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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1148.95629" ID-Pensoft-Pub="1313-2970-1148-79" ID-Pensoft-UUID="74E19B4BEA41574FA77DA36192DC3F70" ID-ZooBank="9A646626019345F7ACF0B809374C74F3" ModsDocID="1313-2970-1148-79" checkinTime="1676637843037" checkinUser="pensoft" docAuthor="Salini, Santhamma, Gracy, R. G., Akoijam, Romila, Rabbani, Mehaboob K., David, K. Jacob &amp; Roca-Cusachs, Marcos" docDate="2023" docId="73F96188B34E5CBF9E5CB93F54979C9E" docLanguage="en" docName="ZooKeys 1148: 79-117" docOrigin="ZooKeys 1148" docPubDate="2023-02-16" docSource="http://dx.doi.org/10.3897/zookeys.1148.95629" docTitle="Mycterizon bellus Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs 2023, stat. rev." docType="treatment" docVersion="1" id="74E19B4BEA41574FA77DA36192DC3F70" lastPageNumber="79" masterDocId="74E19B4BEA41574FA77DA36192DC3F70" masterDocTitle="Revision of Acesines Stal and Dunnius Distant, resurrection of Mycterizon Breddin (Hemiptera, Heteroptera, Pentatomidae, Pentatominae), and description of a new species from India" masterLastPageNumber="117" masterPageNumber="79" pageNumber="79" updateTime="1676637843037" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Acesines Stal and Dunnius Distant, resurrection of Mycterizon Breddin (Hemiptera, Heteroptera, Pentatomidae, Pentatominae), and description of a new species from India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salini, Santhamma</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1234-8330</mods:nameIdentifier>
<mods:affiliation>ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, India</mods:affiliation>
<mods:nameIdentifier type="email">shalini.nbaii@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gracy, R. G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6764-5167</mods:nameIdentifier>
<mods:affiliation>ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Akoijam, Romila</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5592-2841</mods:nameIdentifier>
<mods:affiliation>ICAR-Regional Station, ICAR Complex for NEH Region, Manipur Centre, Lamphelpat, Imphal, 795004, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rabbani, Mehaboob K.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7306-082X</mods:nameIdentifier>
<mods:affiliation>ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>David, K. Jacob</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5092-141X</mods:nameIdentifier>
<mods:affiliation>ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Roca-Cusachs, Marcos</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9174-6021</mods:nameIdentifier>
<mods:affiliation>IRBio. Institut de Recerca a la Biodiversitat, Universitat de Barcelona, Barcelona, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1148</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>79</mods:start>
<mods:end>117</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1148.95629</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1148.95629</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1148-79</mods:identifier>
<mods:identifier type="ZooBank">9A646626019345F7ACF0B809374C74F3</mods:identifier>
<mods:identifier type="Pensoft-UUID">74E19B4BEA41574FA77DA36192DC3F70</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="79" type="taxonomic account">
<treatment LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E" httpUri="http://treatment.plazi.org/id/73F96188B34E5CBF9E5CB93F54979C9E" lastPageNumber="79" pageId="0" pageNumber="79">
<subSubSection pageId="0" pageNumber="79" type="nomenclature">
<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="73F96188-B34E-5CBF-9E5C-B93F54979C9E" authority="(Distant, 1902)" authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus" status="stat. rev.">Mycterizon bellus (Distant, 1902)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">stat. rev.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">Figs 63-68</figureCitation>
<figureCitation captionStart="Figures 69–75" captionStartId="F14" captionText="Figures 69 - 75. Mycterizon bellus (Distant), stat. rev. 69 genital capsule (ventral) 70 genital capsule (caudal) 71, 72 paramere 73 articulatory apparatus 74 phallus (dorsal) 75 phallus (ventral). Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; cp - capitate processes; ls - lobe-like ingrowths; pa - processes of aedeagus; dcp I - dorsal conjunctival processes (inner one); dcp II - dorsal conjunctival processes (outer one); lcp - lateral conjunctival processes; vcp - ventral conjunctival processes." figureDoi="10.3897/zookeys.1148.95629.figures69-75" httpUri="https://binary.pensoft.net/fig/811002" pageId="0" pageNumber="79">, 69-75</figureCitation>
<figureCitation captionStart="Figures 76–79" captionStartId="F15" captionText="Figures 76 - 79. Mycterizon bellus (Distant), stat. rev. 76 phallus (lateral) 77 terminalia (dorsal) 78 terminalia (ventral) 79 spermatheca. Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; ar - apical receptacle; rs - ring sclerites; lt VIII - laterotergite VIII; lt IX - laterotergite IX; vf VIII - valvifers VIII; vf IX - valvifers IX." figureDoi="10.3897/zookeys.1148.95629.figures76-79" httpUri="https://binary.pensoft.net/fig/811003" pageId="0" pageNumber="79">, 76-79</figureCitation>
<figureCitation captionStart="Figures 80–85" captionStartId="F16" captionText="Figures 80 - 85. Mycterizon bellus (Distant), stat. rev. 80 label data of lectotype 81 lectotype (dorsal habitus) 82 lectotype (ventral habitus) 83 label data of paralectotype 84 paralectotype (dorsal habitus) 85 paralectotype (ventral habitus). Photographed by Mick Webb, NHM, London, UK." figureDoi="10.3897/zookeys.1148.95629.figures80-85" httpUri="https://binary.pensoft.net/fig/811004" pageId="0" pageNumber="79">, 80-85</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="reference_group">
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bella" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bella" status="stat. rev.">Araducta bella</taxonomicName>
Distant, 1900a: 427 (original description). Syntype: ♂, Punduloya, Ceylon, Atkinson Coll. 92-6, 116 (NHM), London, UK, lectotype (present designation); Syntype: ♀, Punduloya, Ceylon, Atkinson Coll. 92-6, 116 (NHM), London, UK, paralectotype (present designation).
</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus" status="stat. rev.">Dunnius bellus</taxonomicName>
Distant, 1902: 233, fig. 146 (redescription, figures of habitat in dorsal view, and mesothorax, metathorax and abdomen in ventral view; distribution);
<bibRefCitation author="Bergroth, E" journalOrPublisher="Memoires de la Societe Entomologique de Belgique" pageId="0" pageNumber="79" pagination="131 - 200" refId="B2" refString="Bergroth, E, 1908. Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum. Memoires de la Societe Entomologique de Belgique 15: 131 - 200" title="Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum." volume="15" year="1908">Bergroth (1908)</bibRefCitation>
: 173 (catalogue);
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.15205" author="Kirkaldy, GW" journalOrPublisher="Journal of the Linnean Society of Zoology" pageId="0" pageNumber="79" refId="B20" refString="Kirkaldy, GW, 1909. Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families (Vol. I). Cimicidae. Felix L. Dames, Berlin, [xl +] 392 pp. https://doi.org/10.5962/bhl.title.15205" title="Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families (Vol. I). Cimicidae. Felix L. Dames, Berlin, [xl +] 392 pp." url="https://doi.org/10.5962/bhl.title.15205" year="1909">Kirkaldy (1909)</bibRefCitation>
: 137 (catalogue);
<bibRefCitation author="Distant, WL" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="0" pageNumber="79" refId="B12" refString="Distant, WL, 1918. Rhynchota (Vol. VII). Homoptera: appendix. Heteroptera: addenda. In: Shipley AE, Marshall AK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, [VII +] 210 pp." title="Rhynchota (Vol. VII). Homoptera: appendix. Heteroptera: addenda. In: Shipley AE, Marshall AK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, [VII +] 210 pp." year="1918">Distant (1918)</bibRefCitation>
: 145 (synonymisation, additional distribution);
<bibRefCitation author="Chatterjee, NC" journalOrPublisher="Indian Forest Records" pageId="0" pageNumber="79" pagination="1 - 31" refId="B6" refString="Chatterjee, NC, 1934. Entomological investigations on the spike disease of sandal (24). Pentatomidae (Hemipt.). Indian Forest Records 20: 1 - 31" title="Entomological investigations on the spike disease of sandal (24). Pentatomidae (Hemipt.)." volume="20" year="1934">Chatterjee (1934)</bibRefCitation>
: 22 (host record, new records);
<bibRefCitation author="Chandra, A" editor="Roonwal, ML" journalOrPublisher="Parts 9 - 21. Orders Isoptera, Embioptera, Psocoptera, Anoplura, Ephemeroptera, Odonata, Thysanoptera and Hemiptera. Government of India Press, Calcutta" pageId="0" pageNumber="79" pagination="83 - 102" refId="B5" refString="Chandra, A, 1953. Order Hemiptera (in part). Family Pentatomidae. In: Roonwal, ML, Mathur, RN, Bhasin, GD, Singh, B, Pant, GD, Chandra, A, Katiyar, KN, Mathur, RK, Eds., A Systematic Catalogue of the Main Identified Entomological Collection at the Forest Research Institute, Dehra Dun. Parts 9-21. Orders Isoptera, Embioptera, Psocoptera, Anoplura, Ephemeroptera, Odonata, Thysanoptera and Hemiptera. Government of India Press, Calcutta: 83 - 102" title="Order Hemiptera (in part). Family Pentatomidae." volumeTitle="A Systematic Catalogue of the Main Identified Entomological Collection at the Forest Research Institute, Dehra Dun." year="1953">Chandra (1953)</bibRefCitation>
(distribution);
<bibRefCitation author="Mathur, RN" journalOrPublisher="Indian Forest Bulletin" pageId="0" pageNumber="79" pagination="1 - 88" refId="B24" refString="Mathur, RN, Singh, B, 1960. A list of insect pests of forest plants in India and the adjacent countries (arranged alphabetically according to the plant genera and species, for the use of forest officers). Part 9. List of insect pests of plant genera S (Sabia to Syzygium). Indian Forest Bulletin 171 (8): 1 - 88" title="A list of insect pests of forest plants in India and the adjacent countries (arranged alphabetically according to the plant genera and species, for the use of forest officers). Part 9. List of insect pests of plant genera S (Sabia to Syzygium)." volume="171" year="1960">Mathur and Singh (1960)</bibRefCitation>
: 20 (host);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3924.1.1" author="Salini, S" journalOrPublisher="Zootaxa" pageId="0" pageNumber="79" pagination="1 - 76" refId="B33" refString="Salini, S, Viraktamath, CA, 2015. Genera of Pentatomidae (Hemiptera: Pentatomoidea) from south India- an illustrated key to genera and checklist of species. Zootaxa 3924 (1): 1 - 76, DOI: https://doi.org/10.11646/zootaxa.3924.1.1" title="Genera of Pentatomidae (Hemiptera: Pentatomoidea) from south India- an illustrated key to genera and checklist of species." url="https://doi.org/10.11646/zootaxa.3924.1.1" volume="3924" year="2015">Salini and Viraktamath (2015)</bibRefCitation>
: 12 (key to genera), 24 (checklist), 46, fig. 54 (illustration of dorsal habitus), 68, fig. 185 (illustration of external scent efferent system);
<bibRefCitation DOI="https://doi.org/10.1201/9780429061400-8" author="Salini, S" editor="Ramani, S" journalOrPublisher="A. Viraktamath's 75 th birthday, CRC Press, Taylor and Francis Group, Boca Ralton, London, New York" pageId="0" pageNumber="79" pagination="121 - 146" refId="B31" refString="Salini, S, 2019. Pentatomidae (Hemiptera: Heteroptera: Pentatomidea) of India. In: Ramani, S, Mohanraj, P, Yeshwanth, HM, Eds., Indian Insects Diversity and Science, a Festschrift for Professor C. A. Viraktamath's 75th birthday, CRC Press, Taylor and Francis Group, Boca Ralton, London, New York: 121 - 146, DOI: https://doi.org/10.1201/9780429061400-8" title="Pentatomidae (Hemiptera: Heteroptera: Pentatomidea) of India." url="https://doi.org/10.1201/9780429061400-8" volumeTitle="Indian Insects Diversity and Science, a Festschrift for Professor C." year="2019">Salini (2019)</bibRefCitation>
: 126, fig. 8.20 (illustration of dorsal habitus).
</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus" status="stat. rev.">Mycterizon bellus</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.21874" author="Breddin, G" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="79" pagination="250 - 309" refId="B3" refString="Breddin, G, 1909. Rhynchoten von Ceylon gesammelt von Dr. Walter Horn. Annales de la Societe Entomologique de Belgique 53: 250 - 309, DOI: https://doi.org/10.5962/bhl.part.21874" title="Rhynchoten von Ceylon gesammelt von Dr. Walter Horn." url="https://doi.org/10.5962/bhl.part.21874" volume="53" year="1909">Breddin (1909)</bibRefCitation>
: 279 (synonymisation);
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.15205" author="Kirkaldy, GW" journalOrPublisher="Journal of the Linnean Society of Zoology" pageId="0" pageNumber="79" refId="B20" refString="Kirkaldy, GW, 1909. Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families (Vol. I). Cimicidae. Felix L. Dames, Berlin, [xl +] 392 pp. https://doi.org/10.5962/bhl.title.15205" title="Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families (Vol. I). Cimicidae. Felix L. Dames, Berlin, [xl +] 392 pp." url="https://doi.org/10.5962/bhl.title.15205" year="1909">Kirkaldy (1909)</bibRefCitation>
: 371 (catalogue).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="type locality">
<paragraph pageId="0" pageNumber="79">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bella" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta bella</emphasis>
</taxonomicName>
. Ceylon (= Sri Lanka), Punduloya.
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius bellus</emphasis>
</taxonomicName>
. Ceylon (= Sri Lanka), Punduloya.
</paragraph>
<caption doi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79" start="Figures 63–68" startId="F13">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figures 63-68.</emphasis>
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon bellus</emphasis>
</taxonomicName>
(Distant), stat. rev.
<emphasis bold="true" pageId="0" pageNumber="79">63</emphasis>
habitus (dorsal)
<emphasis bold="true" pageId="0" pageNumber="79">64</emphasis>
habitus (ventral)
<emphasis bold="true" pageId="0" pageNumber="79">65</emphasis>
external scent efferent system (arrow showing the peritreme)
<emphasis bold="true" pageId="0" pageNumber="79">66</emphasis>
close up of posterior end of ventral side of male abdomen
<emphasis bold="true" pageId="0" pageNumber="79">67</emphasis>
close up of posterior end of ventral side of female abdomen (terminalia)
<emphasis bold="true" pageId="0" pageNumber="79">68</emphasis>
genital capsule (dorsal). Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="79">63, 64</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="79">65</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="79">66-68</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1148.95629.figures69-75" httpUri="https://binary.pensoft.net/fig/811002" pageId="0" pageNumber="79" start="Figures 69–75" startId="F14">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figures 69-75.</emphasis>
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon bellus</emphasis>
</taxonomicName>
(Distant), stat. rev.
<emphasis bold="true" pageId="0" pageNumber="79">69</emphasis>
genital capsule (ventral)
<emphasis bold="true" pageId="0" pageNumber="79">70</emphasis>
genital capsule (caudal)
<emphasis bold="true" pageId="0" pageNumber="79">71, 72</emphasis>
paramere
<emphasis bold="true" pageId="0" pageNumber="79">73</emphasis>
articulatory apparatus
<emphasis bold="true" pageId="0" pageNumber="79">74</emphasis>
phallus (dorsal)
<emphasis bold="true" pageId="0" pageNumber="79">75</emphasis>
phallus (ventral). Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; cp - capitate processes; ls - lobe-like ingrowths; pa - processes of aedeagus; dcp I - dorsal conjunctival processes (inner one); dcp II - dorsal conjunctival processes (outer one); lcp - lateral conjunctival processes; vcp - ventral conjunctival processes.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="type material">
<paragraph pageId="0" pageNumber="79">Type material.</paragraph>
<paragraph pageId="0" pageNumber="79">
Not examined, only type image was seen. The original description of
<taxonomicName lsidName="A. bella" pageId="0" pageNumber="79" rank="species" species="bella">
<emphasis italics="true" pageId="0" pageNumber="79">A. bella</emphasis>
</taxonomicName>
is based on a male specimen whereas that of
<taxonomicName lsidName="D. bellus" pageId="0" pageNumber="79" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">D. bellus</emphasis>
</taxonomicName>
was based on a female specimen. The syntype male specimen of
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bella" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta bella</emphasis>
</taxonomicName>
Distant, 1900a: 427 (Ceylon: Punduloya, Atkinson Coll., 92-6, NHMUK 013588914) herein designated as lectotype (Figs
<figureCitation captionStart="Figures 80–85" captionStartId="F16" captionText="Figures 80 - 85. Mycterizon bellus (Distant), stat. rev. 80 label data of lectotype 81 lectotype (dorsal habitus) 82 lectotype (ventral habitus) 83 label data of paralectotype 84 paralectotype (dorsal habitus) 85 paralectotype (ventral habitus). Photographed by Mick Webb, NHM, London, UK." figureDoi="10.3897/zookeys.1148.95629.figures80-85" httpUri="https://binary.pensoft.net/fig/811004" pageId="0" pageNumber="79">80-82</figureCitation>
) and the female syntype specimen of
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bella" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta bella</emphasis>
</taxonomicName>
Distant, 1900a: 427 (Ceylon: Punduloya, Atkinson Coll., 92-6, NHMUK 013588915) is herein designated as paraletotype (Figs
<figureCitation captionStart="Figures 80–85" captionStartId="F16" captionText="Figures 80 - 85. Mycterizon bellus (Distant), stat. rev. 80 label data of lectotype 81 lectotype (dorsal habitus) 82 lectotype (ventral habitus) 83 label data of paralectotype 84 paralectotype (dorsal habitus) 85 paralectotype (ventral habitus). Photographed by Mick Webb, NHM, London, UK." figureDoi="10.3897/zookeys.1148.95629.figures80-85" httpUri="https://binary.pensoft.net/fig/811004" pageId="0" pageNumber="79">83-85</figureCitation>
). Additionally, 31 specimens of this species are available in NHM (M Webb, pers. comm.). The image of the type is provided by courtesy of M Webb.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2008-10-22" collectingMethod="ex light trap" collectorName="Yeshwanth, H. M." country="India" location="Tirupathi" specimenCount="1" specimenCount-female="1" stateProvince="Andhra Pradesh">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="India">India</collectingCountry>
:
<collectingRegion country="India" name="Andhra Pradesh">Andhra Pradesh</collectingRegion>
</emphasis>
:
<specimenCount type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:A7553BB2CB33190F4A360A649B7538DF" country="India" name="Tirupathi" stateProvince="Andhra Pradesh">Tirupathi</location>
,
<collectingDate value="2008-10-22">22.x.2008</collectingDate>
,
<collectorName>Yeshwanth, H. M.</collectorName>
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp) (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2009-04-03" collectingMethod="ex light trap" collectorName="Yeshwanth" country="India" latitude="14.429833" location="Nellore" longLatPrecision="1" longitude="79.9987" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Andhra Pradesh">
<specimenCount type="female">3♀</specimenCount>
,
<specimenCount type="male">3♂</specimenCount>
ARS,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:B4FB227194E151B8D8CDFCFE3AEA9190" country="India" latitude="14.429833" longLatPrecision="1" longitude="79.9987" name="Nellore" stateProvince="Andhra Pradesh">Nellore</location>
,
<geoCoordinate degrees="14" direction="north" minutes="25.790" orientation="latitude" precision="1" value="14.429833">14°25.790'N</geoCoordinate>
,
<geoCoordinate degrees="079" direction="east" minutes="59.922" orientation="longitude" precision="1" value="79.9987">079°59.922'E</geoCoordinate>
,
<collectingDate value="2009-04-03">03.iv.2009</collectingDate>
,
<collectorName>Yeshwanth</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp) (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2009-04-05" collectorName="Yeshwanth, Goa" country="India" latitude="17.697584" location="Anakapalli" longLatPrecision="1" longitude="83.00422" specimenCount="2" specimenCount-male="2" stateProvince="Andhra Pradesh">
<specimenCount type="male">2♂</specimenCount>
, ARS,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:FF3F726388C6DED1924647B9552DA42B" country="India" latitude="17.697584" longLatPrecision="1" longitude="83.00422" name="Anakapalli" stateProvince="Andhra Pradesh">Anakapalli</location>
,
<geoCoordinate degrees="17" direction="north" minutes="41.855" orientation="latitude" precision="1" value="17.697584">17°41.855'N</geoCoordinate>
,
<geoCoordinate degrees="083" direction="east" minutes="00.253" orientation="longitude" precision="1" value="83.00422">083°00.253'E</geoCoordinate>
,
<collectingDate value="2009-04-05">5.iv.2009</collectingDate>
,
<collectorName>Yeshwanth</collectorName>
(NIM);
<emphasis bold="true" pageId="0" pageNumber="79">
<collectorName>Goa</collectorName>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="India" location="Karnataka" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount type="female">1♀</specimenCount>
, MRC. coll.
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="India" name="Karnataka">Karnataka</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2007-09-28" collectingMethod="ex light trap" collectorName="Naveen Kumar" country="India" location="Udupi" municipality="Brahmavara" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount type="female">3♀</specimenCount>
,
<specimenCount type="male">1♂</specimenCount>
,
<collectingMunicipality>Brahmavara</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:12B37230795A10E9E4CEA4A6003ED15B" country="India" municipality="Brahmavara" name="Udupi" stateProvince="Karnataka">Udupi</location>
,
<collectingDate value="2007-09-28">28.ix.2007</collectingDate>
,
<collectorName>Naveen Kumar</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp) (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2008-02-12" collectingMethod="ex light trap" collectorName="Naveen Kumar" country="India" location="Belgaum" municipality="Arabhavi" specimenCount="2" specimenCount-female="2" stateProvince="Karnataka">
<specimenCount type="female">2♀</specimenCount>
,
<collectingMunicipality>Arabhavi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:520FCE47C7CC2E4C455D3CCD2368675B" country="India" municipality="Arabhavi" name="Belgaum" stateProvince="Karnataka">Belgaum</location>
,
<collectingDate value="2008-02-12">12.ii.2008</collectingDate>
,
<collectorName>Naveen Kumar</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp) (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2008-02-13" collectorName="Yadava Babu" country="India" location="Belgaum" municipality="Arabhavi" specimenCount="2" specimenCount-male="2" stateProvince="Karnataka">
<specimenCount type="male">2♂</specimenCount>
, same data as former except
<collectingDate value="2008-02-13">13.ii.2008</collectingDate>
,
<collectorName>Yadava Babu</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2008-11-19" collectorName="Yeshwanth" country="India" latitude="11.70125" location="Hangla" longLatPrecision="1" longitude="76.62808" municipality="Bandipur" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Bandipur</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:EFEAD8A5844718842C92CE9787AF389D" country="India" latitude="11.70125" longLatPrecision="1" longitude="76.62808" municipality="Bandipur" name="Hangla" stateProvince="Karnataka">Hangla</location>
,
<collectingDate value="2008-11-19">19.xi.2008</collectingDate>
,
<geoCoordinate degrees="11" direction="north" minutes="42.075" orientation="latitude" precision="1" value="11.70125">11°42.075'N</geoCoordinate>
,
<geoCoordinate degrees="076" direction="east" minutes="37.685" orientation="longitude" precision="1" value="76.62808">076°37.685'E</geoCoordinate>
,
<collectorName>Yeshwanth</collectorName>
(NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2009-11-12" collectingMethod="ex light trap" collectorName="Markanja, D, Umeshkumar" country="India" location="Kannada" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount type="female">1♀</specimenCount>
,
<collectorName>Markanja, D</collectorName>
.
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:135E1E392522CDF72147F8F66FD135C4" country="India" name="Kannada" stateProvince="Karnataka">Kannada</location>
,
<collectingDate value="2009-11-12">12.xi.2009</collectingDate>
,
<collectorName>Umeshkumar</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp) (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2008-12-01" collectorName="Umeshkumar" country="India" location="Hessaraghata" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:21127EE3A7F5F8A65C64A7240B413166" country="India" name="Hessaraghata" stateProvince="Karnataka">Hessaraghata</location>
,
<collectingDate value="2008-12-01">1.xii.2008</collectingDate>
,
<collectorName>Umeshkumar</collectorName>
(NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2014-05-29" collectingMethod="ex light trap" collectorName="Naveen" country="India" elevation="79" latitude="13.416667" location="Udupi" longLatPrecision="1291" longitude="74.75" municipality="Brahmavara" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Karnataka">
<specimenCount type="male">7♂</specimenCount>
,
<specimenCount type="female">2♀</specimenCount>
,
<collectingMunicipality>Brahmavara</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:3A8191BF83322B8217BE15E88E860FB1" country="India" latitude="13.416667" longLatPrecision="1291" longitude="74.75" municipality="Brahmavara" name="Udupi" stateProvince="Karnataka">Udupi</location>
,
<geoCoordinate degrees="13" direction="north" minutes="25" orientation="latitude" precision="925" value="13.416667">13°25'N</geoCoordinate>
,
<geoCoordinate degrees="74" direction="east" minutes="45" orientation="longitude" precision="925" value="74.75">74°45'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="7.9" unit="m" value="79.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="7.9" unit="m" value="79.0">79 m</elevation>
</quantity>
,
<collectingDate value="2014-05-29">29.v.2014</collectingDate>
,
<collectorName>Naveen</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2018-04-06" collectorName="David, K. J." country="India" location="Vittal" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount type="female">1♀</specimenCount>
, CPCRI,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:77158A71EC9EB6FD8C33CB4B7A02B4D5" country="India" name="Vittal" stateProvince="Karnataka">Vittal</location>
,
<collectingDate value="2018-04-06">06.iv.2018</collectingDate>
,
<collectorName>David, K. J.</collectorName>
(NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2008-01-24" collectingMethod="ex light trap" collectorName="Naveen Kumar" country="Kozhikode" latitude="11.2779665" location="Kerala" longLatPrecision="1" longitude="75.84412" specimenCount="2" specimenCount-female="2" stateProvince="Kerala">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="India" name="Kerala">Kerala</collectingRegion>
</emphasis>
:
<specimenCount type="female">2♀</specimenCount>
,
<collectingCountry name="India">Kozhikode</collectingCountry>
,
<collectingDate value="2008-01-24">24.i.2008</collectingDate>
,
<geoCoordinate degrees="11" direction="north" minutes="16.678" orientation="latitude" precision="1" value="11.2779665">11°16.678'N</geoCoordinate>
,
<geoCoordinate degrees="075" direction="east" minutes="50.647" orientation="longitude" precision="1" value="75.84412">075°50.647'E</geoCoordinate>
,
<collectorName>Naveen Kumar</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp) (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2012-11-18" collectingMethod="ex light trap" collectorName="Devaraj" country="Calicut" latitude="11.416667" location="Calicut" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
, ZSI,
<collectingCountry name="India">Calicut</collectingCountry>
,
<geoCoordinate degrees="11" direction="north" minutes="25" orientation="latitude" precision="925" value="11.416667">11°25'N</geoCoordinate>
, 75°77'E,
<collectingDate value="2012-11-18">18.xi.2012</collectingDate>
,
<collectorName>Devaraj</collectorName>
, R.,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp), NIM-2022 (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2012-12-06" collectingMethod="ex light trap" collectorName="Umeshkumar, S." country="India" latitude="12.516666" location="Kasaragod" longLatPrecision="1293" longitude="74.95" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount type="male">1♂</specimenCount>
, CPCRI,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:076F7B20CF0F03F91F12C270CB57BE6C" country="India" latitude="12.516666" longLatPrecision="1293" longitude="74.95" name="Kasaragod" stateProvince="Kerala">Kasaragod</location>
,
<geoCoordinate degrees="12" direction="north" minutes="31" orientation="latitude" precision="925" value="12.516666">12°31'N</geoCoordinate>
,
<geoCoordinate degrees="74" direction="east" minutes="57" orientation="longitude" precision="925" value="74.95">74°57'E</geoCoordinate>
,
<collectingDate value="2012-12-06">6.xii.2012</collectingDate>
,
<collectorName>Umeshkumar, S.</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(sodium vapour lamp), NIM-2022 (NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2012-12-04" collectorName="Umeshkumar, S." country="India" latitude="12.516666" location="Kasaragod" longLatPrecision="1293" longitude="74.95" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount type="male">1♂</specimenCount>
, same data as before except date
<collectingDate value="2012-12-04">4.xii.2012</collectingDate>
(NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2014-04-24" collectingMethod="ex light trap" collectorName="Naveen" country="Calicut" location="Calicut" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kerala">
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
, ZSI,
<collectingCountry name="India">Calicut</collectingCountry>
,
<collectingDate value="2014-04-24">24.iv.2014</collectingDate>
, V.
<collectorName>Naveen</collectorName>
,
<collectingMethod>ex light trap</collectingMethod>
(NIM)
</materialsCitation>
;
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="India" location="Tamil Nadu" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu">
<specimenCount type="female">1♀</specimenCount>
, no label (NIM);
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2016-06-16" collectorName="Ramesh Kumar, A" country="India" location="Javadi Hills" municipality="Yelagiri" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu">
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Yelagiri</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:73F96188B34E5CBF9E5CB93F54979C9E:D80CD212A2D20611B40EC8029FD907FF" country="India" municipality="Yelagiri" name="Javadi Hills" stateProvince="Tamil Nadu">Javadi Hills</location>
,
<collectingDate value="2016-06-16">16.vi.2016</collectingDate>
,
<collectorName>Ramesh Kumar, A</collectorName>
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1148.95629.figures76-79" httpUri="https://binary.pensoft.net/fig/811003" pageId="0" pageNumber="79" start="Figures 76–79" startId="F15">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figures 76-79.</emphasis>
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon bellus</emphasis>
</taxonomicName>
(Distant), stat. rev.
<emphasis bold="true" pageId="0" pageNumber="79">76</emphasis>
phallus (lateral)
<emphasis bold="true" pageId="0" pageNumber="79">77</emphasis>
terminalia (dorsal)
<emphasis bold="true" pageId="0" pageNumber="79">78</emphasis>
terminalia (ventral)
<emphasis bold="true" pageId="0" pageNumber="79">79</emphasis>
spermatheca. Scale bar:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
. Abbreviations: ad - aedeagus; ar - apical receptacle; rs - ring sclerites; lt VIII - laterotergite VIII; lt IX - laterotergite IX; vf VIII - valvifers VIII; vf IX - valvifers IX.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="redescription">
<paragraph pageId="0" pageNumber="79">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">Colouration</emphasis>
(Figs
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">63</figureCitation>
,
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">64</figureCitation>
). Dorsum ochraceous with black irregular spots scattered over pronotum, scutellum and hemelytra; head ochraceous except narrow lateral margins of mandibular plates, two oblique, narrow longitudinal lines on disc of clypeus, black; antennae ochraceous; lateral margins of pronotum adjacent to humeri (including humeri), black; disc of scutellum with roughly anchor-shaped spot and one short oblique stripe each at basal angles of scutellum, black; connexivum with narrow, transverse, black stripe on anterior and posterior margin of connexivial segments, black; sometimes anterior 1/2 margin of each pleurite III-VII with longitudinal stripe, black; distal end of hemelytra with moderate size spot, ochraceous, sometimes with purplish shade; membrane smoky brown; ventral side ochraceous with the following markings: thoracic pleura laterally with irregular markings mainly formed by the coarse punctures, one roughly circular spot mesad at an oblique angle to each spiracles; short, narrow and transverse stripe below each spiracles on either side of abdominal sternites III-VII and anterior and posterior angles of each abdominal sternites III-VII, black; labium ochraceous with apex black; legs ochraceous except small nearly circular spots densely distributed on femora and tibiae (finer on tibiae) and apical 1/2 of claws, black.
</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">
<emphasis italics="true" pageId="0" pageNumber="79">Integument and vestiture</emphasis>
.
</emphasis>
Body above covered with coarse, black punctures confluent to form irregular markings on dorsum especially on pronotum and scutellum. Ventral side of body with coarse and dense black punctures mostly concentrated towards lateral sides on thoracic sternites. Femora of legs with coarse, black, dense punctures and tibiae with fine black punctures. The punctures finer on abdominal sternites, concentrated towards laterally. Ventral side of genital capsule including posterolateral lobes with fine pale brown punctures; terminalia with valvifers VIII and laterotergite VIII with sparse, coarse, brown punctures.
</paragraph>
<paragraph pageId="0" pageNumber="79">Body glabrous, except the following: antennae with fine, golden pubescence; legs with setae moderately elongate and semi erect; tibiae with dense, brown, short, semi erect setae on ventral side and dorsally with moderately elongate brown setae, dense mat of setae at the posterior end of tibiae; tarsomeres on dorsal side with moderately elongate golden setae and dense mat of short, golden setae on ventral side of all tarsomeres; labium with sparse, golden setae. Genital capsule (middle margin of dorsal rim, posterolateral lobes and the caudal 1/2 of genital capsule on ventral side) with moderately elongate, dense, semi erect, golden setae. Female genitalia (valvifers VIII, valvifers IX, laterotergite VIII and IX) with moderately long, golden semi-erect setae.</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">
<emphasis italics="true" pageId="0" pageNumber="79">Structure</emphasis>
.
</emphasis>
See the generic redescription.
</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">Male genitalia</emphasis>
(Figs
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">66</figureCitation>
,
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">68</figureCitation>
-
<figureCitation captionStart="Figures 76–79" captionStartId="F15" captionText="Figures 76 - 79. Mycterizon bellus (Distant), stat. rev. 76 phallus (lateral) 77 terminalia (dorsal) 78 terminalia (ventral) 79 spermatheca. Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; ar - apical receptacle; rs - ring sclerites; lt VIII - laterotergite VIII; lt IX - laterotergite IX; vf VIII - valvifers VIII; vf IX - valvifers IX." figureDoi="10.3897/zookeys.1148.95629.figures76-79" httpUri="https://binary.pensoft.net/fig/811003" pageId="0" pageNumber="79">76</figureCitation>
). Genital capsule (Figs
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">68</figureCitation>
-
<figureCitation captionStart="Figures 69–75" captionStartId="F14" captionText="Figures 69 - 75. Mycterizon bellus (Distant), stat. rev. 69 genital capsule (ventral) 70 genital capsule (caudal) 71, 72 paramere 73 articulatory apparatus 74 phallus (dorsal) 75 phallus (ventral). Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; cp - capitate processes; ls - lobe-like ingrowths; pa - processes of aedeagus; dcp I - dorsal conjunctival processes (inner one); dcp II - dorsal conjunctival processes (outer one); lcp - lateral conjunctival processes; vcp - ventral conjunctival processes." figureDoi="10.3897/zookeys.1148.95629.figures69-75" httpUri="https://binary.pensoft.net/fig/811002" pageId="0" pageNumber="79">70</figureCitation>
) nearly quadrangular with posterolateral lobes (= caudal lobes) well developed and broadly rounded. Dorsal rim widely and deeply excavated concave with almost straight middle margin; infoldings of dorsal rim submedially with rounded lobe-like ingrowths (ls) towards posterior aperture. Ventral rim medially shallowly excavated and broadly concave.
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">Paramere</emphasis>
(Figs
<figureCitation captionStart="Figures 69–75" captionStartId="F14" captionText="Figures 69 - 75. Mycterizon bellus (Distant), stat. rev. 69 genital capsule (ventral) 70 genital capsule (caudal) 71, 72 paramere 73 articulatory apparatus 74 phallus (dorsal) 75 phallus (ventral). Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; cp - capitate processes; ls - lobe-like ingrowths; pa - processes of aedeagus; dcp I - dorsal conjunctival processes (inner one); dcp II - dorsal conjunctival processes (outer one); lcp - lateral conjunctival processes; vcp - ventral conjunctival processes." figureDoi="10.3897/zookeys.1148.95629.figures69-75" httpUri="https://binary.pensoft.net/fig/811002" pageId="0" pageNumber="79">71</figureCitation>
,
<figureCitation captionStart="Figures 69–75" captionStartId="F14" captionText="Figures 69 - 75. Mycterizon bellus (Distant), stat. rev. 69 genital capsule (ventral) 70 genital capsule (caudal) 71, 72 paramere 73 articulatory apparatus 74 phallus (dorsal) 75 phallus (ventral). Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; cp - capitate processes; ls - lobe-like ingrowths; pa - processes of aedeagus; dcp I - dorsal conjunctival processes (inner one); dcp II - dorsal conjunctival processes (outer one); lcp - lateral conjunctival processes; vcp - ventral conjunctival processes." figureDoi="10.3897/zookeys.1148.95629.figures69-75" httpUri="https://binary.pensoft.net/fig/811002" pageId="0" pageNumber="79">72</figureCitation>
). Sclerotised, crown broad, subquadrate with one moderately elongate finger-like process at distal end and another short, stout, apically rounded process forming an acute angle with the distal finger-like process, short neck and small plate-like apodeme.
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">Articulatory apparatus</emphasis>
(Fig.
<figureCitation captionStart="Figures 69–75" captionStartId="F14" captionText="Figures 69 - 75. Mycterizon bellus (Distant), stat. rev. 69 genital capsule (ventral) 70 genital capsule (caudal) 71, 72 paramere 73 articulatory apparatus 74 phallus (dorsal) 75 phallus (ventral). Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; cp - capitate processes; ls - lobe-like ingrowths; pa - processes of aedeagus; dcp I - dorsal conjunctival processes (inner one); dcp II - dorsal conjunctival processes (outer one); lcp - lateral conjunctival processes; vcp - ventral conjunctival processes." figureDoi="10.3897/zookeys.1148.95629.figures69-75" httpUri="https://binary.pensoft.net/fig/811002" pageId="0" pageNumber="79">73</figureCitation>
). Support bridge complex excavated ventrally cup-like, for which the narrow, elongate, oval capitate processes (cp) attached with short dorsal connectives.
<emphasis bold="true" pageId="0" pageNumber="79">
<emphasis italics="true" pageId="0" pageNumber="79">Phallus</emphasis>
.
</emphasis>
Phallotheca longer than broad, narrowed towards proximal end, dorsal region convex and ventral margin nearly straight (in lateral view); thecal process slightly developed on mid dorsal side at the distal end of phallotheca; a pair of minute, stout, sclerotised, ventrolateral tubercles at the posterior end of the phallotheca; four pairs of conjunctival processes; two pairs on dorsal side (inner pair, dcp I and outer pair dcp II), one pair each on ventral side (vcp) and lateral side (lcp) respectively. inner pair of dorsal conjunctival processes (dcp I) longest, broad and membranous, narrowed towards distal end; outer pair (dcp II) short, narrow, sclerotised with distal apex acuminate and membranous; ventral ones (vcp) moderately elongate, sclerotised and apically rounded; lateral pairs (lcp) shortest and membranous; processes of aedeagus (pa) fused mid longitudinally and sclerotised except median longitudinal part; partially enclosing aedeagus (ad) at the proximal end; aedeagus (ad) moderately elongate, sclerotised and tube-like with oblique phallotreme.
</paragraph>
<caption doi="10.3897/zookeys.1148.95629.figures80-85" httpUri="https://binary.pensoft.net/fig/811004" pageId="0" pageNumber="79" start="Figures 80–85" startId="F16">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figures 80-85.</emphasis>
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon bellus</emphasis>
</taxonomicName>
(Distant), stat. rev.
<emphasis bold="true" pageId="0" pageNumber="79">80</emphasis>
label data of lectotype
<emphasis bold="true" pageId="0" pageNumber="79">81</emphasis>
lectotype (dorsal habitus)
<emphasis bold="true" pageId="0" pageNumber="79">82</emphasis>
lectotype (ventral habitus)
<emphasis bold="true" pageId="0" pageNumber="79">83</emphasis>
label data of paralectotype
<emphasis bold="true" pageId="0" pageNumber="79">84</emphasis>
paralectotype (dorsal habitus)
<emphasis bold="true" pageId="0" pageNumber="79">85</emphasis>
paralectotype (ventral habitus). Photographed by Mick Webb, NHM, London, UK.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">
<emphasis italics="true" pageId="0" pageNumber="79">Female genitalia</emphasis>
.
</emphasis>
(Figs
<figureCitation captionStart="Figures 63–68" captionStartId="F13" captionText="Figures 63 - 68. Mycterizon bellus (Distant), stat. rev. 63 habitus (dorsal) 64 habitus (ventral) 65 external scent efferent system (arrow showing the peritreme) 66 close up of posterior end of ventral side of male abdomen 67 close up of posterior end of ventral side of female abdomen (terminalia) 68 genital capsule (dorsal). Scale bars: 1 mm (63, 64); 0.5 mm (65); 0.1 mm (66 - 68)." figureDoi="10.3897/zookeys.1148.95629.figures63-68" httpUri="https://binary.pensoft.net/fig/811001" pageId="0" pageNumber="79">67</figureCitation>
,
<figureCitation captionStart="Figures 76–79" captionStartId="F15" captionText="Figures 76 - 79. Mycterizon bellus (Distant), stat. rev. 76 phallus (lateral) 77 terminalia (dorsal) 78 terminalia (ventral) 79 spermatheca. Scale bar: 0.1 mm. Abbreviations: ad - aedeagus; ar - apical receptacle; rs - ring sclerites; lt VIII - laterotergite VIII; lt IX - laterotergite IX; vf VIII - valvifers VIII; vf IX - valvifers IX." figureDoi="10.3897/zookeys.1148.95629.figures76-79" httpUri="https://binary.pensoft.net/fig/811003" pageId="0" pageNumber="79">77-79</figureCitation>
). Valvifers VIII (vf VIII) transverse, broad and roughly quadrangular, with medial margins nearly straight; inner posterolateral angles rounded; posterior margin nearly straight; valvifers IX (vf IX) single, transverse, broad sclerite; laterotergites IX (lt IX) oblique, elongate with inner and outer margin straight, caudal margin orbicular or rounded, caudal apex smooth, without any denticle; laterotergite VIII (lt VIII) subtriangular, caudal margin smooth, slightly convex. A pair of ring sclerites (rs) oblong.
<emphasis bold="true" pageId="0" pageNumber="79">
<emphasis italics="true" pageId="0" pageNumber="79">Spermatheca</emphasis>
.
</emphasis>
Spermathecal dilation long, regularly, obliquely fluted; proximal end of sclerotised rod, at the middle of lumen of spermathecal dilation, curved upwards and continued as distal spermathecal duct; proximal flange 1/2 the size of distal flange; apical receptacle (ar) orbicular with three elongate ductules, all facing towards distal rim and surpassing distal rim.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">(mm).</emphasis>
Males (
<emphasis italics="true" pageId="0" pageNumber="79">n</emphasis>
= 5) as median (minimum-maximum). Body length 8.63 (8.08-9.09); head: length 1.90 (1.82-1.98), width (including eyes) 2.91 (2.85-3.01), interocular width 1.74 (1.65-1.78); lengths of antennomeres: I - 0.56 (0.46-0.68), II - 0.68 (0.54-0.77), III - 1.25 (1.20-1.33), IV - 1.58 (1.47-1.68), V - 1.55 (1.50-1.63); lengths of labiomeres: I - 0.63 (0.51-0.73), II - 0.86 (0.79-0.93), III - 0.60 (0.56-0.63), IV - 0.52 (0.46-0.60); pronotum: length 2.12 (1.92-2.33), width (including humeri) 5.07 (4.82-5.32); scutellum: length 4.17 (3.84-4.31), width (at basal angles) 3.33 (3.28-3.48).
</paragraph>
<paragraph pageId="0" pageNumber="79">
Females (
<emphasis italics="true" pageId="0" pageNumber="79">n</emphasis>
= 5) as median (minimum-maximum). Body length 10.26 (9.83-10.73); head: length 2.05 (2.01-2.13), width (including eyes) 3.19 (3.12-3.37), interocular width 1.99 (1.97-2.04); lengths of antennomeres: I -0.59 (0.55-0.64), II - 0.68 (0.56-0.74), III - 1.35 (1.26-1.44), IV - 1.72 (1.71-1.76), V -1.74 (1.73-1.75); lengths of labiomeres: I -0.61 (0.54-0.66), II -0.97 (0.91-1.04), III - 0.61 (0.56-0.75), IV - 0.53 (0.51-0.60); pronotum: length 2.63 (2.40-2.78), width (including humeri) 5.95 (5.62-6.44); scutellum: length 4.94 (4.64-5.31), width (at basal angles) 3.90 (3.70-4.11).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="differential diagnosis and remarks">
<paragraph pageId="0" pageNumber="79">Differential diagnosis and remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
Though the species
<taxonomicName class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta bellus</emphasis>
</taxonomicName>
neither possess the basal abdominal tubercle nor the metasternal cruciform process,
<bibRefCitation DOI="https://doi.org/10.1080/00222930008678310" author="Distant, WL" journalOrPublisher="Annals &amp; Magazine of Natural History" pageId="0" pageNumber="79" pagination="420 - 435" refId="B8" refString="Distant, WL, 1900a. Rhynchotal Notes. - IV. Heteroptera: Pentatominae (part). Annals &amp; Magazine of Natural History 5 (7): 420 - 435, DOI: https://doi.org/10.1080/00222930008678310" title="Rhynchotal Notes. - IV. Heteroptera: Pentatominae (part)." url="https://doi.org/10.1080/00222930008678310" volume="5" year="1900 a">Distant (1900a)</bibRefCitation>
placed
<taxonomicName class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta bellus</emphasis>
</taxonomicName>
in
<taxonomicName authorityName="Walker" authorityYear="1867" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta</emphasis>
</taxonomicName>
; however, he mentioned in the redescription of
<taxonomicName authorityName="Walker" authorityYear="1867" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta</emphasis>
</taxonomicName>
:
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
Metasternum with central, cruciform process, not notched posteriorly. Second abdominal segment with a central, broad, obtuse, spinous tubercle...&quot; Later,
<bibRefCitation author="Distant, WL" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="0" pageNumber="79" refId="B11" refString="Distant, WL, 1902. Rhynchota (Vol. I). Heteroptera. In: Blanford WT (Ed.) The Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, [XXXVII +] 438 pp." title="Rhynchota (Vol. I). Heteroptera. In: Blanford WT (Ed.) The Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, [XXXVII +] 438 pp." year="1902">Distant (1902)</bibRefCitation>
proposed the genus
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
and the original description of
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
again mentioned the characters of the central cruciform process and the basal abdominal tubercle, but he transferred
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius bellus</emphasis>
</taxonomicName>
from
<taxonomicName authorityName="Walker" authorityYear="1867" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.21874" author="Breddin, G" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="79" pagination="250 - 309" refId="B3" refString="Breddin, G, 1909. Rhynchoten von Ceylon gesammelt von Dr. Walter Horn. Annales de la Societe Entomologique de Belgique 53: 250 - 309, DOI: https://doi.org/10.5962/bhl.part.21874" title="Rhynchoten von Ceylon gesammelt von Dr. Walter Horn." url="https://doi.org/10.5962/bhl.part.21874" volume="53" year="1909">Breddin (1909)</bibRefCitation>
recognised
<taxonomicName lsidName="D. bellus" pageId="0" pageNumber="79" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">D. bellus</emphasis>
</taxonomicName>
as a distinct species (see comments under
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon</emphasis>
</taxonomicName>
) and found that it was not congeneric with
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
, hence proposed
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon</emphasis>
</taxonomicName>
to accommodate it. However,
<bibRefCitation author="Distant, WL" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="0" pageNumber="79" refId="B12" refString="Distant, WL, 1918. Rhynchota (Vol. VII). Homoptera: appendix. Heteroptera: addenda. In: Shipley AE, Marshall AK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, [VII +] 210 pp." title="Rhynchota (Vol. VII). Homoptera: appendix. Heteroptera: addenda. In: Shipley AE, Marshall AK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, [VII +] 210 pp." year="1918">Distant (1918)</bibRefCitation>
synonymised this genus with
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
and transferred
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius bellus</emphasis>
</taxonomicName>
back to
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
without justifying his action. The present study is in agreement with
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.21874" author="Breddin, G" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="79" pagination="250 - 309" refId="B3" refString="Breddin, G, 1909. Rhynchoten von Ceylon gesammelt von Dr. Walter Horn. Annales de la Societe Entomologique de Belgique 53: 250 - 309, DOI: https://doi.org/10.5962/bhl.part.21874" title="Rhynchoten von Ceylon gesammelt von Dr. Walter Horn." url="https://doi.org/10.5962/bhl.part.21874" volume="53" year="1909">Breddin (1909)</bibRefCitation>
(see comments under
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon</emphasis>
</taxonomicName>
) and therefore
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon bellus</emphasis>
</taxonomicName>
(Distant, 1902) is removed from synonymy with
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius bellus</emphasis>
</taxonomicName>
(Distant, 1902). The single male syntype of
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Araducta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Araducta bella" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="0" pageNumber="79">Araducta bella</emphasis>
</taxonomicName>
, presently lodged in NHM, London, UK, is herewith designated as lectotype and the female syntype as paralectotype, also currently in NHM, London, UK.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="distribution">
<paragraph pageId="0" pageNumber="79">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="79">
India: Bihar, Odisha, Tamil Nadu (
<bibRefCitation author="Distant, WL" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="0" pageNumber="79" refId="B12" refString="Distant, WL, 1918. Rhynchota (Vol. VII). Homoptera: appendix. Heteroptera: addenda. In: Shipley AE, Marshall AK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, [VII +] 210 pp." title="Rhynchota (Vol. VII). Homoptera: appendix. Heteroptera: addenda. In: Shipley AE, Marshall AK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, [VII +] 210 pp." year="1918">Distant 1918</bibRefCitation>
); Ceylon (= Sri Lanka) (
<bibRefCitation DOI="https://doi.org/10.1080/00222930008678310" author="Distant, WL" journalOrPublisher="Annals &amp; Magazine of Natural History" pageId="0" pageNumber="79" pagination="420 - 435" refId="B8" refString="Distant, WL, 1900a. Rhynchotal Notes. - IV. Heteroptera: Pentatominae (part). Annals &amp; Magazine of Natural History 5 (7): 420 - 435, DOI: https://doi.org/10.1080/00222930008678310" title="Rhynchotal Notes. - IV. Heteroptera: Pentatominae (part)." url="https://doi.org/10.1080/00222930008678310" volume="5" year="1900 a">Distant 1900a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="tribal placement of mycterizon">
<paragraph pageId="0" pageNumber="79">
Tribal placement of
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="79">
The genus
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
originally included four species, viz.,
<taxonomicName lsidName="D. bellus" pageId="0" pageNumber="79" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">D. bellus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="D. fulvescens" pageId="0" pageNumber="79" rank="species" species="fulvescens">
<emphasis italics="true" pageId="0" pageNumber="79">D. fulvescens</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="D. minor" pageId="0" pageNumber="79" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="79">D. minor</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="D. sordida" pageId="0" pageNumber="79" rank="species" species="sordida">
<emphasis italics="true" pageId="0" pageNumber="79">D. sordida</emphasis>
</taxonomicName>
(Kirby) (Rider, 2022). The aforementioned nomenclatural changes and new taxa descriptions (
<taxonomicName lsidName="A. sordida" pageId="0" pageNumber="79" rank="species" species="sordida">
<emphasis italics="true" pageId="0" pageNumber="79">A. sordida</emphasis>
</taxonomicName>
(Kirby, 1891), comb. nov;
<taxonomicName lsidName="M. bellus" pageId="0" pageNumber="79" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">M. bellus</emphasis>
</taxonomicName>
(Distant, 1902), stat. rev.;
<taxonomicName lsidName="D. laticeps" pageId="0" pageNumber="79" rank="species" species="laticeps">
<emphasis italics="true" pageId="0" pageNumber="79">D. laticeps</emphasis>
</taxonomicName>
(Zheng &amp; Liu, 1987), comb. nov.;
<taxonomicName lsidName="D. tridentatus" pageId="0" pageNumber="79" rank="species" species="tridentatus">
<emphasis italics="true" pageId="0" pageNumber="79">D. tridentatus</emphasis>
</taxonomicName>
(Xiong &amp; Liu, 1995), comb. nov.;
<taxonomicName lsidName="D. trifasciatus" pageId="0" pageNumber="79" rank="species" species="trifasciatus">
<emphasis italics="true" pageId="0" pageNumber="79">D. trifasciatus</emphasis>
</taxonomicName>
(Xiong &amp; Liu, 1995), comb. nov.; and
<taxonomicName authorityName="Salini &amp; Rabbani" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius barpetensis" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="barpetensis">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius barpetensis</emphasis>
</taxonomicName>
Salini &amp; Rabbani, sp. nov.) led to the addition of two more species to the genus
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Dunnius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dunnius" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Dunnius</emphasis>
</taxonomicName>
. The genus
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon</emphasis>
</taxonomicName>
, monotypic with
<taxonomicName lsidName="M. bellus" pageId="0" pageNumber="79" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">M. bellus</emphasis>
</taxonomicName>
, is designated as type species by monotypy and presently considered as a member of
<taxonomicName lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Menidini">Menidini</taxonomicName>
, although its tribal placement is dubious.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.21874" author="Breddin, G" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="79" pagination="250 - 309" refId="B3" refString="Breddin, G, 1909. Rhynchoten von Ceylon gesammelt von Dr. Walter Horn. Annales de la Societe Entomologique de Belgique 53: 250 - 309, DOI: https://doi.org/10.5962/bhl.part.21874" title="Rhynchoten von Ceylon gesammelt von Dr. Walter Horn." url="https://doi.org/10.5962/bhl.part.21874" volume="53" year="1909">Breddin (1909)</bibRefCitation>
opined that
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon</emphasis>
</taxonomicName>
is closely related to
<taxonomicName authorityName="Stal" authorityYear="1864" class="Insecta" family="Pentatomidae" genus="Antestia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Antestia" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Antestia</emphasis>
</taxonomicName>
, a member of
<taxonomicName authorityName="Distant" authorityYear="1902" lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Antestiini">Antestiini</taxonomicName>
. The tribe
<taxonomicName lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Menidini">Menidini</taxonomicName>
is a large and variable one, the monophyly of which has not yet been proved. It was formally named by
<bibRefCitation author="Atkinson, ET" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="79" pagination="118 - 184" refId="B1" refString="Atkinson, ET, 1888. Notes on Indian Rhynchota: Heteroptera. No. 2 &amp; 3. Journal of the Asiatic Society of Bengal 57 (2): 118 - 184" title="Notes on Indian Rhynchota: Heteroptera. No. 2 &amp; 3." volume="57" year="1888">Atkinson (1888)</bibRefCitation>
as Menidaria and was defined based on a wide variety of characters: basal abdominal segment is obtusely convex or with a slender spine or rounded or compressed; if tuberculate, tibiae rounded; very rarely obtusely tuberculate at middle; metasternum not elevated, mesosternal ridge with uniform breadth throughout its length; labium usually extending beyond mid coxae, either reaching or extending beyond basal abdominal segment; third labial segment sometimes slightly longer than second; tibiae usually sulcate, if rounded, basal abdominal sternum with a spine, which usually does not extend beyond mid coxae.
</paragraph>
<paragraph pageId="0" pageNumber="79">
Based on the examination of Indian
<taxonomicName lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Menidini">Menidini</taxonomicName>
and perusal of literature revealed that the members of this tribe do not appear to share a common ancestor and can be broadly assembled into three groups based on the combination of characters like shape of the scutellum, presence/absence of abdominal spine/tubercle, development of metasternal carinae:
</paragraph>
<paragraph pageId="0" pageNumber="79">Group 1 - spatulate scutellum, elongate abdominal spine present, metasternal carina narrow throughout its length;</paragraph>
<paragraph pageId="0" pageNumber="79">Group 2 - subtriangular scutellum, elongate abdominal spine present, metasternal carina narrow throughout its length;</paragraph>
<paragraph pageId="0" pageNumber="79">Group 3 - subtriangular scutellum, abdominal tubercle present, apex of which is placed in metasternal carina (broadening posteriorly).</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Salini &amp; Gracy &amp; Akoijam &amp; Rabbani &amp; David &amp; Roca-Cusachs" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1902" class="Insecta" family="Pentatomidae" genus="Mycterizon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mycterizon bellus" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="bellus">
<emphasis italics="true" pageId="0" pageNumber="79">Mycterizon bellus</emphasis>
</taxonomicName>
does not fit into any of the above-mentioned groups, but rather possesses a unique combination of characters such as a spatulate scutellum, a narrow metasternal carina, and the absence of either an abdominal spine or a tubercle. It also possesses other characters such as the head flat above, as broad as 1.5 times the head length, the external scent efferent system with peritreme well developed, short, nearly reaching mid-metapleuron, spout-like, obliquely elevated to the pleural surface, the shape of the genital capsule including a dorsal rim (slightly concave medially) and ventral rim (broadly concave, without denticles on infoldings), the shape of parameres (broad crown-shaped with stout, straight, finger-like process), and the phallus with elongate conjunctival processes. Considering these remarkable differences, it cannot be considered as a member of
<taxonomicName lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Menidini">Menidini</taxonomicName>
. Therefore, it is removed from the tribe and its tribal status cannot be decided except through phylogenetic studies using both morphological and molecular characters of the taxa included in
<taxonomicName lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Menidini">Menidini</taxonomicName>
and other related tribes.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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