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<document id="CBFA8E9F96D4F8053D0F375788E6CFF3" ID-DOI="10.18590/euscorpius.2003.vol2003.iss10.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="4672850" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617910371128" checkinUser="carolina" docAuthor="Kovařík, František" docDate="2003" docId="4703879F4513270DFC98A8C620E68693" docLanguage="en" docName="Euscorpius.10.1-19.pdf" docOrigin="Euscorpius 10 (10)" docSource="https://mds.marshall.edu/euscorpius/vol2003/iss10/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Isometrus (Reddyanus) problematicus Kovařík 2003, sp. n." docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="BB3AFFE7451F2700FFFDAE1023128705" masterDocTitle="A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia" masterLastPageNumber="19" masterPageNumber="1" pageNumber="11" updateTime="1698902588460" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A2821CB659979C1A87F5ADB224AFA393">A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia</mods:title>
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<mods:title id="B01EAF1C3967579876D5B899D2E14414">Euscorpius</mods:title>
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<subSubSection id="87B065024513270CFC98A8C627FB8017" pageId="12" pageNumber="11" type="nomenclature">
<paragraph id="CF1536894513270CFC98A8C6266881F5" blockId="12.[869,1402,1750,1810]" box="[869,1402,1750,1777]" pageId="12" pageNumber="11">
<heading id="945D81E54513270CFC98A8C6266881F5" box="[869,1402,1750,1777]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="11" reason="4">
<emphasis id="FDDEEA9B4513270CFC98A8C6266881F5" bold="true" box="[869,1402,1750,1777]" pageId="12" pageNumber="11">
<taxonomicName id="08AA4D0A4513270CFC98A8C6262281F4" authority="Kovařík, 2003" authorityName="Kovařík" authorityYear="2003" box="[869,1328,1750,1777]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="problematicus" status="sp. nov." subGenus="Reddyanus">
<emphasis id="FDDEEA9B4513270CFC98A8C6262281F4" bold="true" box="[869,1328,1750,1777]" italics="true" pageId="12" pageNumber="11">Isometrus (Reddyanus) problematicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6ED57E04513270CFAC5A8C6266881F5" box="[1336,1402,1750,1776]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="CF1536894513270CFC0BA8EB27FB8017" blockId="12.[869,1402,1750,1810]" box="[1014,1257,1787,1810]" pageId="12" pageNumber="11">
(Figs 1 and 6,
<tableCitation id="822803324513270CFB72A8EB27CC8017" box="[1167,1246,1787,1810]" captionStart="Table 1" captionStartId="9.[192,257,1030,1054]" captionText="Table 1: Measurements (mm) of the type specimens of the new Isometrus species." pageId="12" pageNumber="11">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="87B065024513270DFCC3A92E21DB8672" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11" type="materials_examined">
<paragraph id="CF1536894513270CFCC3A92E27A78073" blockId="12.[830,1440,1854,1910]" pageId="12" pageNumber="11">
<typeStatus id="1011882B4513270CFCC3A92E20658050" box="[830,887,1854,1877]" pageId="12" pageNumber="11">TYPE</typeStatus>
LOCALITY AND
<materialsCitation id="7FC23CD44513270CFBC9A95227728073" country="India" location="Pondichery State" municipality="Karaikal" pageId="12" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="1011882B4513270CFBC9A952277B8050" box="[1076,1129,1858,1877]" pageId="12" pageNumber="11">TYPE</typeStatus>
REPOSITORY.
<emphasis id="FDDEEA9B4513270CFAF3A92E26598053" bold="true" box="[1294,1355,1854,1878]" pageId="12" pageNumber="11">
<collectingCountry id="B7BD76194513270CFAF3A92E26598053" box="[1294,1355,1854,1878]" name="India" pageId="12" pageNumber="11">India</collectingCountry>
</emphasis>
, south,
<location id="CA7560524513270CFCC3A94F20E48073" LSID="urn:lsid:plazi:treatment:4703879F4513270DFC98A8C620E68693:CA7560524513270CFCC3A94F20E48073" box="[830,1014,1887,1910]" country="India" municipality="Karaikal" name="Pondichery State" pageId="12" pageNumber="11">Pondichery State</location>
,
<collectingMunicipality id="2F71ACF34513270CFBFCA94F27728073" box="[1025,1120,1887,1910]" pageId="12" pageNumber="11">Karaikal</collectingMunicipality>
</materialsCitation>
;
<collectionCode id="A9BBAE4C4513270CFB96A94F27A38073" box="[1131,1201,1887,1910]" pageId="12" pageNumber="11">FKCP</collectionCode>
.
</paragraph>
<paragraph id="CF1536894512270DFF3DAED721DB8672" blockId="13.[192,802,198,437]" pageId="13" pageNumber="12">
<materialsCitation id="7FC23CD44512270DFF3DAED721D78672" collectingDate="1957-09" collectingDateMax="1958-11" collectingDateMin="1957-09" collectionCode="FKCP" collectorName="Karaikal, T. R. S. N. &amp; P. Susai Nathan &amp; Nilgiri Hills &amp; Kallas" country="India" location="Coimbatore" municipality="Pondichery State" pageId="13" pageNumber="12" specimenCount="3" specimenCount-female="3" stateProvince="Tamil Nadu" typeStatus="holotype">
TYPE MATERIAL.
<emphasis id="FDDEEA9B4512270DFE60AED622C887DB" bold="true" box="[413,474,198,222]" pageId="13" pageNumber="12">
<collectingCountry id="B7BD76194512270DFE60AED622C887DB" box="[413,474,198,222]" name="India" pageId="13" pageNumber="12">India</collectingCountry>
</emphasis>
, south,
<collectingMunicipality id="2F71ACF34512270DFDAEAED7200F87DB" box="[595,797,199,222]" pageId="13" pageNumber="12">Pondichery State</collectingMunicipality>
,
<collectorName id="625F535F4512270DFF3DAEF6229C87F8" box="[192,398,230,253]" pageId="13" pageNumber="12">Karaikal, T.R.S.N.</collectorName>
coll., 2002,
<specimenCount id="D9ACFD004512270DFDDBAEF6215A87F8" box="[550,584,230,253]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
(
<typeStatus id="1011882B4512270DFDA1AEF621AD87F8" box="[604,703,230,253]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
),
<collectionCode id="A9BBAE4C4512270DFD25AEF6200C87F8" box="[728,798,230,253]" pageId="13" pageNumber="12">FKCP</collectionCode>
; south,
<collectingRegion id="0D6EF86B4512270DFEECAF1422F7861E" box="[273,485,260,283]" country="India" name="Tamil Nadu" pageId="13" pageNumber="12">Tamil Nadu State</collectingRegion>
,
<location id="CA7560524512270DFE07AF142169861E" LSID="urn:lsid:plazi:treatment:4703879F4513270DFC98A8C620E68693:CA7560524512270DFE07AF142169861E" box="[506,635,260,283]" country="India" municipality="Pondichery State" name="Coimbatore" pageId="13" pageNumber="12" stateProvince="Tamil Nadu">Coimbatore</location>
,
<date id="BB1410494512270DFD6DAF1421F9861E" box="[656,747,260,283]" pageId="13" pageNumber="12" value="1957-09">
<collectingDate id="AB50E9A14512270DFD6DAF1421F9861E" box="[656,747,260,283]" pageId="13" pageNumber="12" value="1957-09">IX.1957</collectingDate>
</date>
,
<specimenCount id="D9ACFD004512270DFCFDAF142030861E" box="[768,802,260,283]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
(
<typeStatus id="1011882B4512270DFF3AAF33223B863F" box="[199,297,291,314]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
), leg.
<collectorName id="625F535F4512270DFE7BAF33215D863F" box="[390,591,291,314]" pageId="13" pageNumber="12">P. Susai Nathan</collectorName>
, CASC (Stahnke colection);
<collectorName id="625F535F4512270DFEC7AF5222AD865C" box="[314,447,322,345]" pageId="13" pageNumber="12">Nilgiri Hills</collectorName>
,
<collectorName id="625F535F4512270DFE36AF522100865C" box="[459,530,322,345]" pageId="13" pageNumber="12">Kallas</collectorName>
,
<date id="BB1410494512270DFDE3AF52216A865C" box="[542,632,322,345]" pageId="13" pageNumber="12" value="1958-11">
<collectingDate id="AB50E9A14512270DFDE3AF52216A865C" box="[542,632,322,345]" pageId="13" pageNumber="12" value="1958-11">XI.1958</collectingDate>
</date>
,
<specimenCount id="D9ACFD004512270DFD79AF5221B4865C" box="[644,678,322,345]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
(
<typeStatus id="1011882B4512270DFD48AF522004865C" box="[693,790,322,345]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
), leg.
<collectorName id="625F535F4512270DFF13AF7022898672" box="[238,411,352,375]" pageId="13" pageNumber="12">P. Susai Nathan</collectorName>
, CASC (Stahnke colection)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87B065024512270DFF3DAF6F22D986B0" pageId="13" pageNumber="12" type="etymology">
<paragraph id="CF1536894512270DFF3DAF6F22D986B0" blockId="13.[192,802,198,437]" pageId="13" pageNumber="12">ETYMOLOGY. The name reflects past identification problems (see affinities).</paragraph>
</subSubSection>
<subSubSection id="87B065024512270DFF3DAFCB214685CC" pageId="13" pageNumber="12" type="diagnosis">
<paragraph id="CF1536894512270DFF3DAFCB214685CC" blockId="13.[192,802,475,713]" pageId="13" pageNumber="12">
DIAGNOSIS: Total length
<quantity id="08529B6C4512270DFE19AFCB212486F7" box="[484,566,475,498]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="12" unit="mm" value="22.0">22 mm</quantity>
(female). Pedipalps, legs and metasomal segments yellow to reddish brown with black spots. First metasomal segment bears 10 carinae, second through fourth segments bear eight carinae, fifth segment bears five carinae. Telson bulbous, without carinae. Subaculear tooth large, obliquely triangular, dorsally with three pairs of granules. Pectinal teeth number 912.
</paragraph>
</subSubSection>
<subSubSection id="87B065024512270DFF3DACFF20E68693" pageId="13" pageNumber="12" type="description">
<paragraph id="CF1536894512270DFF3DACFF223984D8" blockId="13.[192,803,751,1694]" pageId="13" pageNumber="12">
DESCRIPTION: The total length of the female
<typeStatus id="1011882B4512270DFD57ACFF201A8403" box="[682,776,751,774]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
is
<quantity id="08529B6C4512270DFF3DAD1E22028420" box="[192,272,782,805]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="12" unit="mm" value="22.0">22 mm</quantity>
. Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps, and numbers of pectinal teeth are given in
<tableCitation id="822803324512270DFDA0AD5B21BD8467" box="[605,687,843,866]" captionStart="Table 1" captionStartId="9.[192,257,1030,1054]" captionText="Table 1: Measurements (mm) of the type specimens of the new Isometrus species." pageId="13" pageNumber="12">Table 1</tableCitation>
. There are 12 teeth in the pecten. The
<typeStatus id="1011882B4512270DFE15AD7A215C8484" box="[488,590,874,897]" pageId="13" pageNumber="12" type="paratype">paratypes</typeStatus>
have 910 pectinal teeth. For the position and distribution of trichobothria on the chela of pedipalps see Fig. 6. The male is unknown.
</paragraph>
<paragraph id="CF1536894512270DFF3DADF422678373" blockId="13.[192,803,751,1694]" pageId="13" pageNumber="12">COLORATION: The chelicera is reticulated. The carapace and mesosoma are dominantly black with a weak yellow pattern. The femur, patella and chela of pedipalps, legs and metasomal segments are yellow to reddish brown with black spots.</paragraph>
<paragraph id="CF1536894512270DFF3DAA6E226A82AC" blockId="13.[192,803,751,1694]" pageId="13" pageNumber="12">MESOSOMA AND CARAPACE: The carapace is without carinae but bears large granules. The mesosoma bears one median carina and is granulated. The ventral side of the seventh segment bears four carinae and is granulated. METASOMA AND TELSON: The first segment bears 10 carinae, the second through fourth segments bear eight carinae, and the fifth segment bears five carinae. The telson is bulbous, without carinae. The subaculear tooth is large, obliquely triangular, dorsally with three pairs of granules (Fig. 1).</paragraph>
<paragraph id="CF1536894512270DFF3DABA021238147" blockId="13.[192,803,751,1694]" pageId="13" pageNumber="12">PEDIPALPS: The femur and patella of pedipalps bear complete carinae and are weakly granulated. The manus of pedipalps is smooth, with only traces of carinae and several granules. The sixth cutting edge of movable fingers bears one external granule.</paragraph>
<paragraph id="CF1536894512270DFF3DA85A2242819B" blockId="13.[192,803,751,1694]" pageId="13" pageNumber="12">LEGS: The femur and patella bear complete carinae and are granulated. The legs are hirsute and without bristlecombs.</paragraph>
<paragraph id="CF1536894512270DFF3DA8D420E68693" blockId="13.[192,802,1732,1908]" lastBlockId="13.[830,1441,199,406]" pageId="13" pageNumber="12">
AFFINITIES. The described features distinguish
<taxonomicName id="08AA4D0A4512270DFCEFA8D4224B81FF" authority="Kovařík, 2003" authorityName="Kovařík" authorityYear="2003" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="problematicus" status="sp. nov.">
<emphasis id="FDDEEA9B4512270DFCEFA8D4224B81FF" italics="true" pageId="13" pageNumber="12">I. problematicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6ED57E04512270DFE9CA8F3228981FF" box="[353,411,1763,1786]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
from all other species of the genus. They are recounted in the key below. The most similar species is
<taxonomicName id="08AA4D0A4512270DFED1A93021248032" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" box="[300,566,1824,1847]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="FDDEEA9B4512270DFED1A93022B38032" box="[300,417,1824,1847]" italics="true" pageId="13" pageNumber="12">I. basilicus</emphasis>
<bibRefCitation id="AB3B4B784512270DFE55A93021248032" author="KARSCH" box="[424,566,1824,1847]" pageId="13" pageNumber="12" pagination="97 - 136" refId="ref15475" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="B7BD76194512270DFD85A93021F18032" box="[632,739,1824,1847]" name="Sri Lanka" pageId="13" pageNumber="12">Sri Lanka</collectingCountry>
, from which
<taxonomicName id="08AA4D0A4512270DFEF4A92F22A88053" authority="Kovařík, 2003" authorityName="Kovařík" authorityYear="2003" box="[265,442,1855,1878]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="problematicus" status="sp. nov.">
<emphasis id="FDDEEA9B4512270DFEF4A92F22A88053" box="[265,442,1855,1878]" italics="true" pageId="13" pageNumber="12">I. problematicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6ED57E04512270DFE3FA92F22EF8053" box="[450,509,1855,1878]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
can be easily separated by smaller size and shorter/broader metasomal segments. One of the
<typeStatus id="1011882B4512270DFC46AED7273387DB" box="[955,1057,199,222]" pageId="13" pageNumber="12" type="paratype">paratypes</typeStatus>
was originally labeled
<emphasis id="FDDEEA9B4512270DFADFAED726B387DB" box="[1314,1441,199,222]" italics="true" pageId="13" pageNumber="12">
<taxonomicName id="08AA4D0A4512270DFADFAED7268F87DB" authorityName="Karsch" authorityYear="1879" box="[1314,1437,199,222]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="basilicus">I. basilicus</taxonomicName>
.
</emphasis>
The other
<typeStatus id="1011882B4512270DFC40AEF6270B87F8" box="[957,1049,230,253]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
was labeled as
<taxonomicName id="08AA4D0A4512270DFB1FAEF6268887F8" authorityName="Pocock" authorityYear="1899" box="[1250,1434,230,253]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="brachycentrus">
<emphasis id="FDDEEA9B4512270DFB1FAEF6268887F8" box="[1250,1434,230,253]" italics="true" pageId="13" pageNumber="12">I. brachycentrus</emphasis>
</taxonomicName>
, whose juveniles may be mistaken for
<taxonomicName id="08AA4D0A4512270DFB10AF1426B2861E" authority="Kovařík, 2003" authorityName="Kovařík" authorityYear="2003" box="[1261,1440,260,283]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="problematicus" status="sp. nov.">
<emphasis id="FDDEEA9B4512270DFB10AF1426B2861E" box="[1261,1440,260,283]" italics="true" pageId="13" pageNumber="12">I. problematicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6ED57E04512270DFCC3AF332069863F" box="[830,891,291,314]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
although they have markedly narrower manus of pedipalps and their legs and metasomal segments are conspicuously spotted in a way different from adults of
<taxonomicName id="08AA4D0A4512270DFCC3AF6F20FC8693" authorityName="Pocock" authorityYear="1899" box="[830,1006,383,406]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="brachycentrus">
<emphasis id="FDDEEA9B4512270DFCC3AF6F20FC8693" box="[830,1006,383,406]" italics="true" pageId="13" pageNumber="12">I. brachycentrus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>