<superScript id="38AC87220522FFD44D8B65D1FB2DFD57" attach="left" box="[1231,1243,733,747]" fontSize="6" pageId="3" pageNumber="518">th</superScript>
|
||
May 1969
|
||
</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD44983640AFEEEFC33" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<materialsCitation id="7FB120370522FFD44983640AFBD2FCF9" ID-GBIF-Occurrence="2235579192" collectingDate="1969-03-26" collectionCode="FMNH" collectorName="W. Ronald Heyer" country="Thailand" location="Sakaerat Environmental Research Station" pageId="3" pageNumber="518" specimenCode="FMNH 180144" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
|
||
<typeStatus id="106294C80522FFD44983640AFECFFCA1" box="[199,313,774,797]" pageId="3" pageNumber="518" type="paratype">
|
||
<emphasis id="FDADF6780522FFD44983640AFECFFCA1" box="[199,313,774,797]" italics="true" pageId="3" pageNumber="518">Paratypes</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<emphasis id="FDADF6780522FFD44808640AFEACFCA1" box="[332,346,774,797]" italics="true" pageId="3" pageNumber="518">n</emphasis>
|
||
=2). (1)
|
||
<specimenCode id="9F7F82110522FFD448FA640AFD98FCA2" box="[446,622,774,798]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="518" type="Museum">
|
||
FMNH
|
||
<date id="BB670CAA0522FFD44B5E640AFD98FCA2" box="[538,622,774,798]" pageId="3" pageNumber="518" value="44-01-18">180144</date>
|
||
</specimenCode>
|
||
(
|
||
<figureCitation id="57E236EF0522FFD44BC56409FD3FFCA2" box="[641,713,773,798]" captionStart="FIGURE 4" captionStartId="10.[151,250,1595,1617]" captionTargetBox="[151,1436,193,1574]" captionTargetId="figure@10.[151,1436,193,1574]" captionTargetPageId="10" captionText="FIGURE 4. Lycodon gibsonae spec. nov., FMNH 180144, preserved paratype, compared with an adult L. fasciatus of the same size (FMNH 270716, Nan Province, Thailand). A: dorsal view of body; B: ventral view of body. Compare the shape of the bands which are well defined posteriorly in L. gibsonae spec. nov. whereas in Lycodon fasciatus posterior bands are much more strongly speckled with dark dots than anterior bands, up to turning nearly entirely brown. Photographs by Joshua Mata." httpUri="https://zenodo.org/record/2632385/files/figure.png" pageId="3" pageNumber="518">Fig. 4</figureCitation>
|
||
), an adult male (dissected; hemipenes in situ), from the same locality as the holotype; collected by
|
||
<collectorName id="622C4FBC0522FFD44B786421FD08FCF9" box="[572,766,812,837]" pageId="3" pageNumber="518">W. Ronald Heyer</collectorName>
|
||
on
|
||
<collectingDate id="AB23F5420522FFD44A6D6420FC15FCF9" box="[809,995,808,837]" pageId="3" pageNumber="518" value="1969-03-26">
|
||
<date id="BB670CAA0522FFD44A6D6420FC15FCF9" box="[809,995,808,837]" pageId="3" pageNumber="518" value="1969-03-26">
|
||
26
|
||
<superScript id="38AC87220522FFD44A016424FCA7FC8A" attach="left" box="[837,849,808,822]" fontSize="6" pageId="3" pageNumber="518">th</superScript>
|
||
March 1969
|
||
</date>
|
||
</collectingDate>
|
||
.—(2)
|
||
</materialsCitation>
|
||
<materialsCitation id="7FB120370522FFD44D686421FEE2FC33" ID-GBIF-Occurrence="2235579193" collectingDate="1969-04-14" collectionCode="FMNH" collectorName="W. Ronald Heyer" country="Thailand" location="Sakaerat Environmental Research Station" pageId="3" pageNumber="518" specimenCode="FMNH 180145" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
|
||
<specimenCode id="9F7F82110522FFD44D686421FB2EFCF9" box="[1068,1240,812,837]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="518" type="Museum">
|
||
FMNH
|
||
<date id="BB670CAA0522FFD44DC06420FB2EFCF9" box="[1156,1240,812,837]" pageId="3" pageNumber="518" value="45-01-18">180145</date>
|
||
</specimenCode>
|
||
(
|
||
<figureCitation id="57E236EF0522FFD44DAC6420FADBFCF9" box="[1256,1325,812,837]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1935]" captionTargetBox="[313,1273,1152,1901]" captionTargetId="figure@5.[313,1273,1152,1901]" captionTargetPageId="5" captionText="FIGURE 2. Lycodon gibsonae spec. nov., FMNH 180145, preserved paratype, dorsal view. Photograph by Joshua Mata." httpUri="https://zenodo.org/record/2632379/files/figure.png" pageId="3" pageNumber="518">Fig. 2</figureCitation>
|
||
), an adult male (dissected; hemipenes in situ), from the same locality as the holotype; collected by
|
||
<collectorName id="622C4FBC0522FFD44DC06458FABEFCD0" box="[1156,1352,851,876]" pageId="3" pageNumber="518">W. Ronald Heyer</collectorName>
|
||
on
|
||
<collectingDate id="AB23F5420522FFD44C30645FFEE2FC33" pageId="3" pageNumber="518" value="1969-04-14">
|
||
<date id="BB670CAA0522FFD44C30645FFEE2FC33" pageId="3" pageNumber="518" value="1969-04-14">
|
||
14
|
||
<superScript id="38AC87220522FFD44CD46443FA6AFCE1" attach="none" box="[1424,1436,847,861]" fontSize="6" pageId="3" pageNumber="518">th</superScript>
|
||
April 1969
|
||
</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10522FFD449836497FC7DFB68" pageId="3" pageNumber="518" type="diagnosis">
|
||
<paragraph id="CF662A6A0522FFD449836497FDE3FBF8" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<emphasis id="FDADF6780522FFD449836497FEB7FC0F" bold="true" box="[199,321,923,947]" italics="true" pageId="3" pageNumber="518">Diagnosis.</emphasis>
|
||
A species of the genus
|
||
<taxonomicName id="08D951E90522FFD44B1D6497FD4FFC08" box="[601,697,923,948]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="genus">
|
||
<emphasis id="FDADF6780522FFD44B1D6497FD4FFC08" box="[601,697,923,948]" italics="true" pageId="3" pageNumber="518">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
, characterized by: (1) loreal entering orbit; (2) 17–17–15 dorsal scale rows; (3) upper 3 or 4 and vertebral dorsal scale rows keeled; (4) about 223–226
|
||
<collectingRegion id="0D1DE4880522FFD44D1E64CCFB62FC6B" box="[1114,1172,960,983]" country="Austria" name="Wien" pageId="3" pageNumber="518">VEN</collectingRegion>
|
||
in males; (5) about 91– 92 SC in males; (6) relative tail length about
|
||
<quantity id="0821878F0522FFD44BD764EFFD19FC40" box="[659,751,995,1020]" metricMagnitude="-3" metricUnit="m" metricValue="5.0292" pageId="3" pageNumber="518" unit="in" value="0.198">0.198 in</quantity>
|
||
males; (7) 17–18 cream, well-defined bands on a dark body; (8) first dorsal band starting at
|
||
<collectingRegion id="0D1DE4880522FFD448AA6304FDDEFBA3" box="[494,552,1032,1055]" country="Austria" name="Wien" pageId="3" pageNumber="518">VEN</collectingRegion>
|
||
13–15; (9) width of the first band occupying 4, 5 or 6 vertebral scales; and (10) maximum SVL
|
||
<quantity id="0821878F0522FFD4480F6327FE5EFBFF" box="[331,424,1067,1092]" metricMagnitude="-1" metricUnit="m" metricValue="9.059999999999999" pageId="3" pageNumber="518" unit="mm" value="906.0">906 mm</quantity>
|
||
in males.
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD449836342FC7DFB68" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
This species can be recognized by its ventral and subcaudal scale counts (both higher than in
|
||
<taxonomicName id="08D951E90522FFD44DA2635CFA9EFBDB" authorityName="Anderson" authorityYear="1879" box="[1254,1384,1103,1127]" class="Reptilia" family="Colubridae" genus="Ophites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780522FFD44DA2635CFA9EFBDB" box="[1254,1384,1103,1127]" italics="true" pageId="3" pageNumber="518">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
), by the first band on the body (broader in terms of vertebral scales than in
|
||
<taxonomicName id="08D951E90522FFD44AEE6378FBDDFB37" authorityName="Anderson" authorityYear="1879" box="[938,1067,1139,1163]" class="Reptilia" family="Colubridae" genus="Ophites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780522FFD44AEE6378FBDDFB37" box="[938,1067,1139,1163]" italics="true" pageId="3" pageNumber="518">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
), and on the number of bands on the body (lower than in
|
||
<emphasis id="FDADF6780522FFD448E76394FDC3FB13" box="[419,565,1175,1199]" italics="true" pageId="3" pageNumber="518">
|
||
<taxonomicName id="08D951E90522FFD448E76394FDDCFB13" authorityName="Anderson" authorityYear="1879" box="[419,554,1175,1199]" class="Reptilia" family="Colubridae" genus="Ophites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="fasciatus">L. fasciatus</taxonomicName>
|
||
).
|
||
</emphasis>
|
||
Most other characters match those of
|
||
<taxonomicName id="08D951E90522FFD44AA56394FB94FB13" authorityName="Anderson" authorityYear="1879" box="[993,1122,1175,1199]" class="Reptilia" family="Colubridae" genus="Ophites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780522FFD44AA56394FB94FB13" box="[993,1122,1175,1199]" italics="true" pageId="3" pageNumber="518">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
. Detailed comparisons with other species of the genus
|
||
<taxonomicName id="08D951E90522FFD448FB63B7FDE9FB68" box="[447,543,1211,1236]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="genus">
|
||
<emphasis id="FDADF6780522FFD448FB63B7FDE9FB68" box="[447,543,1211,1236]" italics="true" pageId="3" pageNumber="518">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
appear below in the Discussion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10522FFD4498363D2FC3AFA10" pageId="3" pageNumber="518" type="etymology">
|
||
<paragraph id="CF662A6A0522FFD4498363D2FD7AFAD8" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<emphasis id="FDADF6780522FFD4498363D2FEB1FB4B" bold="true" box="[199,327,1246,1271]" italics="true" pageId="3" pageNumber="518">Etymology.</emphasis>
|
||
This species is named after Mrs. Nancy Lynne Gibson, Executive director of the “Bird Conservation Society of
|
||
<collectingCountry id="B7CE6AFA0522FFD448F6620FFDEEFAA0" box="[434,536,1283,1308]" name="Thailand" pageId="3" pageNumber="518">Thailand</collectingCountry>
|
||
” and President of the executive committee of the “Love Wildlife Foundation”, for her involvement in the protection of the wildlife in
|
||
<collectingCountry id="B7CE6AFA0522FFD44A68622BFC66FAFC" box="[812,912,1319,1344]" name="Thailand" pageId="3" pageNumber="518">Thailand</collectingCountry>
|
||
and, especially, of the primary forest which harbour the
|
||
<typeStatus id="106294C80522FFD4485A6240FEB8FAD8" box="[286,334,1356,1380]" pageId="3" pageNumber="518">type</typeStatus>
|
||
locality of this new species.
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD44983627CFC3AFA10" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
We suggest the following common names: Gibson’s Wolf Snake (English);
|
||
<taxonomicName id="08D951E90522FFD44D006263FA6EFA34" authority="de Gibson (French)" box="[1092,1432,1391,1416]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="518" phylum="Chordata" rank="genus">Lycodon de Gibson (French)</taxonomicName>
|
||
; Gibsons Wolfszahnnatter (German); Ngoo Plong-chanuan Gibson (Thai).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10522FFD1498362BBFAE2FA45" lastPageId="6" lastPageNumber="521" pageId="3" pageNumber="518" type="description">
|
||
<paragraph id="CF662A6A0522FFD4498362BBFCE4F914" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<emphasis id="FDADF6780522FFD4498362BBFDF0FA73" bold="true" box="[199,518,1462,1487]" italics="true" pageId="3" pageNumber="518">
|
||
Description of the
|
||
<typeStatus id="106294C80522FFD448D962BAFDF4FA73" box="[413,514,1462,1487]" pageId="3" pageNumber="518" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body robust, slightly compressed laterally; neck forming a natural angle with the head; head subrectangular, elongate, flattened, well distinct from the neck; snout elongate, strongly flattened, oblique in profile, projecting over the lower jaw, 27.5 % of HL, or 2.2 times as long as diameter of eye; a weak canthus rostralis; upper maxilla strongly arched with an angulous apex, distinctly bent inwards anteriorly (one maxilla dissected); large oval nostril that opens laterally and nearly entirely divides the nasal at its center; eye average, its diameter barely greater than the distance between the edge of the lip and its lower margin, with a vertical pupil; tail comparatively long, thin and tapering.
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD4498361BFFD64F970" blockId="3.[151,1437,662,1961]" box="[199,658,1715,1740]" pageId="3" pageNumber="518">
|
||
SVL
|
||
<quantity id="0821878F0522FFD4484761BFFE95F977" box="[259,355,1715,1740]" metricMagnitude="-1" metricUnit="m" metricValue="9.059999999999999" pageId="3" pageNumber="518" unit="mm" value="906.0">906 mm</quantity>
|
||
; TaL>
|
||
<quantity id="0821878F0522FFD448FC61BFFDE0F977" box="[440,534,1715,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.94" pageId="3" pageNumber="518" unit="mm" value="194.0">194 mm</quantity>
|
||
(mutilate).
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD4498361D4FC6CF884" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<emphasis id="FDADF6780522FFD4498361D4FECEF953" box="[199,312,1752,1775]" italics="true" pageId="3" pageNumber="518">Dentition.</emphasis>
|
||
Based on specimen FMNH
|
||
<date id="BB670CAA0522FFD44B3361DBFD39F953" box="[631,719,1751,1775]" pageId="3" pageNumber="518" value="44-01-18">180144</date>
|
||
: total of 13 maxillary teeth, with the following formula: 4 small anterior teeth + 3 strongly enlarged teeth, thick + a wide gap, longer than the larger anterior teeth + 3 small teeth + a small gap + 3 strongly enlarged posterior teeth, also rather straight.
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0522FFD449836049FB86F815" blockId="3.[151,1437,662,1961]" pageId="3" pageNumber="518">
|
||
<emphasis id="FDADF6780522FFD449836049FE86F8E2" box="[199,368,1861,1886]" italics="true" pageId="3" pageNumber="518">Body scalation</emphasis>
|
||
. Dorsal scales in 17–17–15 rows, smooth on 1
|
||
<superScript id="38AC87220522FFD44AC8604EFC60F8EC" attach="none" box="[908,918,1858,1872]" fontSize="6" pageId="3" pageNumber="518">st</superScript>
|
||
–4
|
||
<superScript id="38AC87220522FFD44AF7604EFC49F8EC" attach="left" box="[947,959,1858,1872]" fontSize="6" pageId="3" pageNumber="518">th</superScript>
|
||
dorsal rows, moderately keeled on 5
|
||
<superScript id="38AC87220522FFD44C2C604EFA82F8EC" attach="none" box="[1384,1396,1858,1872]" fontSize="6" pageId="3" pageNumber="518">th</superScript>
|
||
–9
|
||
<superScript id="38AC87220522FFD44CD4604EFA6AF8EC" attach="none" box="[1424,1436,1858,1872]" fontSize="6" pageId="3" pageNumber="518">th</superScript>
|
||
rows along the entire body; scales of 1
|
||
<superScript id="38AC87220522FFD44B016065FDB9F8CB" attach="left" box="[581,591,1897,1911]" fontSize="6" pageId="3" pageNumber="518">st</superScript>
|
||
dorsal scale row slightly enlarged; scales of the vertebral row not enlarged; no apical pits detected; 225
|
||
<collectingRegion id="0D1DE4880522FFD44894609DFDFCF814" box="[464,522,1937,1960]" country="Austria" name="Wien" pageId="3" pageNumber="518">VEN</collectingRegion>
|
||
(+ 2 preventrals),> 73 paired SC; cloacal plate single.
|
||
</paragraph>
|
||
<caption id="9BA67AE20525FFD349D360B5FDECF852" httpUri="https://zenodo.org/record/2632377/files/figure.png" pageId="4" pageNumber="519" startId="4.[151,250,1977,1999]" targetBox="[398,1194,194,1960]" targetPageId="4">
|
||
<paragraph id="CF662A6A0525FFD349D360B5FDECF852" blockId="4.[151,1435,1977,2030]" pageId="4" pageNumber="519">
|
||
<emphasis id="FDADF6780525FFD349D360B5FEE4F873" bold="true" box="[151,274,1977,1999]" pageId="4" pageNumber="519">FIGURE 1.</emphasis>
|
||
<taxonomicName id="08D951E90525FFD3485C60B5FE26F873" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[280,464,1977,1999]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="519" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780525FFD3485C60B5FE26F873" box="[280,464,1977,1999]" italics="true" pageId="4" pageNumber="519">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030525FFD3489260B5FDCAF873" box="[470,572,1977,1999]" pageId="4" pageNumber="519" rank="species">
|
||
<emphasis id="FDADF6780525FFD3489260B5FDC1F873" bold="true" box="[470,567,1977,1999]" pageId="4" pageNumber="519">spec. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, FMNH 180146, preserved holotype, A: dorsal view; B: ventral view; C: lateral view of head. Photographs by Joshua Mata.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CF662A6A0524FFD24983679BFD5BFF0C" blockId="5.[199,685,151,176]" box="[199,685,151,176]" pageId="5" pageNumber="520">Dorsal scale row formula (recount system):</paragraph>
|
||
<paragraph id="CF662A6A0524FFD2498367E9FE87FEF7" blockId="5.[151,403,223,333]" pageId="5" pageNumber="520">
|
||
3+4
|
||
<emphasis id="FDADF6780524FFD249BE67D3FEEFFF42" bold="true" box="[250,281,223,254]" italics="true" pageId="5" pageNumber="520">Ą</emphasis>
|
||
3 (143) 17 ————— 15 3+4
|
||
<emphasis id="FDADF6780524FFD249BE6622FEEFFEF1" bold="true" box="[250,281,302,333]" italics="true" pageId="5" pageNumber="520">Ą</emphasis>
|
||
3 (144)
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0524FFD24983667AFD84FBF0" blockId="5.[151,1436,374,1100]" pageId="5" pageNumber="520">
|
||
<emphasis id="FDADF6780524FFD24983667AFE82FE33" box="[199,372,374,399]" italics="true" pageId="5" pageNumber="520">Head scalation</emphasis>
|
||
. Complement of upper head scales complete, including two internasals, two prefrontals, two supraoculars, one frontal, and two parietals. Rostral triangular, rather short, much wider than high, well visible from above but barely separating internasals; nasal rectangular, vertically divided into two nearly equal parts by a furrow below and above the nostril piercing in between, altogether about 1.7 times as long as high; internasals small, trapezoidal, about 1.5 times wider than long, widely in contact each with the other and with prefrontals; prefrontals large, subrectangular, 2.2 times longer than internasals, in broad contact with loreal; frontal rather small, hexagonal or ogive-shaped, its apex pointing backwards, 1.2 times longer than wide and about 1.3 times as long as the prefrontals suture; parietals large, elongate, subtriangular, 1.5 times longer than frontal (suture 1.2 times longer than frontal), each edged posteriorly with three large scales, i.e. 2 upper temporals and 1 larger paraparietal; supraocular entire, short and broad, 1.4 times longer than wide, about 0.6 times as wide as frontal; 1 / 1 loreal, pentagonal or subrectangular, very elongate, strongly narrowed posteriorly, about 3.0 times longer than high, entering orbit on a short height, in contact with 2
|
||
<superScript id="38AC87220524FFD24BF3640DFD31FCB3" attach="left" box="[695,711,769,783]" fontSize="6" pageId="5" pageNumber="520">nd</superScript>
|
||
and 3
|
||
<superScript id="38AC87220524FFD24A4E640DFCE1FCB3" attach="left" box="[778,791,769,783]" fontSize="6" pageId="5" pageNumber="520">rd</superScript>
|
||
SL, the preocular, the prefrontal, and the posterior part of nasal; 8 / 8 SL, of which the first six are taller than wide, 1
|
||
<superScript id="38AC87220524FFD24A746424FCCCFC8A" attach="left" box="[816,826,808,822]" fontSize="6" pageId="5" pageNumber="520">st</superScript>
|
||
and 2
|
||
<superScript id="38AC87220524FFD24AC46424FC66FC8A" attach="left" box="[896,912,808,822]" fontSize="6" pageId="5" pageNumber="520">nd</superScript>
|
||
SL in contact with nasal, 3
|
||
<superScript id="38AC87220524FFD24D806424FB27FC8A" attach="none" box="[1220,1233,808,822]" fontSize="6" pageId="5" pageNumber="520">rd</superScript>
|
||
–5
|
||
<superScript id="38AC87220524FFD24DA96424FB0FFC8A" attach="left" box="[1261,1273,808,822]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
SL on the left and 3
|
||
<superScript id="38AC87220524FFD249916443FF14FCE1" attach="none" box="[213,226,847,861]" fontSize="6" pageId="5" pageNumber="520">rd</superScript>
|
||
–4
|
||
<superScript id="38AC87220524FFD249BA6443FEFCFCE1" attach="left" box="[254,266,847,861]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
SL on the right right side touching the eye, 6
|
||
<superScript id="38AC87220524FFD24A4B6443FCEDFCE1" attach="left" box="[783,795,847,861]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
and 7
|
||
<superScript id="38AC87220524FFD24A266443FC98FCE1" attach="left" box="[866,878,847,861]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
SL largest; 1 / 1 small, subrectangular preocular, located above the posterior part of loreal, reaching the top of the head and broadly in contact with the frontal; 1 large subocular at right, keeping 5
|
||
<superScript id="38AC87220524FFD24B6C6496FDC2FC14" attach="left" box="[552,564,922,936]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
SL from entering orbit; 2 / 2 postoculars, the upper one slightly larger; 2 anterior temporals on each side, followed by 3 secondary temporals, lower anterior temporal much larger than upper one, posterior temporals smaller; 10 / 10 IL, first pair in contact, 1
|
||
<superScript id="38AC87220524FFD24AB464E8FC0CFC4E" attach="none" box="[1008,1018,996,1010]" fontSize="6" pageId="5" pageNumber="520">st</superScript>
|
||
–5
|
||
<superScript id="38AC87220524FFD24D5364E8FBD5FC4E" attach="left" box="[1047,1059,996,1010]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
IL in contact with anterior chin shields, 6
|
||
<superScript id="38AC87220524FFD248456300FEFBFBA6" attach="none" box="[257,269,1036,1050]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
–8
|
||
<superScript id="38AC87220524FFD2486D6300FEC3FBA6" attach="left" box="[297,309,1036,1050]" fontSize="6" pageId="5" pageNumber="520">th</superScript>
|
||
infralabials largest; mental triangular, longer than wide; anterior and posterior pair of sublinguals of about same length, but anterior pair wider.
|
||
</paragraph>
|
||
<caption id="9BA67AE20524FFD249D36075FA97F833" box="[151,1377,1913,1935]" httpUri="https://zenodo.org/record/2632379/files/figure.png" pageId="5" pageNumber="520" startId="5.[151,250,1913,1935]" targetBox="[313,1273,1152,1901]" targetPageId="5">
|
||
<paragraph id="CF662A6A0524FFD249D36075FA97F833" blockId="5.[151,1377,1913,1935]" box="[151,1377,1913,1935]" pageId="5" pageNumber="520">
|
||
<emphasis id="FDADF6780524FFD249D36075FEE4F833" bold="true" box="[151,274,1913,1935]" pageId="5" pageNumber="520">FIGURE 2.</emphasis>
|
||
<taxonomicName id="08D951E90524FFD2485D6075FE27F833" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[281,465,1913,1935]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="520" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780524FFD2485D6075FE27F833" box="[281,465,1913,1935]" italics="true" pageId="5" pageNumber="520">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030524FFD248936075FDC8F833" box="[471,574,1913,1935]" pageId="5" pageNumber="520" rank="species">
|
||
<emphasis id="FDADF6780524FFD248936075FDCEF833" bold="true" box="[471,568,1913,1935]" pageId="5" pageNumber="520">spec. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, FMNH 180145, preserved paratype, dorsal view. Photograph by Joshua Mata.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CF662A6A0524FFD1498360B8FD6DFEA1" blockId="5.[151,1436,1972,2033]" lastBlockId="6.[151,1437,151,680]" lastPageId="6" lastPageNumber="521" pageId="5" pageNumber="520">
|
||
<emphasis id="FDADF6780524FFD2498360B8FE07F871" box="[199,497,1972,1997]" italics="true" pageId="5" pageNumber="520">Coloration in preservative</emphasis>
|
||
. Body and tail dark chestnut brown, with 17 well-defined crossbands on body and eight on tail, cream, strongly speckled with dark brown spots; these crossbands, about 4–5 dorsal scale long at mid- height of sides, widen at their ventrolateral limit, up to about 5–7 dorsal scale long; the first crossband, beginning at the level of
|
||
<collectingRegion id="0D1DE4880527FFD1485967B1FEA1FF68" box="[285,343,189,212]" country="Austria" name="Wien" pageId="6" pageNumber="521">VEN</collectingRegion>
|
||
14, is narrow, whereas the second crossband begins at
|
||
<collectingRegion id="0D1DE4880527FFD14A8167B1FC09FF68" box="[965,1023,189,212]" country="Austria" name="Wien" pageId="6" pageNumber="521">VEN</collectingRegion>
|
||
31; posterior crossbands closer each to the other, separated by 8–10 DSR at mid-height of sides and about 6 DSR just above the venter; crossbands of the tail cream, also speckled with dark brown.
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0527FFD14983662BFD66FE34" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="521">The head is nearly uniformly dark brown, slightly paler on the sides of the snout; supralabials speckled with pale brown, especially behind the level of the eye; no nuchal collar visible. The throat is cream, strongly speckled with dark brown on the chin and infralabials.</paragraph>
|
||
<paragraph id="CF662A6A0527FFD149836698FE25FE48" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="521">The venter is cream, ornate throughout with both strong, narrow dark brown blotches in its middle and the dark brown dorsal color between the cream. The under surface of the tail is as the venter near its base, then mostly dark brown speckled with cream.</paragraph>
|
||
<paragraph id="CF662A6A0527FFD14983650CFE6BFD14" blockId="6.[151,1437,151,680]" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD14983650CFECFFDA4" bold="true" box="[199,313,512,536]" italics="true" pageId="6" pageNumber="521">Variation.</emphasis>
|
||
Both
|
||
<typeStatus id="106294C80527FFD148C7650CFE18FDA4" box="[387,494,512,536]" pageId="6" pageNumber="521" type="paratype">paratypes</typeStatus>
|
||
(FMNH
|
||
<date id="BB670CAA0527FFD14B1966F3FD44FDA4" box="[605,690,511,536]" pageId="6" pageNumber="521" value="44-01-18">180144</date>
|
||
–45) agree in most respects with the description of the
|
||
<typeStatus id="106294C80527FFD14C7766F3FA6FFDA4" box="[1331,1433,511,536]" pageId="6" pageNumber="521" type="holotype">holotype</typeStatus>
|
||
. Dorsal bands of specimen FMNH
|
||
<date id="BB670CAA0527FFD14B5F6528FD99FD81" box="[539,623,548,573]" pageId="6" pageNumber="521" value="45-01-18">180145</date>
|
||
are only slightly speckled with dark brown. Both
|
||
<typeStatus id="106294C80527FFD14DE16529FAE7FD81" box="[1189,1297,549,573]" pageId="6" pageNumber="521" type="paratype">paratypes</typeStatus>
|
||
have a faint collar. It appears that these specimens represent animals younger than the
|
||
<typeStatus id="106294C80527FFD14AA6654BFBB1FDDC" box="[994,1095,583,608]" pageId="6" pageNumber="521" type="holotype">holotype</typeStatus>
|
||
, so some of the differences in coloration may indicate an ontogenetical shift. A comparison of the most important morphological characters are summarized in
|
||
<tableCitation id="825B1FD10527FFD148006583FE6FFD14" box="[324,409,655,680]" captionStart="TABLE 2" captionStartId="6.[151,239,724,747]" captionText="TABLE 2. Morphological characters of the type specimens of Lycodon gibsonae spec. nov." pageId="6" pageNumber="521">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="9BA67AE20527FFD149D365D8FB9FFD57" box="[151,1129,724,747]" pageId="6" pageNumber="521">
|
||
<paragraph id="CF662A6A0527FFD149D365D8FB9FFD57" blockId="6.[151,1129,724,747]" box="[151,1129,724,747]" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD149D365D8FEFFFD57" bold="true" box="[151,265,724,747]" pageId="6" pageNumber="521">TABLE 2.</emphasis>
|
||
Morphological characters of the type specimens of
|
||
<taxonomicName id="08D951E90527FFD14A7065D8FC03FD57" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[820,1013,724,747]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780527FFD14A7065D8FC03FD57" box="[820,1013,724,747]" italics="true" pageId="6" pageNumber="521">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030527FFD14AB865D8FB9FFD57" box="[1020,1129,724,747]" pageId="6" pageNumber="521" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CF662A6A0527FFD149DB640AFB86FAE8" pageId="6" pageNumber="521">
|
||
<table id="BDD9D8CA0527002849DB640AFB08FAE8" box="[159,1278,774,1364]" gridcols="4" gridrows="14" pageId="6" pageNumber="521">
|
||
<tr id="71E928280527002849DB640AFB08FC87" box="[159,1278,774,827]" gridrow="0" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB640AFED5FC87" box="[159,291,774,827]" gridcol="0" gridrow="0" pageId="6" pageNumber="521">Characters</th>
|
||
<th id="323841540527002848A4640AFD8AFC87" box="[480,636,774,827]" gridcol="1" gridrow="0" pageId="6" pageNumber="521">FMNH 180146 holotype</th>
|
||
<th id="32384154052700284A65640AFC4BFC87" box="[801,957,774,827]" gridcol="2" gridrow="0" pageId="6" pageNumber="521">FMNH 180144 paratype</th>
|
||
<th id="32384154052700284D27640AFB08FC87" box="[1123,1278,774,827]" gridcol="3" gridrow="0" pageId="6" pageNumber="521">FMNH 180145 paratype</th>
|
||
</tr>
|
||
<tr id="71E928280527002849DB6442FB08FCD8" box="[159,1278,846,868]" gridrow="1" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB6442FED5FCD8" box="[159,291,846,868]" gridcol="0" gridrow="1" pageId="6" pageNumber="521">Sex</th>
|
||
<td id="323841540527002848A46442FD8AFCD8" box="[480,636,846,868]" gridcol="1" gridrow="1" pageId="6" pageNumber="521">Male</td>
|
||
<td id="32384154052700284A656442FC4BFCD8" box="[801,957,846,868]" gridcol="2" gridrow="1" pageId="6" pageNumber="521">Male</td>
|
||
<td id="32384154052700284D276442FB08FCD8" box="[1123,1278,846,868]" gridcol="3" gridrow="1" pageId="6" pageNumber="521">Male</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB6474FB08FC32" box="[159,1278,888,910]" gridrow="2" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB6474FED5FC32" box="[159,291,888,910]" gridcol="0" gridrow="2" pageId="6" pageNumber="521">SVL</th>
|
||
<td id="323841540527002848A46474FD8AFC32" box="[480,636,888,910]" gridcol="1" gridrow="2" pageId="6" pageNumber="521">906</td>
|
||
<td id="32384154052700284A656474FC4BFC32" box="[801,957,888,910]" gridcol="2" gridrow="2" pageId="6" pageNumber="521">522</td>
|
||
<td id="32384154052700284D276474FB08FC32" box="[1123,1278,888,910]" gridcol="3" gridrow="2" pageId="6" pageNumber="521">430</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB64ADFB08FC0B" box="[159,1278,929,951]" gridrow="3" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB64ADFED5FC0B" box="[159,291,929,951]" gridcol="0" gridrow="3" pageId="6" pageNumber="521">TaL</th>
|
||
<td id="323841540527002848A464ADFD8AFC0B" box="[480,636,929,951]" gridcol="1" gridrow="3" pageId="6" pageNumber="521">----</td>
|
||
<td id="32384154052700284A6564ADFC4BFC0B" box="[801,957,929,951]" gridcol="2" gridrow="3" pageId="6" pageNumber="521">129</td>
|
||
<td id="32384154052700284D2764ADFB08FC0B" box="[1123,1278,929,951]" gridcol="3" gridrow="3" pageId="6" pageNumber="521">106</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB64C6FB08FC5C" box="[159,1278,970,992]" gridrow="4" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB64C6FED5FC5C" box="[159,291,970,992]" gridcol="0" gridrow="4" pageId="6" pageNumber="521">Rel TL</th>
|
||
<td id="323841540527002848A464C6FD8AFC5C" box="[480,636,970,992]" gridcol="1" gridrow="4" pageId="6" pageNumber="521">----</td>
|
||
<td id="32384154052700284A6564C6FC4BFC5C" box="[801,957,970,992]" gridcol="2" gridrow="4" pageId="6" pageNumber="521">0.198</td>
|
||
<td id="32384154052700284D2764C6FB08FC5C" box="[1123,1278,970,992]" gridcol="3" gridrow="4" pageId="6" pageNumber="521">0.198</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB64F8FB08FBB6" box="[159,1278,1012,1034]" gridrow="5" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB64F8FED5FBB6" box="[159,291,1012,1034]" gridcol="0" gridrow="5" pageId="6" pageNumber="521">DSR</th>
|
||
<td id="323841540527002848A464F8FD8AFBB6" box="[480,636,1012,1034]" gridcol="1" gridrow="5" pageId="6" pageNumber="521">17–17–15</td>
|
||
<td id="32384154052700284A6564F8FC4BFBB6" box="[801,957,1012,1034]" gridcol="2" gridrow="5" pageId="6" pageNumber="521">17–17–15</td>
|
||
<td id="32384154052700284D2764F8FB08FBB6" box="[1123,1278,1012,1034]" gridcol="3" gridrow="5" pageId="6" pageNumber="521">17–17–15</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB6311FB08FB8F" box="[159,1278,1053,1075]" gridrow="6" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB6311FED5FB8F" box="[159,291,1053,1075]" gridcol="0" gridrow="6" pageId="6" pageNumber="521">VEN</th>
|
||
<td id="323841540527002848A46311FD8AFB8F" box="[480,636,1053,1075]" gridcol="1" gridrow="6" pageId="6" pageNumber="521">225</td>
|
||
<td id="32384154052700284A656311FC4BFB8F" box="[801,957,1053,1075]" gridcol="2" gridrow="6" pageId="6" pageNumber="521">223</td>
|
||
<td id="32384154052700284D276311FB08FB8F" box="[1123,1278,1053,1075]" gridcol="3" gridrow="6" pageId="6" pageNumber="521">226</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB634AFB08FBE0" box="[159,1278,1094,1116]" gridrow="7" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB634AFED5FBE0" box="[159,291,1094,1116]" gridcol="0" gridrow="7" pageId="6" pageNumber="521">SC</th>
|
||
<td id="323841540527002848A4634AFD8AFBE0" box="[480,636,1094,1116]" gridcol="1" gridrow="7" pageId="6" pageNumber="521">incomplete</td>
|
||
<td id="32384154052700284A65634AFC4BFBE0" box="[801,957,1094,1116]" gridcol="2" gridrow="7" pageId="6" pageNumber="521">92</td>
|
||
<td id="32384154052700284D27634AFB08FBE0" box="[1123,1278,1094,1116]" gridcol="3" gridrow="7" pageId="6" pageNumber="521">91</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB637CFB08FB3A" box="[159,1278,1136,1158]" gridrow="8" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB637CFED5FB3A" box="[159,291,1136,1158]" gridcol="0" gridrow="8" pageId="6" pageNumber="521">Paraparietals</th>
|
||
<td id="323841540527002848A4637CFD8AFB3A" box="[480,636,1136,1158]" gridcol="1" gridrow="8" pageId="6" pageNumber="521">7/6</td>
|
||
<td id="32384154052700284A65637CFC4BFB3A" box="[801,957,1136,1158]" gridcol="2" gridrow="8" pageId="6" pageNumber="521">7/6</td>
|
||
<td id="32384154052700284D27637CFB08FB3A" box="[1123,1278,1136,1158]" gridcol="3" gridrow="8" pageId="6" pageNumber="521">8/7</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB6395FB08FB13" box="[159,1278,1177,1199]" gridrow="9" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB6395FED5FB13" box="[159,291,1177,1199]" gridcol="0" gridrow="9" pageId="6" pageNumber="521">Dorsal bands</th>
|
||
<td id="323841540527002848A46395FD8AFB13" box="[480,636,1177,1199]" gridcol="1" gridrow="9" pageId="6" pageNumber="521">17</td>
|
||
<td id="32384154052700284A656395FC4BFB13" box="[801,957,1177,1199]" gridcol="2" gridrow="9" pageId="6" pageNumber="521">18</td>
|
||
<td id="32384154052700284D276395FB08FB13" box="[1123,1278,1177,1199]" gridcol="3" gridrow="9" pageId="6" pageNumber="521">17</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB63CEFB08FB64" box="[159,1278,1218,1240]" gridrow="10" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB63CEFED5FB64" box="[159,291,1218,1240]" gridcol="0" gridrow="10" pageId="6" pageNumber="521">Tail bands</th>
|
||
<td id="323841540527002848A463CEFD8AFB64" box="[480,636,1218,1240]" gridcol="1" gridrow="10" pageId="6" pageNumber="521">8</td>
|
||
<td id="32384154052700284A6563CEFC4BFB64" box="[801,957,1218,1240]" gridcol="2" gridrow="10" pageId="6" pageNumber="521">9</td>
|
||
<td id="32384154052700284D2763CEFB08FB64" box="[1123,1278,1218,1240]" gridcol="3" gridrow="10" pageId="6" pageNumber="521">9</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB63E0FB08FABE" box="[159,1278,1260,1282]" gridrow="11" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB63E0FED5FABE" box="[159,291,1260,1282]" gridcol="0" gridrow="11" pageId="6" pageNumber="521">First band</th>
|
||
<td id="323841540527002848A463E0FD8AFABE" box="[480,636,1260,1282]" gridcol="1" gridrow="11" pageId="6" pageNumber="521">15</td>
|
||
<td id="32384154052700284A6563E0FC4BFABE" box="[801,957,1260,1282]" gridcol="2" gridrow="11" pageId="6" pageNumber="521">15</td>
|
||
<td id="32384154052700284D2763E0FB08FABE" box="[1123,1278,1260,1282]" gridcol="3" gridrow="11" pageId="6" pageNumber="521">13</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB6219FB08FA97" box="[159,1278,1301,1323]" gridrow="12" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB6219FED5FA97" box="[159,291,1301,1323]" gridcol="0" gridrow="12" pageId="6" pageNumber="521">Broad base</th>
|
||
<td id="323841540527002848A46219FD8AFA97" box="[480,636,1301,1323]" gridcol="1" gridrow="12" pageId="6" pageNumber="521">10</td>
|
||
<td id="32384154052700284A656219FC4BFA97" box="[801,957,1301,1323]" gridcol="2" gridrow="12" pageId="6" pageNumber="521">10.5</td>
|
||
<td id="32384154052700284D276219FB08FA97" box="[1123,1278,1301,1323]" gridcol="3" gridrow="12" pageId="6" pageNumber="521">11</td>
|
||
</tr>
|
||
<tr id="71E928280527002849DB6232FB08FAE8" box="[159,1278,1342,1364]" gridrow="13" pageId="6" pageNumber="521">
|
||
<th id="323841540527002849DB6232FED5FAE8" box="[159,291,1342,1364]" gridcol="0" gridrow="13" pageId="6" pageNumber="521">Broad vert</th>
|
||
<td id="323841540527002848A46232FD8AFAE8" box="[480,636,1342,1364]" gridcol="1" gridrow="13" pageId="6" pageNumber="521">4.5</td>
|
||
<td id="32384154052700284A656232FC4BFAE8" box="[801,957,1342,1364]" gridcol="2" gridrow="13" pageId="6" pageNumber="521">4.5</td>
|
||
<td id="32384154052700284D276232FB08FAE8" box="[1123,1278,1342,1364]" gridcol="3" gridrow="13" pageId="6" pageNumber="521">6</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0527FFD149836294FAE2FA45" blockId="6.[151,1437,1428,2033]" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD149836294FEBCFA0C" bold="true" box="[199,330,1432,1456]" italics="true" pageId="6" pageNumber="521">Hemipenis.</emphasis>
|
||
In everted position, the organ is single, not forked, stout, and reaches the level of the 10
|
||
<superScript id="38AC87220527FFD14C696298FACFFA1E" attach="left" box="[1325,1337,1428,1442]" fontSize="6" pageId="6" pageNumber="521">th</superScript>
|
||
SC. It is smooth along a short distance proximally, with a fold near its base, then entirely densely covered with short spines distally, with spines longer at the botton of the spinose part; tip smooth; sulcus straight, not prominent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10527FFD14983610FFB94F8A5" pageId="6" pageNumber="521" type="distribution">
|
||
<paragraph id="CF662A6A0527FFD14983610FFE1EF911" blockId="6.[151,1437,1428,2033]" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD14983610FFDB1F9A1" box="[199,583,1539,1565]" italics="true" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD14983610FFEACF9A0" bold="true" box="[199,346,1539,1564]" italics="true" pageId="6" pageNumber="521">Distribution.</emphasis>
|
||
<taxonomicName id="08D951E90527FFD1482A6108FDB1F9A1" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[366,583,1540,1565]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">Lycodon gibsonae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="E69E4B030527FFD14B1F6108FD2DF9A0" box="[603,731,1540,1564]" pageId="6" pageNumber="521" rank="species">
|
||
<emphasis id="FDADF6780527FFD14B1F6108FD2DF9A0" bold="true" box="[603,731,1540,1564]" pageId="6" pageNumber="521">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is at present known only from the Sankambaeng (or Sankamphaeng) Range, a short mountain range of up to
|
||
<quantity id="0821878F0527FFD14A576124FC99F9FC" box="[787,879,1576,1601]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="521" unit="m" value="1350.0">1,350 m</quantity>
|
||
a.s.l. in elevation which marks the south-western edge of
|
||
<collectingRegion id="0D1DE4880527FFD149BC6140FEB0F9D9" box="[248,326,1612,1637]" country="Thailand" name="Nakhon Ratchasima" pageId="6" pageNumber="521">Khorat</collectingRegion>
|
||
Plateau, Province of
|
||
<collectingRegion id="0D1DE4880527FFD14B016140FCD8F9D9" box="[581,814,1612,1637]" country="Thailand" name="Nakhon Ratchasima" pageId="6" pageNumber="521">Nakhon Ratchasima</collectingRegion>
|
||
,
|
||
<collectingCountry id="B7CE6AFA0527FFD14A786140FC55F9D9" box="[828,931,1612,1637]" name="Thailand" pageId="6" pageNumber="521">Thailand</collectingCountry>
|
||
. This mountain range, as well as the Dong Phaya Yen Mountains that edge the western limit of the
|
||
<collectingRegion id="0D1DE4880527FFD14A4F617CFCAFF935" box="[779,857,1648,1673]" country="Thailand" name="Nakhon Ratchasima" pageId="6" pageNumber="521">Khorat</collectingRegion>
|
||
Plateau, receive heavy rainfall and are still covered with tropical evergreen forest.
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0527FFD1498361BBFB94F8A5" blockId="6.[151,1437,1428,2033]" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD1498361BBFE66F973" bold="true" box="[199,400,1719,1744]" italics="true" pageId="6" pageNumber="521">Ecological notes.</emphasis>
|
||
The specimens were collected in the relatively cool evergreen forest covering the southwestern edges of the
|
||
<collectingRegion id="0D1DE4880527FFD148C161D0FE25F949" box="[389,467,1756,1781]" country="Thailand" name="Nakhon Ratchasima" pageId="6" pageNumber="521">Khorat</collectingRegion>
|
||
Plateau (
|
||
<bibRefCitation id="AB48579B0527FFD14B7861D0FCB6F948" author="Inger, R. F. & Colwell, R. K." box="[572,832,1756,1781]" pageId="6" pageNumber="521" pagination="229 - 253" refId="ref6449" refString="Inger, R. F. & Colwell, R. K. (1977) Organization of contiguous communities of amphibians and reptiles in Thailand. Ecological Monographs, 47, 229 - 253. https: // doi. org / 10.2307 / 1942516" type="journal article" year="1977">Inger & Colwell, 1977</bibRefCitation>
|
||
). As with other species of the genus, they were found to be terrestrial and nocturnal. Nothing else is known about the biology of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C379E10527FFD049836028FA6BFBAC" lastPageId="7" lastPageNumber="522" pageId="6" pageNumber="521" type="description">
|
||
<paragraph id="CF662A6A0527FFD149836028FA95F870" blockId="6.[151,1437,1428,2033]" pageId="6" pageNumber="521">
|
||
<emphasis id="FDADF6780527FFD149836028FEA9F880" bold="true" box="[199,351,1828,1852]" italics="true" pageId="6" pageNumber="521">Comparisons.</emphasis>
|
||
It should be noted that, since no females of
|
||
<taxonomicName id="08D951E90527FFD14A186028FBD5F881" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[860,1059,1828,1853]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780527FFD14A186028FBD5F881" box="[860,1059,1828,1853]" italics="true" pageId="6" pageNumber="521">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030527FFD14D696028FB68F880" box="[1069,1182,1828,1852]" pageId="6" pageNumber="521" rank="species">
|
||
<emphasis id="FDADF6780527FFD14D696028FB68F880" bold="true" box="[1069,1182,1828,1852]" pageId="6" pageNumber="521">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are known, only male characteristics were used in the comparisons.
|
||
<taxonomicName id="08D951E90527FFD14BC76044FCB0F8DD" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[643,838,1864,1889]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780527FFD14BC76044FCB0F8DD" box="[643,838,1864,1889]" italics="true" pageId="6" pageNumber="521">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030527FFD14A0A6044FC4AF8DC" box="[846,956,1864,1888]" pageId="6" pageNumber="521" rank="species">
|
||
<emphasis id="FDADF6780527FFD14A0A6044FC4AF8DC" bold="true" box="[846,956,1864,1888]" pageId="6" pageNumber="521">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from most other species of the genus
|
||
<taxonomicName id="08D951E90527FFD149D36060FF05F839" box="[151,243,1900,1925]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="genus">
|
||
<emphasis id="FDADF6780527FFD149D36060FF05F839" box="[151,243,1900,1925]" italics="true" pageId="6" pageNumber="521">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
, except those of the
|
||
<taxonomicName id="08D951E90527FFD1488F6061FDB2F838" authorityName="Anderson" authorityYear="1879" box="[459,580,1900,1924]" class="Reptilia" family="Colubridae" genus="Ophites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780527FFD1488F6061FDB2F838" box="[459,580,1900,1924]" italics="true" pageId="6" pageNumber="521">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
-group as well as
|
||
<taxonomicName id="08D951E90527FFD14BBA6061FC7CF839" authorityName="Boie" authorityYear="1827" box="[766,906,1900,1925]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="FDADF6780527FFD14BBA6061FC7CF839" box="[766,906,1900,1925]" italics="true" pageId="6" pageNumber="521">L. subcinctus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="08D951E90527FFD14AD26061FC1CF839" authorityName="Wostl, Hamidy, Kurniawan & E.N.Smith" authorityYear="2017" box="[918,1002,1900,1925]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="sidiki">
|
||
<emphasis id="FDADF6780527FFD14AD26061FC1CF839" box="[918,1002,1900,1925]" italics="true" pageId="6" pageNumber="521">L. sidiki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="08D951E90527FFD14AB36061FB9AF839" authorityName="Lanza" authorityYear="1999" box="[1015,1132,1900,1925]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="FDADF6780527FFD14AB36061FB9AF839" box="[1015,1132,1900,1925]" italics="true" pageId="6" pageNumber="521">L. dumerili</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="08D951E90527FFD14DE16061FAFEF839" box="[1189,1288,1900,1925]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="butleri">
|
||
<emphasis id="FDADF6780527FFD14DE16061FAFEF839" box="[1189,1288,1900,1925]" italics="true" pageId="6" pageNumber="521">L. butleri</emphasis>
|
||
</taxonomicName>
|
||
, by having its loreal contacting the eye.
|
||
<taxonomicName id="08D951E90527FFD148EB609CFD82F815" authorityName="Vogel & David" authorityYear="2019" box="[431,628,1936,1961]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gibsonae">
|
||
<emphasis id="FDADF6780527FFD148EB609CFD82F815" box="[431,628,1936,1961]" italics="true" pageId="6" pageNumber="521">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="08D951E90527FFD14A41609DFC62F815" authorityName="Boie" authorityYear="1827" box="[773,916,1936,1961]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="FDADF6780527FFD14A41609DFC62F815" box="[773,916,1936,1961]" italics="true" pageId="6" pageNumber="521">L. subcinctus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="08D951E90527FFD14A8F609DFBD4F815" authorityName="Wostl, Hamidy, Kurniawan & E.N.Smith" authorityYear="2017" box="[971,1058,1936,1961]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="sidiki">
|
||
<emphasis id="FDADF6780527FFD14A8F609DFBD4F815" box="[971,1058,1936,1961]" italics="true" pageId="6" pageNumber="521">L. sidiki</emphasis>
|
||
</taxonomicName>
|
||
by having a preocular (vs. absent) and from
|
||
<taxonomicName id="08D951E90527FFD149BA60B9FE83F871" authorityName="Lanza" authorityYear="1999" box="[254,373,1972,1997]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="FDADF6780527FFD149BA60B9FE83F871" box="[254,373,1972,1997]" italics="true" pageId="6" pageNumber="521">L. dumerili</emphasis>
|
||
</taxonomicName>
|
||
in having 8 infralabials (vs. 9) and by the lower number of subcaudals (91 or 92 vs. 111–120).
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0527FFD0498360D4FD49FE6C" blockId="6.[151,1437,1428,2033]" lastBlockId="7.[151,1437,151,1040]" lastPageId="7" lastPageNumber="522" pageId="6" pageNumber="521">
|
||
<taxonomicName id="08D951E90527FFD1498360D4FE65F84D" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[199,403,2008,2033]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780527FFD1498360D4FE65F84D" box="[199,403,2008,2033]" italics="true" pageId="6" pageNumber="521">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030527FFD148D860D4FDF9F84C" box="[412,527,2008,2032]" pageId="6" pageNumber="521" rank="species">
|
||
<emphasis id="FDADF6780527FFD148D860D4FDF9F84C" bold="true" box="[412,527,2008,2032]" pageId="6" pageNumber="521">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
belongs to the group of
|
||
<taxonomicName id="08D951E90527FFD14A6360D4FC04F84C" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[807,1010,2008,2033]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780527FFD14A6360D4FC04F84C" box="[807,1010,2008,2033]" italics="true" pageId="6" pageNumber="521">Lycodon fasciatus</emphasis>
|
||
</taxonomicName>
|
||
. It differs from
|
||
<taxonomicName id="08D951E90527FFD14DEE60D5FADBF84C" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[1194,1325,2008,2032]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="521" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780527FFD14DEE60D5FADBF84C" box="[1194,1325,2008,2032]" italics="true" pageId="6" pageNumber="521">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
by (1) its much larger size (SVL of the largest male of
|
||
<taxonomicName id="08D951E90526FFD04BCF6795FCFBFF0C" authorityName="Vogel & David" authorityYear="2019" box="[651,781,151,176]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae">
|
||
<emphasis id="FDADF6780526FFD04BCF6795FCFBFF0C" box="[651,781,151,176]" italics="true" pageId="7" pageNumber="522">L. gibsonae</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="0821878F0526FFD04A58679BFC8CFF13" box="[796,890,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="9.059999999999999" pageId="7" pageNumber="522" unit="mm" value="906.0">906 mm</quantity>
|
||
, SVL of the largest specimen among
|
||
<specimenCount id="D9DFE1E30526FFD04C5E679BFA88FF0C" box="[1306,1406,151,176]" pageId="7" pageNumber="522" type="male">26 males</specimenCount>
|
||
of
|
||
<taxonomicName id="08D951E90526FFD049D367B1FEEAFF68" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[151,284,188,212]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD049D367B1FEEAFF68" box="[151,284,188,212]" italics="true" pageId="7" pageNumber="522">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="0821878F0526FFD0486B67B0FE62FF68" box="[303,404,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="6.74" pageId="7" pageNumber="522" unit="mm" value="674.0">674 mm</quantity>
|
||
), (2) the number of ventrals (223–226 vs.
|
||
<quantity id="0821878F0526FFD04ACF67B0FBFBFF69" box="[907,1037,188,213]" metricMagnitude="0" metricUnit="m" metricValue="5.2324" metricValueMax="5.4102" metricValueMin="5.0546" pageId="7" pageNumber="522" unit="in" value="206.0" valueMax="213.0" valueMin="199.0">199–213 in</quantity>
|
||
<specimenCount id="D9DFE1E30526FFD04D5C67B0FB77FF69" box="[1048,1153,188,213]" pageId="7" pageNumber="522" type="male">25 males</specimenCount>
|
||
of
|
||
<taxonomicName id="08D951E90526FFD04DEA67B1FAC5FF68" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[1198,1331,188,212]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD04DEA67B1FAC5FF68" box="[1198,1331,188,212]" italics="true" pageId="7" pageNumber="522">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
), (3) the number of subcaudals (91 or
|
||
<quantity id="0821878F0526FFD048B767ECFDC6FF44" box="[499,560,223,248]" metricMagnitude="0" metricUnit="m" metricValue="2.3368" pageId="7" pageNumber="522" unit="in" value="92.0">92 in</quantity>
|
||
<specimenCount id="D9DFE1E30526FFD04B7F67ECFD3FFF44" box="[571,713,223,248]" pageId="7" pageNumber="522" type="generic">2 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="08D951E90526FFD04BB267D3FC33FF44" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[758,965,223,248]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD04BB267D3FC33FF44" box="[758,965,223,248]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04A9567ECFBB1FF44" box="[977,1095,224,248]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04A9567ECFBB1FF44" bold="true" box="[977,1095,224,248]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
vs.
|
||
<quantity id="0821878F0526FFD04D3967ECFB12FF44" box="[1149,1252,223,248]" metricMagnitude="0" metricUnit="m" metricValue="2.0827999999999998" metricValueMax="2.286" metricValueMin="1.8796" pageId="7" pageNumber="522" unit="in" value="82.0" valueMax="90.0" valueMin="74.0">74–90 in</quantity>
|
||
<specimenCount id="D9DFE1E30526FFD04DAB67ECFAA1FF44" box="[1263,1367,223,248]" pageId="7" pageNumber="522" type="male">20 males</specimenCount>
|
||
of
|
||
<taxonomicName id="08D951E90526FFD04CC167EDFF0DFEA0" baseAuthorityName="Anderson" baseAuthorityYear="1879" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD04CC167EDFF0DFEA0" italics="true" pageId="7" pageNumber="522">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
), (4) the number of bands on the body (17 or 18 vs.
|
||
<quantity id="0821878F0526FFD04A016608FC5EFEA1" box="[837,936,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" metricValueMax="10.922" metricValueMin="4.826" pageId="7" pageNumber="522" unit="in" value="31.0" valueMax="43.0" valueMin="19.0">19–43 in</quantity>
|
||
<taxonomicName id="08D951E90526FFD04AEB6608FB8FFEA0" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[943,1145,260,285]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" sensu="stricto" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD04AEB6608FB8FFEA0" box="[943,1145,260,285]" italics="true" pageId="7" pageNumber="522">Lycodon fasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04DC46609FAF9FEA1" box="[1152,1295,260,285]" pageId="7" pageNumber="522" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
), and (5) the pattern of the bands on the posterior part of the body: in
|
||
<taxonomicName id="08D951E90526FFD04A4B6625FC67FEFC" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[783,913,295,320]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD04A4B6625FC67FEFC" box="[783,913,295,320]" italics="true" pageId="7" pageNumber="522">L. gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FDADF6780526FFD04ADC6624FBE4FEFC" bold="true" box="[920,1042,296,320]" pageId="7" pageNumber="522">
|
||
<taxonomicNameLabel id="E69E4B030526FFD04ADC6624FBFAFEFC" box="[920,1036,296,320]" pageId="7" pageNumber="522" rank="species">spec. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
bands remain well defined, largely cream and not more strongly speckled with dark brown than anterior bands whereas, in
|
||
<taxonomicName id="08D951E90526FFD04D3B6641FAF4FED8" authorityName="Vogel & David" authorityYear="2019" box="[1151,1282,332,356]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD04D3B6641FAF4FED8" box="[1151,1282,332,356]" italics="true" pageId="7" pageNumber="522">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
, bands of the posterior part of the body and tail are much more strongly dark speckled than anterior bands, up to turning nearly entirely brown. There are also a few other minor points, which will be discussed elsewhere in a review of the
|
||
<taxonomicName id="08D951E90526FFD049D366BBFE97FE6C" authorityName="Vogel & David" authorityYear="2019" box="[151,353,439,464]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD049D366BBFE97FE6C" box="[151,353,439,464]" italics="true" pageId="7" pageNumber="522">Lycodon fasciatus</emphasis>
|
||
</taxonomicName>
|
||
complex (Vogel
|
||
<emphasis id="FDADF6780526FFD04B6466B5FDAFFE6C" box="[544,601,439,464]" italics="true" pageId="7" pageNumber="522">et al.</emphasis>
|
||
in prep).
|
||
</paragraph>
|
||
<paragraph id="CF662A6A0526FFD0498366D0FA6BFBAC" blockId="7.[151,1437,151,1040]" pageId="7" pageNumber="522">
|
||
Regarding other species of the
|
||
<taxonomicName id="08D951E90526FFD04B5B66D0FD1FFE48" authorityName="Vogel & David" authorityYear="2019" box="[543,745,476,501]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="FDADF6780526FFD04B5B66D0FD1FFE48" box="[543,745,476,501]" italics="true" pageId="7" pageNumber="522">Lycodon fasciatus</emphasis>
|
||
</taxonomicName>
|
||
group,
|
||
<taxonomicName id="08D951E90526FFD04A7966D0FBFEFE49" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[829,1032,476,501]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD04A7966D0FBFEFE49" box="[829,1032,476,501]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04D4A66D0FB76FE48" box="[1038,1152,476,500]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04D4A66D0FB76FE48" bold="true" box="[1038,1152,476,500]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="08D951E90526FFD04C5666D1FA8CFE49" box="[1298,1402,476,501]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="butleri">
|
||
<emphasis id="FDADF6780526FFD04C5666D1FA8CFE49" box="[1298,1402,476,501]" italics="true" pageId="7" pageNumber="522">L. butleri</emphasis>
|
||
</taxonomicName>
|
||
by the number of bands on the body, less than 19 vs. 30 or more in
|
||
<taxonomicName id="08D951E90526FFD04AC8650DFC01FDA4" box="[908,1015,511,536]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="butleri">
|
||
<emphasis id="FDADF6780526FFD04AC8650DFC01FDA4" box="[908,1015,511,536]" italics="true" pageId="7" pageNumber="522">L. butleri</emphasis>
|
||
</taxonomicName>
|
||
, and by the pattern of the venter. In
|
||
<taxonomicName id="08D951E90526FFD049D36528FE94FD81" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[151,354,548,573]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD049D36528FE94FD81" box="[151,354,548,573]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD0482D6528FE2DFD80" box="[361,475,548,572]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD0482D6528FE2DFD80" bold="true" box="[361,475,548,572]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, bands are more conspicuous posteriorly than on the anterior part of the venter, where the bands are faint, while in
|
||
<taxonomicName id="08D951E90526FFD0489F6545FDB3FDDC" box="[475,581,583,608]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="butleri">
|
||
<emphasis id="FDADF6780526FFD0489F6545FDB3FDDC" box="[475,581,583,608]" italics="true" pageId="7" pageNumber="522">L. butleri</emphasis>
|
||
</taxonomicName>
|
||
the bands are more conspicuous anteriorly, while the venter is turning dark posteriorly with faint bands.
|
||
<taxonomicName id="08D951E90526FFD0489B6560FD5AFD39" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[479,684,620,645]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD0489B6560FD5AFD39" box="[479,684,620,645]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04BF06560FCDEFD38" box="[692,808,620,644]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04BF06560FCDEFD38" bold="true" box="[692,808,620,644]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="08D951E90526FFD04A846561FB98FD39" box="[960,1134,620,645]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="carvenicolus">
|
||
<emphasis id="FDADF6780526FFD04A846561FB98FD39" box="[960,1134,620,645]" italics="true" pageId="7" pageNumber="522">L. carvenicolus</emphasis>
|
||
</taxonomicName>
|
||
by the number of ventrals and subcaudals of the males (223–226 against 245 and 91 or 92 against 113 respectively), and in the number of bands on the body (17 or 18 against 36–45). The main difference with
|
||
<taxonomicName id="08D951E90526FFD04AF265B9FB75FD71" box="[950,1155,692,717]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="FDADF6780526FFD04AF265B9FB75FD71" box="[950,1155,692,717]" italics="true" pageId="7" pageNumber="522">L. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
is the fact that the loreal scale is touching the eye in
|
||
<taxonomicName id="08D951E90526FFD0489065DBFD57FD4C" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[468,673,727,752]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD0489065DBFD57FD4C" box="[468,673,727,752]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04BED65D4FCEBFD4C" box="[681,797,728,752]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04BED65D4FCEBFD4C" bold="true" box="[681,797,728,752]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
whereas it is separate from it in
|
||
<taxonomicName id="08D951E90526FFD04DD065D5FA97FD4C" box="[1172,1377,727,752]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="FDADF6780526FFD04DD065D5FA97FD4C" box="[1172,1377,727,752]" italics="true" pageId="7" pageNumber="522">L. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
. The venter is banded in
|
||
<taxonomicName id="08D951E90526FFD0483D65F0FDB0FCA9" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[377,582,764,789]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD0483D65F0FDB0FCA9" box="[377,582,764,789]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04B0B65F0FD35FCA8" box="[591,707,764,788]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04B0B65F0FD35FCA8" bold="true" box="[591,707,764,788]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
whereas it is not in
|
||
<emphasis id="FDADF6780526FFD04AEA65F1FAAFFCA9" box="[942,1369,764,789]" italics="true" pageId="7" pageNumber="522">
|
||
<taxonomicName id="08D951E90526FFD04AEA65F1FB76FCA9" box="[942,1152,764,789]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="cardamomensis">L. cardamomensis</taxonomicName>
|
||
.
|
||
<taxonomicName id="08D951E90526FFD04DC865F0FAAFFCA9" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[1164,1369,764,789]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">Lycodon gibsonae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04C2665F0FF3EFC84" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04C2665F0FF3EFC84" bold="true" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="08D951E90526FFD04819642DFE1EFC84" authorityName="Vogel & Luo" authorityYear="2011" box="[349,488,799,824]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gongshan">
|
||
<emphasis id="FDADF6780526FFD04819642DFE1EFC84" box="[349,488,799,824]" italics="true" pageId="7" pageNumber="522">L. gongshan</emphasis>
|
||
</taxonomicName>
|
||
by its shorter tail (0.198 against 0.231–0.255), by the lower number of bands on the body (17 or 18 against 37–41), and more infralabials (10 against 8 or 9). It differs from
|
||
<taxonomicName id="08D951E90526FFD04D266449FB17FCE0" authorityName="Vogel & David" authorityYear="2010" box="[1122,1249,837,860]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="synaptor">
|
||
<emphasis id="FDADF6780526FFD04D266449FB17FCE0" box="[1122,1249,837,860]" italics="true" pageId="7" pageNumber="522">L. synaptor</emphasis>
|
||
</taxonomicName>
|
||
by the condition of the loreal scale, entering orbit in
|
||
<taxonomicName id="08D951E90526FFD04B70646BFCF7FC3C" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[564,769,871,896]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD04B70646BFCF7FC3C" box="[564,769,871,896]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04A4E6464FC88FC3C" box="[778,894,872,896]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04A4E6464FC88FC3C" bold="true" box="[778,894,872,896]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
vs. separate from it in
|
||
<taxonomicName id="08D951E90526FFD04DCF6465FAFCFC3C" authorityName="Vogel & David" authorityYear="2010" box="[1163,1290,873,896]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="synaptor">
|
||
<emphasis id="FDADF6780526FFD04DCF6465FAFCFC3C" box="[1163,1290,873,896]" italics="true" pageId="7" pageNumber="522">L. synaptor</emphasis>
|
||
</taxonomicName>
|
||
. It has more infralabials (10 against 8), a lower number of bands on the body (17 or 18 against 30 or 31), and a different shape of the bands (width of the first band at the base 10 or 11 against 3 ventrals). Finally,
|
||
<taxonomicName id="08D951E90526FFD04D1064A3FAD7FC74" authority="Vogel & David, 2019" authorityName="Vogel & David" authorityYear="2019" box="[1108,1313,943,968]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="gibsonae" status="sp. nov.">
|
||
<emphasis id="FDADF6780526FFD04D1064A3FAD7FC74" box="[1108,1313,943,968]" italics="true" pageId="7" pageNumber="522">Lycodon gibsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E69E4B030526FFD04C6D64BCFA6AFC74" box="[1321,1436,944,968]" pageId="7" pageNumber="522" rank="species">
|
||
<emphasis id="FDADF6780526FFD04C6D64BCFA6AFC74" bold="true" box="[1321,1436,944,968]" pageId="7" pageNumber="522">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="08D951E90526FFD0486164D9FE15FC51" authorityName="Zhang, Jiang, Vogel & Rao" authorityYear="2011" box="[293,483,980,1005]" class="Reptilia" family="Colubridae" genus="Lycodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="522" phylum="Chordata" rank="species" species="liuchengchaoi">
|
||
<emphasis id="FDADF6780526FFD0486164D9FE15FC51" box="[293,483,980,1005]" italics="true" pageId="7" pageNumber="522">L. liuchengchaoi</emphasis>
|
||
</taxonomicName>
|
||
by the number of ventrals (223–226 vs. 202–204) and subcaudals in males (91or 92 vs. 68), the number of infralabials (10 against 7–9), and the number of bands on the body (17 or 18 against 40–47).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |