611 lines
129 KiB
XML
611 lines
129 KiB
XML
<document id="F20D8EB227227912735D1DFE0EDA8B95" ID-DOI="10.11646/zootaxa.3722.2.7" ID-GBIF-Dataset="dc599a29-bc40-4802-8365-d090c8254627" ID-ISSN="1175-5326" ID-Zenodo-Dep="216795" ID-ZooBank="5D21B65C-1917-4513-BDF2-168835BEC884" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460691639014" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="BF5B3062FFDFFFE1F99D5C964233EAFB" docLanguage="en" docName="zt03722p266.pdf" docOrigin="Zootaxa 3722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteroscydmus Franz" docType="treatment" docVersion="8" lastPageNumber="255" masterDocId="4362481AFFD8FFEBF90A5D694329EC38" masterDocTitle="Revision of Neotropical genera Microraphes Franz, Heteroscydmus Franz and Mimoscydmus Franz (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="266" masterPageNumber="245" pageNumber="252" updateTime="1698351829303" updateUser="plazi">
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<mods:titleInfo id="DFC335E0627783124000B4BEA4750CD2">
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<mods:title id="59FCFEDFF5478F084AAC7800ECB863DA">Revision of Neotropical genera Microraphes Franz, Heteroscydmus Franz and Mimoscydmus Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="A5FEFC969129A5283BB4D266965E896A">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="26040AF9A22FE67933E595D90692BE21">Zootaxa</mods:title>
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<mods:part id="2D4B5ED220B13305B4F0AD5A9DA52360">
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<mods:date id="EE59D9650E86F52D014B50D70E3E2CC8">2013</mods:date>
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<mods:number id="48E964955A4BFA4FD97E9439DAF34D7A">3722</mods:number>
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<mods:identifier id="10B0160F8D86FD02F6ADF1AC6C46B3B3" type="DOI">10.11646/zootaxa.3722.2.7</mods:identifier>
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<treatment id="BF5B3062FFDFFFE1F99D5C964233EAFB" ID-DOI="http://doi.org/10.5281/zenodo.6154648" ID-GBIF-Taxon="119579879" ID-Zenodo-Dep="6154648" LSID="urn:lsid:plazi:treatment:BF5B3062FFDFFFE1F99D5C964233EAFB" httpUri="http://treatment.plazi.org/id/BF5B3062FFDFFFE1F99D5C964233EAFB" lastPageId="10" lastPageNumber="255" pageId="7" pageNumber="252">
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<subSubSection id="7FE8D2FFFFDFFFECF99D5C96479BEE64" pageId="7" pageNumber="252" type="nomenclature">
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<paragraph id="374D8174FFDFFFECF99D5C96428AEE22" blockId="7.[151,419,511,538]" box="[151,419,511,538]" pageId="7" pageNumber="252">
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<heading id="6C053618FFDFFFECF99D5C96428AEE22" bold="true" box="[151,419,511,538]" fontSize="11" level="1" pageId="7" pageNumber="252" reason="1">
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<taxonomicName id="F0F2FAF7FFDFFFECF99D5C96428AEE22" ID-CoL="8XLSJ" authority="Franz" authorityName="Franz" box="[151,419,511,538]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="252" phylum="Arthropoda" rank="genus">
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<emphasis id="05865D66FFDFFFECF99D5C96428AEE22" bold="true" box="[151,419,511,538]" pageId="7" pageNumber="252">
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<emphasis id="05865D66FFDFFFECF99D5C964278EE21" bold="true" box="[151,337,511,537]" italics="true" pageId="7" pageNumber="252">Heteroscydmus</emphasis>
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Franz
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="374D8174FFDFFFECF99D5F2E479BEE64" blockId="7.[151,1202,582,604]" box="[151,1202,582,604]" pageId="7" pageNumber="252">
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<taxonomicName id="F0F2FAF7FFDFFFECF99D5F2E42CEEE64" authority="Franz, 1980: 208" authorityName="Franz" authorityPageNumber="208" authorityYear="1980" box="[151,487,582,604]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="252" phylum="Arthropoda" rank="genus">
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<emphasis id="05865D66FFDFFFECF99D5F2E421BEE64" box="[151,306,583,604]" italics="true" pageId="7" pageNumber="252">Heteroscydmus</emphasis>
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Franz, 1980: 208
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</taxonomicName>
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.
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<typeStatus id="E8493FD6FFDFFFECF8FB5F2E410AEE64" box="[497,547,583,604]" pageId="7" pageNumber="252">Type</typeStatus>
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species:
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<taxonomicName id="F0F2FAF7FFDFFFECFB8A5F2E4707EE64" authority="Franz, 1980" authorityName="Franz" authorityYear="1980" box="[640,1070,582,604]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="252" phylum="Arthropoda" rank="species" species="yapacariensis">
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<emphasis id="05865D66FFDFFFECFB8A5F2E4087EE64" box="[640,942,583,604]" italics="true" pageId="7" pageNumber="252">Heteroscydmus yapacariensis</emphasis>
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Franz, 1980
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</taxonomicName>
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(monotypy).
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</paragraph>
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</subSubSection>
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<subSubSection id="7FE8D2FFFFDFFFE3F99D5FE04718E8D0" lastPageId="8" lastPageNumber="253" pageId="7" pageNumber="252" type="description">
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<paragraph id="374D8174FFDFFFECF99D5FE04095E869" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
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<emphasis id="05865D66FFDFFFECF99D5FE0425FEE9A" bold="true" box="[151,374,649,674]" pageId="7" pageNumber="252">Revised diagnosis.</emphasis>
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Male and female: head about as long as broad, with vertex not expanded dorso-caudad; occipital constriction only slightly narrower than vertex; thick and long bristles absent on head and prothorax; frontoclypeal groove absent; submentum not demarcated laterally from hypostomae by sutures; maxillary palpomere III strongly elongate; antennae with distinctly delimited club composed of antennomeres IX–XI; pronotum without lateral edges; base of pronotum with fine transverse groove, without pits and without sub-lateral carinae; basisternal part of prosternum much shorter than procoxal cavities; prosternum with rudimentary intercoxal process developed as a low (i.e., only slightly expanding ventrally) subtriangular expansion between procoxal cavities; prothoracic hypomeral ridges complete; pronotosternal sutures visible only along sides of basisternal part of prosternum; mesoventral intercoxal process long, narrow and moderately strongly expanding ventrally (but not keel-shaped); mesoventrite with asetose lateral impressions behind anterior ridge, without setose impressions; mesothorax without lateral foveae; mesocoxal projection without posterior lobe; metacoxae narrowly separated by metaventral intercoxal process composed of a pair of long spines; each elytron with two rudiments of asetose basal fovea. Male: aedeagus asymmetrical, with free parameres.
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</paragraph>
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<paragraph id="374D8174FFDFFFECF9CD593440FDE8A2" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
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<emphasis id="05865D66FFDFFFECF9CD5934425DE84E" bold="true" box="[199,372,1117,1142]" pageId="7" pageNumber="252">Redescription.</emphasis>
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Body of male (
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<figureCitation id="AFC99DF1FFDFFFECFB41593441B1E84E" box="[587,664,1117,1142]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="7" pageNumber="252">Fig. 2</figureCitation>
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) strongly convex, elongate and slender, with moderately long appendages, BL
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<quantity id="F00A2C91FFDFFFECF85859EB42D1E8A1" box="[338,504,1154,1178]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="7.3" metricValueMin="7.0" pageId="7" pageNumber="252" unit="mm" value="0.715" valueMax="0.73" valueMin="0.7">0.70–0.73 mm</quantity>
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; cuticle glossy, light brown, vestiture fine.
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</paragraph>
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<paragraph id="374D8174FFDFFFECF9CD59CC4654E976" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
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Head (
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<figureCitation id="AFC99DF1FFDFFFECF81D59CC4245E886" box="[279,364,1189,1214]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="7" pageNumber="252">Figs. 2</figureCitation>
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,
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<figureCitation id="AFC99DF1FFDFFFECF87459CF4291E886" box="[382,440,1190,1214]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">9–10</figureCitation>
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) with anterior part (in front of occipital constriction) about as long as broad, with moderately large eyes; occipital constriction (
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<figureCitation id="AFC99DF1FFDFFFECFB9259A0403DE8DA" box="[664,788,1225,1250]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Figs. 9–10</figureCitation>
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;
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<emphasis id="05865D66FFDFFFECFA1559A3406FE8D9" box="[799,838,1226,1249]" italics="true" pageId="7" pageNumber="252">occ</emphasis>
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) only slightly narrower than vertex; tempora (
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<figureCitation id="AFC99DF1FFDFFFECFC5B59A046BEE8DA" box="[1361,1431,1225,1250]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 9</figureCitation>
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;
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<emphasis id="05865D66FFDFFFECF99D5987439AE93D" box="[151,179,1262,1285]" italics="true" pageId="7" pageNumber="252">tm</emphasis>
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) long and slightly convergent caudad, without bristles; vertex (
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<figureCitation id="AFC99DF1FFDFFFECFAA0598440DDE93E" box="[938,1012,1261,1286]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 9</figureCitation>
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;
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<emphasis id="05865D66FFDFFFECFD0C59874733E93D" box="[1030,1050,1262,1285]" italics="true" pageId="7" pageNumber="252">vt</emphasis>
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) broader than long, convex, not projecting dorso-caudad; frons (
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<figureCitation id="AFC99DF1FFDFFFECF8F55878416DE912" box="[511,580,1297,1322]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 9</figureCitation>
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;
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<emphasis id="05865D66FFDFFFECFB585878414CE911" box="[594,613,1297,1321]" italics="true" pageId="7" pageNumber="252">fr</emphasis>
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) transverse and subtrapezoidal with anterior margin distinctly expanding in middle, posteriorly confluent with vertex; frontoclypeal groove absent; antennal insertions broadly separated.
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</paragraph>
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<paragraph id="374D8174FFDFFFECF9CD58304000EA1D" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
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Labrum transverse with rounded anterior margin. Mandibles (
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<figureCitation id="AFC99DF1FFDFFFECFA7D58304095E94A" box="[887,956,1369,1394]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 9</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFACD583040C0E949" box="[967,1001,1369,1393]" italics="true" pageId="7" pageNumber="252">md</emphasis>
|
||
) symmetrical, with slender and curved distal parts, one mesal tooth located sub-apically; prostheca in the studied specimen not visible. Each maxilla (
|
||
<figureCitation id="AFC99DF1FFDFFFECFC655814439DE982" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
) with subtriangular basistipes (
|
||
<figureCitation id="AFC99DF1FFDFFFECFB2658C841AFE982" box="[556,646,1441,1466]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFB9258C84190E981" box="[664,697,1441,1465]" italics="true" pageId="7" pageNumber="252">bst</emphasis>
|
||
), elongate galea (
|
||
<figureCitation id="AFC99DF1FFDFFFECFA9958C840C5E982" box="[915,1004,1441,1466]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFAF458C8470BE981" box="[1022,1058,1441,1465]" italics="true" pageId="7" pageNumber="252">gal</emphasis>
|
||
) and lacinia (
|
||
<figureCitation id="AFC99DF1FFDFFFECFDC558C84600E982" box="[1231,1321,1441,1466]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFC3158C84674E981" box="[1339,1373,1441,1465]" italics="true" pageId="7" pageNumber="252">lac</emphasis>
|
||
) and moderately long maxillary palp (
|
||
<figureCitation id="AFC99DF1FFDFFFECFB0358AC4174E9E6" box="[521,605,1477,1502]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFB6358AF41B1E9E5" box="[617,664,1478,1501]" italics="true" pageId="7" pageNumber="252">mxp</emphasis>
|
||
) composed of only slightly elongate palpomere I, strongly elongate, pedunculate palpomere II, elongate palpomere III broadest in distal third, and small, elongate, subconical and pointed palpomere IV with distinctly delimited apical part.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFDFFFECF9CD5B584268EA8D" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
|
||
Labium (
|
||
<figureCitation id="AFC99DF1FFDFFFECF83A5B5842ACEA72" box="[304,389,1585,1610]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
) with transverse submentum (
|
||
<figureCitation id="AFC99DF1FFDFFFECFBEE5B584012EA72" box="[740,827,1585,1610]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFA405B5B405EEA71" box="[842,887,1586,1609]" italics="true" pageId="7" pageNumber="252">smn</emphasis>
|
||
) not demarcated posteriorly and fused laterally with postcardinal parts of hypostomae; subtrapezoidal mentum (
|
||
<figureCitation id="AFC99DF1FFDFFFECFA965B3C40CEEA56" box="[924,999,1621,1646]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig.10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFAF15B3F4734EA55" box="[1019,1053,1622,1645]" italics="true" pageId="7" pageNumber="252">mn</emphasis>
|
||
); and short prementum bearing narrowly separated at bases small 3-segmented labial palps (
|
||
<figureCitation id="AFC99DF1FFDFFFECFA365B1040A6EAAA" box="[828,911,1657,1682]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFA915B104098EAA9" box="[923,945,1657,1681]" italics="true" pageId="7" pageNumber="252">lp</emphasis>
|
||
). Hypostomal ridges (
|
||
<figureCitation id="AFC99DF1FFDFFFECFDA65B1047D6EAAA" box="[1196,1279,1657,1682]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFC015B10460DEAA9" box="[1291,1316,1657,1681]" italics="true" pageId="7" pageNumber="252">hr</emphasis>
|
||
) short and evenly arcuate.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFDFFFECF9CD5BA84092EB1A" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
|
||
Gular plate (
|
||
<figureCitation id="AFC99DF1FFDFFFECF8575BA8429CEAE2" box="[349,437,1729,1754]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECF8CC5BAB42CBEAE1" box="[454,482,1730,1753]" italics="true" pageId="7" pageNumber="252">gp</emphasis>
|
||
) large and indistinctly narrowing anterad; gular sutures (
|
||
<figureCitation id="AFC99DF1FFDFFFECFD8D5BA847F6EAE2" box="[1159,1247,1729,1754]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECFDFA5BAB4620EAE1" box="[1264,1289,1730,1753]" italics="true" pageId="7" pageNumber="252">gs</emphasis>
|
||
) superficial; posterior tentorial pits (
|
||
<figureCitation id="AFC99DF1FFDFFFECF8975B8C42D9EAC6" box="[413,496,1765,1790]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="7" pageNumber="252">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFDFFFECF8F65B8F4109EAC5" box="[508,544,1766,1789]" italics="true" pageId="7" pageNumber="252">ptp</emphasis>
|
||
) small but distinct, located at base of submentum, anterior to transverse arcuate impression delimiting ventrally 'neck region' from anterior part of head.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFDFFFECF9CD5A4746AEEB7D" blockId="7.[151,1436,649,2006]" box="[199,1415,1837,1862]" pageId="7" pageNumber="252">
|
||
Antennae (
|
||
<figureCitation id="AFC99DF1FFDFFFECF84B5A4442ABEB7E" box="[321,386,1837,1862]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="7" pageNumber="252">Fig. 1</figureCitation>
|
||
) short in relation to body, with distinctly delimited club composed of antennomeres IX–XI.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFDFFFECF9CD5A3B42E9EBEE" blockId="7.[151,1436,649,2006]" pageId="7" pageNumber="252">
|
||
Pronotum (
|
||
<figureCitation id="AFC99DF1FFDFFFECF84F5A3842ACEB52" box="[325,389,1873,1898]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="7" pageNumber="252">Fig. 1</figureCitation>
|
||
) in dorsal view oval with weakly arcuate anterior and posterior margins and sides rounded in anterior half and nearly straight in posterior third, anterior and posterior corners rounded and indistinct; marginal and sub-lateral carinae absent; base of pronotum with fine and arcuate transverse groove, without pits; sides of pronotum without bristles.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5DF1428DED94" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
Prosternum (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8535DFE4283EC88" box="[345,426,151,176]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
) with short basisternal part (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FBFC5DFE4060EC88" box="[758,841,151,176]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FA525DF14050EC88" box="[856,889,152,176]" italics="true" pageId="8" pageNumber="253">bst</emphasis>
|
||
) indistinctly demarcated from procoxal cavities (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F9955DD543D8ECEC" box="[159,241,188,213]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3F9F55DD4420FECEC" box="[255,294,189,212]" italics="true" pageId="8" pageNumber="253">pcc</emphasis>
|
||
) by fine arcuate carina; median part of sternum with weakly developed prosternal intercoxal process visible as a subtriangular slightly convex area between procoxal cavities; procoxal sockets (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FD965DB647C7ECC0" box="[1180,1262,223,248]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FDF15D884609ECC0" box="[1275,1312,225,248]" italics="true" pageId="8" pageNumber="253">pcs</emphasis>
|
||
) closed by broad postero-lateral lobes of prosternum; hypomera (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FA0F5C6D4073ED24" box="[773,858,260,285]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FA635C6D40AAED24" box="[873,899,260,284]" italics="true" pageId="8" pageNumber="253">hy</emphasis>
|
||
) elongate, divided into broad lateral parts and narrow internal (adcoxal) parts, adcoxal parts of hypomera anteriorly fused with prosternum, so that pronotosternal sutures (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F9FD5C254262ED5C" box="[247,331,332,357]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3F8535C244254ED5C" box="[345,381,333,356]" italics="true" pageId="8" pageNumber="253">nss</emphasis>
|
||
) visible only along sides of basisternal parts of prosternum; hypomeral ridges (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FC065C254649ED5C" box="[1292,1376,332,357]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FC675C2546BBED5C" box="[1389,1426,332,356]" italics="true" pageId="8" pageNumber="253">hyr</emphasis>
|
||
) complete, anteriorly connected to pronotosternal sutures; internal parts of hypomera with long median longitudinal grooves (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F9F55CFD4279ED94" box="[255,336,404,429]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3F8575CFD42BAED94" box="[349,403,404,428]" italics="true" pageId="8" pageNumber="253">mlhy</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5CD140E8EDCD" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">Mesonotum very small, subtrapezoidal, with slightly concave lateral margins of subtriangular mesocutellum barely visible between bases of elytra; mesoscutoscutellar suture absent.</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5C96462AEEF5" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
Mesoventrite (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8605C964292EE20" box="[362,443,511,536]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
) with narrow anterior ridge (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FA0A5C96407AEE20" box="[768,851,511,536]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FA555F684051EE20" box="[863,888,513,536]" italics="true" pageId="8" pageNumber="253">ar</emphasis>
|
||
); mesoventral intercoxal process (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FDF05C964664EE20" box="[1274,1357,511,536]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FC535F6846BAEE20" box="[1369,1427,513,536]" italics="true" pageId="8" pageNumber="253">msvp</emphasis>
|
||
) narrow and moderately expanded ventrally, anteriorly connected with anterior ridge and with subtriangular posterior margin, without posterior arms; asetose lateral impressions (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FAB85F2E472EEE58" box="[946,1031,583,608]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FD1F5F214703EE67" box="[1045,1066,584,607]" italics="true" pageId="8" pageNumber="253">ai</emphasis>
|
||
) present, each with porous field near middle; mesanepisternum with short prepectus (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FBED5F054013EEBC" box="[743,826,620,645]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FA4F5F044040EEBC" box="[837,873,621,644]" italics="true" pageId="8" pageNumber="253">pre</emphasis>
|
||
) and posterior part largely hidden in ventral view; mesepimeron not visible in ventral view; sides of mesothorax without foveae; mesocoxal projections (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FC0C5FE64670EE90" box="[1286,1369,655,680]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FC6E5FF846BAEE90" box="[1380,1427,657,680]" italics="true" pageId="8" pageNumber="253">mcp</emphasis>
|
||
) with mesocoxal sockets (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8BA5FDD412AEEF4" box="[432,515,692,717]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FB055FDC416CEEF4" box="[527,581,693,716]" italics="true" pageId="8" pageNumber="253">mscs</emphasis>
|
||
) located on their mesoventral surface, without posterior lobes.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5FBE401FEF0F" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
Metaventrite (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8655FBE42E2EEC8" box="[367,459,727,752]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3F8EB5FB04120EEC8" box="[481,521,729,752]" italics="true" pageId="8" pageNumber="253">vIII</emphasis>
|
||
) slightly longer than wide, anteriorly fused with mesoventrite, posteriorly moderately shallowly bisinuate and with narrow median metaventral intercoxal process (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FD885F9547FFEF2C" box="[1154,1238,764,789]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FDE95F944630EF2C" box="[1251,1305,765,788]" italics="true" pageId="8" pageNumber="253">mtvp</emphasis>
|
||
) composed of two long spines. Metanepisterna and metepimera narrow.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5E2D4601EF64" blockId="8.[151,1436,151,2013]" box="[199,1320,836,861]" pageId="8" pageNumber="253">
|
||
Metafurca (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8435E2D42B3EF64" box="[329,410,836,861]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
) with very short and broad stalk and divergent lateral furcal arms (
|
||
<figureCitation id="AFC99DF1FFD0FFE3FD8D5E2D47F3EF64" box="[1159,1242,836,861]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3FDEC5E2D4631EF64" box="[1254,1304,836,860]" italics="true" pageId="8" pageNumber="253">mtfa</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5E0E43D3EFF0" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
Elytra (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8115E0E4249EFB8" box="[283,352,871,896]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="8" pageNumber="253">Fig. 2</figureCitation>
|
||
) oval, each with two rudimentary and asetose basal foveae located in shallow basal impression; humeral calli well-marked and developed as longitudinal protuberances; elytral apices unmodified, separately rounded.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5EBD4079EFD5" blockId="8.[151,1436,151,2013]" box="[199,848,980,1005]" pageId="8" pageNumber="253">
|
||
<collectingCountry id="4FE5C1E4FFD0FFE3F9CD5EBD43D6EFD5" box="[199,255,980,1005]" name="India" pageId="8" pageNumber="253">Hind</collectingCountry>
|
||
wings well-developed, about twice as long as elytra.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD5E91422DE86F" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
Legs (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F81B5E9E424DE828" box="[273,356,1015,1040]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="8" pageNumber="253">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="AFC99DF1FFD0FFE3F87E5E914293E828" box="[372,442,1016,1040]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">11–12</figureCitation>
|
||
) moderately long and slender; procoxae subglobose, mesocoxae elongate, metacoxae transverse; all trochanters short; all femora weakly clavate; tibiae short and slightly thickening distally; tarsi short and stout.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD590D47BCE845" blockId="8.[151,1436,151,2013]" box="[199,1173,1124,1149]" pageId="8" pageNumber="253">
|
||
Abdominal sternites (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F8B2590D4123E844" box="[440,522,1124,1149]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
) unmodified, suture between VII and VIII barely marked.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD59EE4630E8FC" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
Aedeagus (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F84F59EE42E4E898" box="[325,461,1159,1184]" captionStart="FIGURES 22 – 30" captionStartId="18.[151,264,1128,1150]" captionTargetBox="[153,1431,195,1098]" captionTargetId="figure@18.[151,1436,193,1107]" captionTargetPageId="18" captionText="FIGURES 22 – 30. Aedeagi in ventral (22, 24, 27, 28, 30), dorsal (25) and lateral (23, 26, 29) views. Microraphes pygmaeus Franz, holotype (22); Heteroscydmus yapacariensis Franz, holotype (24 – 26) and paratype (27); Mimoscydmus brasiliensis Franz, holotype (28 – 29); Amimoscydmus baruerii (Franz), holotype (30)." httpUri="https://zenodo.org/record/216801/files/figure.png" pageId="8" pageNumber="253">Figs. 24–27</figureCitation>
|
||
) with symmetrical, lightly sclerotized and thin-walled median lobe, with complex and asymmetrical set of internal sclerites, parameres present, not fused with median lobe, with apical setae.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE3F9CD59A64718E8D0" blockId="8.[151,1436,151,2013]" box="[199,1073,1231,1256]" pageId="8" pageNumber="253">
|
||
Spermatheca (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F86D59A64293E8D0" box="[359,442,1231,1256]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="05865D66FFD0FFE3F8CC59B842CEE8D0" box="[454,487,1233,1256]" italics="true" pageId="8" pageNumber="253">spt</emphasis>
|
||
) globular, darkly sclerotized, located in metathorax.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD0FFE3F9CD599D4251E908" pageId="8" pageNumber="253" type="distribution">
|
||
<paragraph id="374D8174FFD0FFE3F9CD599D4251E908" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
<emphasis id="05865D66FFD0FFE3F9CD599D4104E935" bold="true" box="[199,557,1268,1293]" pageId="8" pageNumber="253">Distribution and composition.</emphasis>
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FB3E599D41C9E934" box="[564,736,1268,1292]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FB3E599D41C9E934" box="[564,736,1268,1292]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
is represented by a single species known from the central part of
|
||
<collectingCountry id="4FE5C1E4FFD0FFE3F9BF587E4221E908" box="[181,264,1303,1328]" name="Bolivia" pageId="8" pageNumber="253">Bolivia</collectingCountry>
|
||
(
|
||
<figureCitation id="AFC99DF1FFD0FFE3F81D587E4242E908" box="[279,363,1303,1328]" captionStart="FIGURES 35 – 38" captionStartId="19.[151,264,673,695]" captionTargetBox="[151,1433,193,649]" captionTargetId="figure@19.[151,1436,193,651]" captionTargetPageId="19" captionText="FIGURES 35 – 38. Distribution of revised taxa. Microraphes pygmaeus Franz (35); Heteroscydmus yapacariensis Franz (36); Mimoscydmus brasiliensis Franz (37); Amimoscydmus baruerii (Franz) (38)." httpUri="https://zenodo.org/record/216803/files/figure.png" pageId="8" pageNumber="253">Fig. 36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD0FFE2F9CD585540A9ED78" lastPageId="9" lastPageNumber="254" pageId="8" pageNumber="253" type="discussion">
|
||
<paragraph id="374D8174FFD0FFE3F9CD5855408BEB10" blockId="8.[151,1436,151,2013]" pageId="8" pageNumber="253">
|
||
<emphasis id="05865D66FFD0FFE3F9CD58554212E96D" bold="true" box="[199,315,1340,1365]" pageId="8" pageNumber="253">Remarks.</emphasis>
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F840585542DFE96C" box="[330,502,1340,1364]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3F840585542DFE96C" box="[330,502,1340,1364]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
in the first paragraph of Franz's description (Franz 1980) was compared to
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F99D58094215E940" box="[151,316,1376,1400]" class="Insecta" family="Staphylinidae" genus="Pseudoeudesis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3F99D58094215E940" box="[151,316,1376,1400]" italics="true" pageId="8" pageNumber="253">Pseudoeudesis</emphasis>
|
||
</taxonomicName>
|
||
, a genus of
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F8EF58364153E940" box="[485,634,1375,1400]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
|
||
, but in the further parts of the text it was correctly assigned to
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F99D58ED4214E9A5" box="[151,317,1412,1437]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
|
||
and stated to be closely similar to
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FBDA58ED405BE9A4" box="[720,882,1412,1436]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FBDA58ED405BE9A4" box="[720,882,1412,1436]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
. One of characters that Franz discussed to place
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F99D58C1426AE9F8" box="[151,323,1448,1472]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3F99D58C1426AE9F8" box="[151,323,1448,1472]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F86C58CE4125E9F8" box="[358,524,1447,1472]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
|
||
and not
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FB6458CE4029E9F8" box="[622,768,1447,1472]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
|
||
is the apical margin of the scape, which according to Franz is not emarginate. In fact, the distal margin of the scape in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FA2758A540F0E9DC" box="[813,985,1484,1508]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FA2758A540F0E9DC" box="[813,985,1484,1508]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
is relatively deeply emarginate dorsally (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F995588643CCEA30" box="[159,229,1519,1544]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 9</figureCitation>
|
||
); this character as a feature to distinguish
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FBC658864077EA30" box="[716,862,1519,1544]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
|
||
(often with deeply emarginated or notched scape) from
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F9D05B7D42A9EA15" box="[218,384,1556,1581]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
|
||
(usually with shallowly emarginate or straight apical margin) was recently discussed by Jałoszyński (2012). Interestingly, also in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FB615B514024EA68" box="[619,781,1592,1616]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FB615B514024EA68" box="[619,781,1592,1616]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
the apical margin of the scape is deeply emarginate. The shape of maxillary palpomere IV, ventral structures of the prothorax and the aedeagus suggest that the placement of
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F99D5BE9426AEAA0" box="[151,323,1664,1688]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3F99D5BE9426AEAA0" box="[151,323,1664,1688]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F8615B164138EAA0" box="[363,529,1663,1688]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
|
||
was correct. Similarly to
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FA375BE940F6EAA0" box="[829,991,1664,1688]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FA375BE940F6EAA0" box="[829,991,1664,1688]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FD105BE94783EAA0" box="[1050,1194,1664,1688]" class="Insecta" family="Staphylinidae" genus="Microraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FD105BE94783EAA0" box="[1050,1194,1664,1688]" italics="true" pageId="8" pageNumber="253">Microraphes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FDB05BE9464FEAA0" box="[1210,1382,1664,1688]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FDB05BE9464FEAA0" box="[1210,1382,1664,1688]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
also has the internal parts of prothoracic hypomera anteriorly fused with the prosternum, so the pronotosternal sutures are visible only along sides of the basisternal part of prosternum, and in ventral view it seems like the notosternal sutures were continuous with the hypomeral ridges that separate the internal part of each hypomeron from the external part, which is merely a lateral deflexed part of the pronotum.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD0FFE2F9CD5A5D40A9ED78" blockId="8.[151,1436,151,2013]" lastBlockId="9.[151,1436,151,320]" lastPageId="9" lastPageNumber="254" pageId="8" pageNumber="253">
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F9CD5A5D425AEB74" box="[199,371,1844,1868]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3F9CD5A5D425AEB74" box="[199,371,1844,1868]" italics="true" pageId="8" pageNumber="253">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FB645A5D4039EB74" box="[622,784,1844,1868]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FB645A5D4039EB74" box="[622,784,1844,1868]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
in the following characters: the posterior tentorial pits located distinctly anterior to transverse impression that ventrally demarcates the 'neck region' from the anterior part of head (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F9F55A15427AEBAC" box="[255,339,1916,1941]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 10</figureCitation>
|
||
) (pits in the impression in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FB9B5A15401AEBAC" box="[657,819,1916,1940]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FB9B5A15401AEBAC" box="[657,819,1916,1940]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
); base of pronotum with short and narrow transverse groove (two pairs of pits in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3F8D95AC9415CEB80" box="[467,629,1952,1976]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3F8D95AC9415CEB80" box="[467,629,1952,1976]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
); internal parts of prothoracic hypomera with long median longitudinal hypomeral grooves (
|
||
<figureCitation id="AFC99DF1FFD0FFE3F88F5AAD42FEEBE4" box="[389,471,1988,2013]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="8" pageNumber="253">Fig. 11</figureCitation>
|
||
) (grooves absent in
|
||
<taxonomicName id="F0F2FAF7FFD0FFE3FBC15AAD4044EBE4" box="[715,877,1988,2012]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD0FFE3FBC15AAD4044EBE4" box="[715,877,1988,2012]" italics="true" pageId="8" pageNumber="253">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
); lack of bristles on sides of pronotum (bristles present in
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2F82A5DF142EBEC88" box="[288,450,152,176]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2F82A5DF142EBEC88" box="[288,450,152,176]" italics="true" pageId="9" pageNumber="254">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
); the mesoventrite without setose impressions (present and fused in middle in
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2F99D5DD54210ECEC" box="[151,313,188,212]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2F99D5DD54210ECEC" box="[151,313,188,212]" italics="true" pageId="9" pageNumber="254">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
); the mesoventral intercoxal process connected anteriorly with the anterior ridge of mesoventrite (anterior margin of process distant from the anterior ridge in
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FA535D8940D2ECC0" box="[857,1019,224,248]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FA535D8940D2ECC0" box="[857,1019,224,248]" italics="true" pageId="9" pageNumber="254">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
); and each elytron with two asetose rudiments of basal foveae (one large and setose fovea in
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FA055C6D4098ED24" box="[783,945,260,284]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FA055C6D4098ED24" box="[783,945,260,284]" italics="true" pageId="9" pageNumber="254">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
). Major differences between
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FDF05C6D46B5ED24" box="[1274,1436,260,284]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FDF05C6D46B5ED24" box="[1274,1436,260,284]" italics="true" pageId="9" pageNumber="254">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
and all genera treated in the present paper are compiled in Table 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD1FFE2F99D5CFA41F6EE2E" pageId="9" pageNumber="254" type="description">
|
||
<paragraph id="374D8174FFD1FFE2F99D5CFA42CDEDE8" blockId="9.[151,570,403,464]" pageId="9" pageNumber="254">
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2F99D5CFA4113ED95" authority="Franz" authorityName="Franz" box="[151,570,403,429]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="species" species="yapacariensis">
|
||
<emphasis id="05865D66FFD1FFE2F99D5CFA4113ED95" bold="true" box="[151,570,403,429]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F99D5CFA42C5ED94" bold="true" box="[151,492,403,428]" italics="true" pageId="9" pageNumber="254">Heteroscydmus yapacariensis</emphasis>
|
||
Franz
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AFC99DF1FFD1FFE2F9955CDE43C7EDE8" box="[159,238,439,464]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="9" pageNumber="254">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="AFC99DF1FFD1FFE2F9F05CD1421CEDE8" box="[250,309,440,464]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="9" pageNumber="254">9–12</figureCitation>
|
||
,
|
||
<figureCitation id="AFC99DF1FFD1FFE2F84A5CD142A0EDE8" box="[320,393,440,464]" captionStart="FIGURES 22 – 30" captionStartId="18.[151,264,1128,1150]" captionTargetBox="[153,1431,195,1098]" captionTargetId="figure@18.[151,1436,193,1107]" captionTargetPageId="18" captionText="FIGURES 22 – 30. Aedeagi in ventral (22, 24, 27, 28, 30), dorsal (25) and lateral (23, 26, 29) views. Microraphes pygmaeus Franz, holotype (22); Heteroscydmus yapacariensis Franz, holotype (24 – 26) and paratype (27); Mimoscydmus brasiliensis Franz, holotype (28 – 29); Amimoscydmus baruerii (Franz), holotype (30)." httpUri="https://zenodo.org/record/216801/files/figure.png" pageId="9" pageNumber="254">24–27</figureCitation>
|
||
,
|
||
<figureCitation id="AFC99DF1FFD1FFE2F89E5CD1429AEDE8" box="[404,435,440,464]" captionStart="FIGURES 31 – 34" captionStartId="18.[151,264,1881,1903]" captionTargetBox="[158,1430,1331,1850]" captionTargetId="figure@18.[151,1436,1292,1860]" captionTargetPageId="18" captionText="FIGURES 31 – 34. Original set of labels of holotypes (31 – 33) and paratype (34). Microraphes pygmaeus Franz (31); Heteroscydmus yapacariensis Franz (32); Mimoscydmus brasiliensis Franz (33); Amimoscydmus baruerii (Franz) (34)." httpUri="https://zenodo.org/record/216802/files/figure.png" pageId="9" pageNumber="254">32</figureCitation>
|
||
,
|
||
<figureCitation id="AFC99DF1FFD1FFE2F8B45CDE42F2EDE8" box="[446,475,439,464]" captionStart="FIGURES 35 – 38" captionStartId="19.[151,264,673,695]" captionTargetBox="[151,1433,193,649]" captionTargetId="figure@19.[151,1436,193,651]" captionTargetPageId="19" captionText="FIGURES 35 – 38. Distribution of revised taxa. Microraphes pygmaeus Franz (35); Heteroscydmus yapacariensis Franz (36); Mimoscydmus brasiliensis Franz (37); Amimoscydmus baruerii (Franz) (38)." httpUri="https://zenodo.org/record/216803/files/figure.png" pageId="9" pageNumber="254">36</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F99D5F6941F6EE2E" blockId="9.[151,735,512,534]" box="[151,735,512,534]" pageId="9" pageNumber="254">
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2F99D5F694150EE2E" authority="Franz, 1980: 208" authorityName="Franz" authorityPageNumber="208" authorityYear="1980" box="[151,633,512,534]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="species" species="yapacariensis">
|
||
<emphasis id="05865D66FFD1FFE2F99D5F6942EDEE2E" box="[151,452,512,534]" italics="true" pageId="9" pageNumber="254">Heteroscydmus yapacariensis</emphasis>
|
||
Franz, 1980: 208
|
||
</taxonomicName>
|
||
, Fig. 195.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD1FFE2F99D5F2B405BEED3" pageId="9" pageNumber="254" type="materials_examined">
|
||
<paragraph id="374D8174FFD1FFE2F99D5F2B405BEED3" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F99D5F2B4274EE63" bold="true" box="[151,349,578,603]" pageId="9" pageNumber="254">Material studied</emphasis>
|
||
.
|
||
<typeStatus id="E8493FD6FFD1FFE2F8615F2B42F1EE63" box="[363,472,578,603]" pageId="9" pageNumber="254" type="holotype">Holotype</typeStatus>
|
||
: ♂: three labels (
|
||
<figureCitation id="AFC99DF1FFD1FFE2FB9C5F2B41C3EE63" box="[662,746,578,603]" captionStart="FIGURES 31 – 34" captionStartId="18.[151,264,1881,1903]" captionTargetBox="[158,1430,1331,1850]" captionTargetId="figure@18.[151,1436,1292,1860]" captionTargetPageId="18" captionText="FIGURES 31 – 34. Original set of labels of holotypes (31 – 33) and paratype (34). Microraphes pygmaeus Franz (31); Heteroscydmus yapacariensis Franz (32); Mimoscydmus brasiliensis Franz (33); Amimoscydmus baruerii (Franz) (34)." httpUri="https://zenodo.org/record/216802/files/figure.png" pageId="9" pageNumber="254">Fig. 32</figureCitation>
|
||
): "Umg. Sta. Cruz /
|
||
<collectingCountry id="4FE5C1E4FFD1FFE2FAC75F2B4718EE63" box="[973,1073,578,603]" name="Bolivia" pageId="9" pageNumber="254">Bolivien</collectingCountry>
|
||
, lg. Franz" with "SA142" on the reverse side [white, printed; reverse handwritten], "
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FBE35F0F40BCEE46" box="[745,917,614,638]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FBE35F0F40BCEE46" box="[745,917,614,638]" italics="true" pageId="9" pageNumber="254">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FABA5F0E4765EE46" box="[944,1100,615,638]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="species" species="yapacariensis">
|
||
<emphasis id="05865D66FFD1FFE2FABA5F0E4765EE46" box="[944,1100,615,638]" italics="true" pageId="9" pageNumber="254">yapacariensis</emphasis>
|
||
</taxonomicName>
|
||
/ m. / det. H. Franz" [white, handwritten and printed], "
|
||
<typeStatus id="E8493FD6FFD1FFE2F8C35FE2413BEE9B" box="[457,530,651,675]" pageId="9" pageNumber="254">Typus</typeStatus>
|
||
" [red, handwritten] (NHMW).
|
||
<typeStatus id="E8493FD6FFD1FFE2FA725FE240C0EE9B" box="[888,1001,651,675]" pageId="9" pageNumber="254" type="paratype">Paratypes</typeStatus>
|
||
: 1 ♀,
|
||
<specimenCount id="21F44AFDFFD1FFE2FD385FE24748EE9A" box="[1074,1121,651,675]" pageId="9" pageNumber="254" type="generic">1 ♂</specimenCount>
|
||
: same data as for
|
||
<typeStatus id="E8493FD6FFD1FFE2FC395FE346B1EE9B" box="[1331,1432,650,675]" pageId="9" pageNumber="254" type="holotype">holotype</typeStatus>
|
||
, except for yellow "
|
||
<typeStatus id="E8493FD6FFD1FFE2F8795FC642CEEEFF" box="[371,487,687,711]" pageId="9" pageNumber="254" type="paratype">Paratypus</typeStatus>
|
||
" label with the female and a yellowish identification label with the male, without annotation stating the
|
||
<typeStatus id="E8493FD6FFD1FFE2F8855FBA42D9EED3" box="[399,496,723,747]" pageId="9" pageNumber="254" type="paratype">paratype</typeStatus>
|
||
status of this specimen (NHMW).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD1FFE2F9CD5F9F424BEA57" pageId="9" pageNumber="254" type="diagnosis">
|
||
<paragraph id="374D8174FFD1FFE2F9CD5F9F42FFEF0B" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F9CD5F9F4268EF37" bold="true" box="[199,321,758,783]" pageId="9" pageNumber="254">Diagnosis.</emphasis>
|
||
This is the only known species of
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FBC65F9F4051EF36" box="[716,888,758,782]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FBC65F9F4051EF36" box="[716,888,758,782]" italics="true" pageId="9" pageNumber="254">Heteroscydmus</emphasis>
|
||
</taxonomicName>
|
||
and can be identified on the basis of the generic characters and the aedeagus.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD5E5747A5EF43" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F9CD5E57425DEF6F" bold="true" box="[199,372,830,855]" pageId="9" pageNumber="254">Redescription.</emphasis>
|
||
Body of male (
|
||
<figureCitation id="AFC99DF1FFD1FFE2FB415E5741B1EF6F" box="[587,664,830,855]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="9" pageNumber="254">Fig. 2</figureCitation>
|
||
) strongly convex, elongate and slender, with moderately long appendages, BL
|
||
<quantity id="F00A2C91FFD1FFE2F8585E0A4290EF42" box="[338,441,867,891]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="254" unit="mm" value="0.7">0.70 mm</quantity>
|
||
; cuticle glossy, body uniformly light brown, vestiture yellowish.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD5EEF408FE86A" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
Head subtrapezoidal, broadest at eyes, HL
|
||
<quantity id="F00A2C91FFD1FFE2FBBC5EEE4036EFA6" box="[694,799,903,927]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="254" unit="mm" value="0.13">0.13 mm</quantity>
|
||
, HW
|
||
<quantity id="F00A2C91FFD1FFE2FA6F5EEF40E7EFA6" box="[869,974,902,927]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="254" unit="mm" value="0.15">0.15 mm</quantity>
|
||
; tempora about as long as eyes, slightly convergent caudad; vertex and frons confluent, strongly convex and together about as long as broad; supraantennal tubercles indistinct. Punctures on head dorsum fine and sparse, inconspicuous; setae short, sparse and suberect. Antennae slender but strikingly short in relation to body (
|
||
<figureCitation id="AFC99DF1FFD1FFE2FA495E9B40A5E833" box="[835,908,1010,1035]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1098,1120]" captionTargetBox="[151,1426,209,1071]" captionTargetId="figure@4.[151,1436,195,1077]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Dorsal habitus of male holotype (1 – 3) and male paratype (4). Microraphes pygmaeus Franz (1); Heteroscydmus yapacariensis Franz (2); Mimoscydmus brasiliensis Franz (3); Amimoscydmus baruerii (Franz) (4)." httpUri="https://zenodo.org/record/216796/files/figure.png" pageId="9" pageNumber="254">Fig. 2</figureCitation>
|
||
), with distinctly delimited club composed of antennomeres IX–XI, AnL
|
||
<quantity id="F00A2C91FFD1FFE2F8C4597E411EE816" box="[462,567,1047,1071]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="254" unit="mm" value="0.23">0.23 mm</quantity>
|
||
; antennomere I–II elongate, III–VIII each slightly transverse; IX–X strongly transverse; XI slightly broader than long, with subconical, blunt apex.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD593641CFE93F" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
Pronotum in dorsal view oval, broadest near anterior third, PL
|
||
<quantity id="F00A2C91FFD1FFE2FA8E593640C0E84E" box="[900,1001,1119,1143]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="9" pageNumber="254" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, PW
|
||
<quantity id="F00A2C91FFD1FFE2FD2C593747A5E84E" box="[1062,1164,1118,1143]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="254" unit="mm" value="0.15">0.15 mm</quantity>
|
||
; anterior margin weakly arcuate; lateral margins strongly rounded in anterior half and nearly straight in posterior third, strongly convergent toward obtuse and blunt hind pronotal angles; posterior margin weakly arcuate; base of pronotum with short an narrow but distinct transverse ante-basal arcuate groove convex anteriorly. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD587B4615E9AF" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
Elytra oval, distinctly more convex than pronotum, broadest near middle, EL
|
||
<quantity id="F00A2C91FFD1FFE2FD26587A47BBE912" box="[1068,1170,1299,1323]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="254" unit="mm" value="0.4">0.40 mm</quantity>
|
||
, EW
|
||
<quantity id="F00A2C91FFD1FFE2FDDA587A461FE912" box="[1232,1334,1299,1323]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="254" unit="mm" value="0.28">0.28 mm</quantity>
|
||
, EI 1.45; humeral calli distinct; basal impressions shallow and short, basal foveae in a dry mount indiscernible; elytral apices separately rounded. Punctures on elytral disc slightly more distinct than those on pronotum but still fine and inconspicuous; setae short, sparse and suberect.
|
||
<collectingCountry id="4FE5C1E4FFD1FFE2FBB8581741C3E9AF" box="[690,746,1406,1431]" name="India" pageId="9" pageNumber="254">Hind</collectingCountry>
|
||
wings well-developed, about twice as long as elytra.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD58CA4067E983" blockId="9.[151,1437,578,1971]" box="[199,846,1442,1467]" pageId="9" pageNumber="254">Legs moderately long and slender, without modifications.</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD58AF40CCEA3A" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
Aedeagus (
|
||
<figureCitation id="AFC99DF1FFD1FFE2F84058AF42FFE9E7" box="[330,470,1478,1503]" captionStart="FIGURES 22 – 30" captionStartId="18.[151,264,1128,1150]" captionTargetBox="[153,1431,195,1098]" captionTargetId="figure@18.[151,1436,193,1107]" captionTargetPageId="18" captionText="FIGURES 22 – 30. Aedeagi in ventral (22, 24, 27, 28, 30), dorsal (25) and lateral (23, 26, 29) views. Microraphes pygmaeus Franz, holotype (22); Heteroscydmus yapacariensis Franz, holotype (24 – 26) and paratype (27); Mimoscydmus brasiliensis Franz, holotype (28 – 29); Amimoscydmus baruerii (Franz), holotype (30)." httpUri="https://zenodo.org/record/216801/files/figure.png" pageId="9" pageNumber="254">Figs. 24–27</figureCitation>
|
||
) elongate, AeL
|
||
<quantity id="F00A2C91FFD1FFE2FB9658AF402EE9E6" box="[668,775,1478,1503]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="254" unit="mm" value="0.05">0.05 mm</quantity>
|
||
, median lobe oval with truncate apex, internal sclerites asymmetrical and complex; parameres slender, each with single apical seta.
|
||
</paragraph>
|
||
<paragraph id="374D8174FFD1FFE2F9CD5B67424BEA57" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F9CD5B674237EA1F" bold="true" box="[199,286,1550,1575]" pageId="9" pageNumber="254">Female</emphasis>
|
||
. Similar to male but differing in slightly smaller eyes, and in consequence tempora longer than eyes (
|
||
<figureCitation id="AFC99DF1FFD1FFE2F9955B5B43CDEA73" box="[159,228,1586,1611]" captionStart="FIGURES 9 – 12" captionStartId="10.[151,264,1293,1315]" captionTargetBox="[156,1416,199,1257]" captionTargetId="figure@10.[151,1436,193,1271]" captionTargetPageId="10" captionText="FIGURES 9 – 12. Morphology of Heteroscydmus yapacariensis Franz, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="9" pageNumber="254">Fig. 9</figureCitation>
|
||
). BL
|
||
<quantity id="F00A2C91FFD1FFE2F82C5B5A42A7EA72" box="[294,398,1587,1611]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="9" pageNumber="254" unit="mm" value="0.73">0.73 mm</quantity>
|
||
; HL
|
||
<quantity id="F00A2C91FFD1FFE2F8CD5B5A4104EA72" box="[455,557,1587,1611]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="254" unit="mm" value="0.13">0.13 mm</quantity>
|
||
, HW
|
||
<quantity id="F00A2C91FFD1FFE2FB7A5B5B41FEEA72" box="[624,727,1586,1611]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="254" unit="mm" value="0.15">0.15 mm</quantity>
|
||
, AnL
|
||
<quantity id="F00A2C91FFD1FFE2FA145B5A40AFEA72" box="[798,902,1587,1611]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="254" unit="mm" value="0.23">0.23 mm</quantity>
|
||
; PL
|
||
<quantity id="F00A2C91FFD1FFE2FAB15B5A470BEA72" box="[955,1058,1587,1611]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="9" pageNumber="254" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, PW
|
||
<quantity id="F00A2C91FFD1FFE2FD555B5B47EEEA72" box="[1119,1223,1586,1611]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="254" unit="mm" value="0.15">0.15 mm</quantity>
|
||
; EL
|
||
<quantity id="F00A2C91FFD1FFE2FDF75B5A464DEA72" box="[1277,1380,1587,1611]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="254" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, EW
|
||
<quantity id="F00A2C91FFD1FFE2F99D5B3E43D4EA56" box="[151,253,1623,1647]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="254" unit="mm" value="0.3">0.30 mm</quantity>
|
||
, EI 1.42.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD1FFE2F9CD5B1340B1EAAA" box="[199,920,1658,1683]" pageId="9" pageNumber="254" type="distribution">
|
||
<paragraph id="374D8174FFD1FFE2F9CD5B1340B1EAAA" blockId="9.[151,1437,578,1971]" box="[199,920,1658,1683]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F9CD5B134270EAAB" bold="true" box="[199,345,1658,1683]" pageId="9" pageNumber="254">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="AFC99DF1FFD1FFE2F8625B134294EAAB" box="[360,445,1658,1683]" captionStart="FIGURES 35 – 38" captionStartId="19.[151,264,673,695]" captionTargetBox="[151,1433,193,649]" captionTargetId="figure@19.[151,1436,193,651]" captionTargetPageId="19" captionText="FIGURES 35 – 38. Distribution of revised taxa. Microraphes pygmaeus Franz (35); Heteroscydmus yapacariensis Franz (36); Mimoscydmus brasiliensis Franz (37); Amimoscydmus baruerii (Franz) (38)." httpUri="https://zenodo.org/record/216803/files/figure.png" pageId="9" pageNumber="254">Fig. 36</figureCitation>
|
||
). Central
|
||
<collectingCountry id="4FE5C1E4FFD1FFE2FB235B134157EAAB" box="[553,638,1658,1683]" name="Bolivia" pageId="9" pageNumber="254">Bolivia</collectingCountry>
|
||
, Santa Cruz Department.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7FE8D2FFFFD1FFE1F9CD5BF74233EAFB" lastPageId="10" lastPageNumber="255" pageId="9" pageNumber="254" type="discussion">
|
||
<paragraph id="374D8174FFD1FFE2F9CD5BF7425DEB8B" blockId="9.[151,1437,578,1971]" pageId="9" pageNumber="254">
|
||
<emphasis id="05865D66FFD1FFE2F9CD5BF74212EA8F" bold="true" box="[199,315,1694,1719]" pageId="9" pageNumber="254">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="E8493FD6FFD1FFE2F8735BF64280EA8F" box="[377,425,1695,1719]" pageId="9" pageNumber="254">type</typeStatus>
|
||
series reported by Franz (1980) consisted of a
|
||
<typeStatus id="E8493FD6FFD1FFE2FAC75BF74706EA8F" box="[973,1071,1694,1719]" pageId="9" pageNumber="254" type="holotype">holotype</typeStatus>
|
||
male and three
|
||
<typeStatus id="E8493FD6FFD1FFE2FDE65BF64671EA8F" box="[1260,1368,1695,1719]" pageId="9" pageNumber="254" type="paratype">paratypes</typeStatus>
|
||
of an unspecified sex. The
|
||
<typeStatus id="E8493FD6FFD1FFE2F88D5BAA429EEAE3" box="[391,439,1731,1755]" pageId="9" pageNumber="254">type</typeStatus>
|
||
series preserved at NHMW consists of a male
|
||
<typeStatus id="E8493FD6FFD1FFE2FAC65BAB4718EAE3" box="[972,1073,1730,1755]" pageId="9" pageNumber="254" type="holotype">holotype</typeStatus>
|
||
, one female and two males, one of them incomplete and entirely embedded in euparal. The aedeagus of the
|
||
<typeStatus id="E8493FD6FFD1FFE2FAD45B8F4769EAC7" box="[990,1088,1766,1791]" pageId="9" pageNumber="254" type="holotype">holotype</typeStatus>
|
||
male is in an erected condition (
|
||
<figureCitation id="AFC99DF1FFD1FFE2F9955A63420EEB1B" box="[159,295,1802,1827]" captionStart="FIGURES 22 – 30" captionStartId="18.[151,264,1128,1150]" captionTargetBox="[153,1431,195,1098]" captionTargetId="figure@18.[151,1436,193,1107]" captionTargetPageId="18" captionText="FIGURES 22 – 30. Aedeagi in ventral (22, 24, 27, 28, 30), dorsal (25) and lateral (23, 26, 29) views. Microraphes pygmaeus Franz, holotype (22); Heteroscydmus yapacariensis Franz, holotype (24 – 26) and paratype (27); Mimoscydmus brasiliensis Franz, holotype (28 – 29); Amimoscydmus baruerii (Franz), holotype (30)." httpUri="https://zenodo.org/record/216801/files/figure.png" pageId="9" pageNumber="254">Figs. 24–26</figureCitation>
|
||
) and was certainly not illustrated by Franz; his figure (1980, Fig. 195) was apparently based on the non-erected aedeagus of one of the
|
||
<typeStatus id="E8493FD6FFD1FFE2FB2F5A4641AFEB7F" box="[549,646,1839,1863]" pageId="9" pageNumber="254" type="paratype">paratype</typeStatus>
|
||
males (
|
||
<figureCitation id="AFC99DF1FFD1FFE2FBD75A47401BEB7F" box="[733,818,1838,1863]" captionStart="FIGURES 22 – 30" captionStartId="18.[151,264,1128,1150]" captionTargetBox="[153,1431,195,1098]" captionTargetId="figure@18.[151,1436,193,1107]" captionTargetPageId="18" captionText="FIGURES 22 – 30. Aedeagi in ventral (22, 24, 27, 28, 30), dorsal (25) and lateral (23, 26, 29) views. Microraphes pygmaeus Franz, holotype (22); Heteroscydmus yapacariensis Franz, holotype (24 – 26) and paratype (27); Mimoscydmus brasiliensis Franz, holotype (28 – 29); Amimoscydmus baruerii (Franz), holotype (30)." httpUri="https://zenodo.org/record/216801/files/figure.png" pageId="9" pageNumber="254">Fig. 27</figureCitation>
|
||
). The other male during the present study was found to belong to a different species and even genus than the
|
||
<typeStatus id="E8493FD6FFD1FFE2FA1A5A3B405FEB53" box="[784,886,1874,1899]" pageId="9" pageNumber="254" type="holotype">holotype</typeStatus>
|
||
. It is a
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FAD95A3A475CEB53" box="[979,1141,1875,1899]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FAD95A3A475CEB53" box="[979,1141,1875,1899]" italics="true" pageId="9" pageNumber="254">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="F0F2FAF7FFD1FFE2FDA85A3A466BEB53" box="[1186,1346,1875,1899]" class="Insecta" family="Staphylinidae" genus="Mimoscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="254" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="05865D66FFD1FFE2FDA85A3A466BEB53" box="[1186,1346,1875,1899]" italics="true" pageId="9" pageNumber="254">Mimoscydmus</emphasis>
|
||
</taxonomicName>
|
||
, but its aedeagus is distorted to a degree that makes a species identification impossible, so ventral details of this specimen were not examined.
|
||
</paragraph>
|
||
<caption id="638DD1FCFFD2FFE1F99D586442A3EA20" httpUri="https://zenodo.org/record/216798/files/figure.png" pageId="10" pageNumber="255" targetBox="[156,1416,199,1257]" targetPageId="10">
|
||
<paragraph id="374D8174FFD2FFE1F99D586442A3EA20" blockId="10.[151,1436,1293,1560]" pageId="10" pageNumber="255">
|
||
<emphasis id="05865D66FFD2FFE1F99D58644260E91B" bold="true" box="[151,329,1293,1315]" pageId="10" pageNumber="255">FIGURES 9–12.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="F0F2FAF7FFD2FFE1F8FD58644044E91B" authority="Franz" authorityName="Franz" box="[503,877,1293,1315]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="255" phylum="Arthropoda" rank="species" species="yapacariensis">
|
||
<emphasis id="05865D66FFD2FFE1F8FD5864400EE91B" box="[503,807,1293,1315]" italics="true" pageId="10" pageNumber="255">Heteroscydmus yapacariensis</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
, paratype female. Head in dorsal (9) and ventral (10) views; prothorax in ventral view (11); pterothorax and abdomen in ventral view (12). Abbreviations: ai, asetose impression; ar, anterior ridge; bst, basisternal part of prosternum; cd, cardo; cl, clypeus; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; fo, foramen occipitale; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nr, neck region; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; ppf, palpifer; pre, prepectus; pstp, prosternal intercoxal process; ptp, posterior tentorial pit; smn, submentum; spt, spermatheca; tm, temple; vIII, metaventrite; vt, vertex.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="374D8174FFD2FFE1F9CD5B294233EAFB" blockId="10.[151,1436,1599,1732]" pageId="10" pageNumber="255">
|
||
Franz named this species "
|
||
<taxonomicName id="F0F2FAF7FFD2FFE1F8FC5B2941BBEA6F" box="[502,658,1600,1623]" class="Insecta" family="Staphylinidae" genus="Heteroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="255" phylum="Arthropoda" rank="species" species="yapacariensis">
|
||
<emphasis id="05865D66FFD2FFE1F8FC5B2941BBEA6F" box="[502,658,1600,1623]" italics="true" pageId="10" pageNumber="255">yapacariensis</emphasis>
|
||
</taxonomicName>
|
||
" after the Rio Yapacani river in
|
||
<collectingCountry id="4FE5C1E4FFD2FFE1FD0E5B564770EA60" box="[1028,1113,1599,1624]" name="Bolivia" pageId="10" pageNumber="255">Bolivia</collectingCountry>
|
||
, misspelled as "Yapacari" in his paper (Franz 1980). In the original description Franz (1980) gives further data related to the collecting site of the
|
||
<typeStatus id="E8493FD6FFD2FFE1F9CA5BE143D9EA98" box="[192,240,1672,1696]" pageId="10" pageNumber="255">type</typeStatus>
|
||
series: a gallery forest along Rio Yapacani (as Yapacari), western Montero, forest litter and rotten wood of fallen trees.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |