treatments-xml/data/AD/5B/CC/AD5BCC59AE2FFFC4FF0DFF2953C0F81E.xml

<document ID-DOI="10.11646/zootaxa.5278.3.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906748" ID-ZooBank="8397A532-CFAB-4BCA-AF98-D76BE5079B04" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683531519212" checkinUser="plazi" docAuthor="Bitar, Murilo Victor Silva, Mendonça, Maria Thayane Da Silva &amp; Fernandes, José Antônio Marin" docDate="2023" docId="AD5BCC59AE2FFFC4FF0DFF2953C0F81E" docLanguage="en" docName="zootaxa.5278.3.9.pdf" docOrigin="Zootaxa 5278 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.3.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessa holochlorata Bitar &amp; Mendonça &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="593" masterDocId="5162B421AE20FFCBFF9AFFBE5775FF88" masterDocTitle="Description of seven new species of Edessa Fabricius, 1803 from Central America (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="600" masterPageNumber="578" pageNumber="593" updateTime="1683532061405" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Description of seven new species of Edessa Fabricius, 1803 from Central America (Heteroptera, Pentatomidae, Edessinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bitar, Murilo Victor Silva</mods:namePart>
<mods:affiliation>murilobioufpa @ gmail. com; https: // orcid. org / 0000 - 0002 - 1109 - 4386</mods:affiliation>
<mods:nameIdentifier type="email">murilobioufpa@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-6504-247X</mods:nameIdentifier>
<mods:affiliation>thaymendonca 4 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6504 - 247 X</mods:affiliation>
<mods:nameIdentifier type="email">thaymendonca4@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fernandes, José Antônio Marin</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7450-5296</mods:nameIdentifier>
<mods:affiliation>joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
<mods:nameIdentifier type="email">joseamf@ufpa.br</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5278</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:start>578</mods:start>
<mods:end>600</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.3.9</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5278.3.9</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7906748</mods:identifier>
<mods:identifier type="ZooBank">8397A532-CFAB-4BCA-AF98-D76BE5079B04</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7906782" ID-Zenodo-Dep="7906782" LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E" httpUri="http://treatment.plazi.org/id/AD5BCC59AE2FFFC4FF0DFF2953C0F81E" lastPageNumber="593" pageId="15" pageNumber="593">
<subSubSection box="[151,472,151,178]" pageId="15" pageNumber="593" type="nomenclature">
<paragraph blockId="15.[151,472,151,214]" box="[151,472,151,178]" pageId="15" pageNumber="593">
<heading bold="true" box="[151,472,151,178]" fontSize="11" level="1" pageId="15" pageNumber="593" reason="1">
<emphasis bold="true" box="[151,472,151,178]" pageId="15" pageNumber="593">
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[151,398,151,178]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="593" phylum="Arthropoda" rank="species" species="holochlorata" status="sp. n.">
<emphasis bold="true" box="[151,398,151,178]" italics="true" pageId="15" pageNumber="593">Edessa holochlorata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[405,472,151,177]" pageId="15" pageNumber="593" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,399,187,214]" pageId="15" pageNumber="593" type="description">
<paragraph blockId="15.[151,472,151,214]" box="[151,399,187,214]" pageId="15" pageNumber="593">
(
<figureCitation box="[159,239,187,214]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Figs. 7</figureCitation>
; 
<figureCitation box="[252,324,187,214]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">9 C, F</figureCitation>
; 
<figureCitation box="[336,390,187,213]" captionStart="FIGURE 10" captionStartId="19.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@19.[203,1385,181,1919]" captionTargetPageId="19" captionText="FIGURE 10. A–C. Distribution map of the species: A, Edessa bituberculata sp. n., Edessa fuscolimbata sp. n., Edessa fuscopunctatasp.n.;B,Edessabivenulatasp.n.,Edessaenargocelidasp.n.,Edessaholochloratasp.n.;C,Edessahelvoalatasp.n." figureDoi="http://doi.org/10.5281/zenodo.7906784" httpUri="https://zenodo.org/record/7906784/files/figure.png" pageId="15" pageNumber="593">10 B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,1368,259,286]" pageId="15" pageNumber="593" type="etymology">
<paragraph blockId="15.[151,1437,259,1942]" box="[151,1368,259,286]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[151,286,259,285]" pageId="15" pageNumber="593">Etymology.</emphasis>
The name refers to the uniform dorsal color of the body (Gr. 
<emphasis box="[974,1041,259,285]" italics="true" pageId="15" pageNumber="593">Holus</emphasis>
- whole; Gr. 
<taxonomicName authorityName="Giglio-Tos" authorityYear="1898" box="[1183,1273,259,285]" class="Insecta" family="Acrididae" genus="Chlorus" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis box="[1183,1273,259,285]" italics="true" pageId="15" pageNumber="593">Chlorus</emphasis>
</taxonomicName>
- green).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="593" type="materials_examined">
<paragraph blockId="15.[151,1437,259,1942]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[199,433,295,321]" pageId="15" pageNumber="593">Material examined.</emphasis>
<materialsCitation collectingDate="1992-11" collectionCode="INBio" collectorName="G. Carballo" country="Costa Rica" elevation="100" location="Limon" pageId="15" pageNumber="593" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
<typeStatus box="[440,544,295,321]" pageId="15" pageNumber="593">Holotype</typeStatus>
male. 
<emphasis bold="true" box="[621,896,294,321]" pageId="15" pageNumber="593">
<collectingCountry box="[621,794,295,321]" name="Costa Rica" pageId="15" pageNumber="593">COSTA RICA</collectingCountry>
, 
<collectingRegion box="[807,891,294,321]" country="Costa Rica" name="Limon" pageId="15" pageNumber="593">Limón</collectingRegion>
:
</emphasis>
1♁, Est. Hitoy-Cerere, 
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<elevation box="[1166,1232,295,321]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="593" unit="m" value="100.0">100m</elevation>
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, Res. Biol. Hitoy-Cerere, 
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<collectingDate box="[240,336,331,357]" bridgedPair="/" pageId="15" pageNumber="593" value="1992-11">XI/1992</collectingDate>
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, 
<collectorName box="[347,480,331,357]" pageId="15" pageNumber="593">G. Carballo</collectorName>
, L-N 184200, 643300, CRI000 926730 (
<collectionCode box="[936,1012,331,357]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="15" pageNumber="593">INBio</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="15.[151,1437,259,1942]" pageId="15" pageNumber="593">
<materialsCitation collectingDate="1995-11" collectionCode="INBio" collectorName="G. Gallardo" country="Costa Rica" elevation="70" location="Amburi" pageId="15" pageNumber="593" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
<typeStatus box="[199,312,368,394]" pageId="15" pageNumber="593">Paratypes</typeStatus>
. 
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<collectingCountry box="[326,502,367,393]" name="Costa Rica" pageId="15" pageNumber="593">COSTA RICA</collectingCountry>
, 
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:
</emphasis>
1♁, 
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, 
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</quantity>
, 
<date box="[841,937,367,394]" bridgedPair="/" pageId="15" pageNumber="593" value="1995-11">
<collectingDate box="[841,937,367,394]" bridgedPair="/" pageId="15" pageNumber="593" value="1995-11">XI/1995</collectingDate>
</date>
, 
<collectorName box="[952,1091,367,393]" pageId="15" pageNumber="593">G. Gallardo</collectorName>
, L_S_385000_578100 #6826. CRI002 429864 (
<collectionCode box="[347,423,403,429]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="15" pageNumber="593">INBio</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="2007-05-19" collectingDateMax="2007-05-20" collectingDateMin="2007-05-19" collectionCode="INBio" collectorName="J. A. Azofeifa &amp; J. Moutero" country="Costa Rica" elevation="80" location="Banano Lodge" municipality="Valle La Estrella" pageId="15" pageNumber="593" specimenCount="1" specimenCount-female="1" stateProvince="Limon" typeStatus="paratype">
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, 
<collectingMunicipality box="[492,683,403,430]" pageId="15" pageNumber="593">Valle La Estrella</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E:202D2B94AE2FFFC4FD2DFE2A542BFE25" box="[695,862,403,430]" country="Costa Rica" municipality="Valle La Estrella" name="Banano Lodge" pageId="15" pageNumber="593" stateProvince="Limon">Banano Lodge</location>
, 
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<elevation box="[874,927,403,429]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="593" unit="m" value="80.0">80m</elevation>
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, 
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<collectingDate box="[939,1104,403,429]" pageId="15" pageNumber="593" value="2007-05-19" valueMax="2007-05-20" valueMin="2007-05-19">19–20/V/2007</collectingDate>
</date>
, 
<collectorName box="[1115,1270,403,430]" pageId="15" pageNumber="593">J. A. Azofeifa</collectorName>
&amp; 
<collectorName box="[1307,1432,404,430]" pageId="15" pageNumber="593">J. Moutero</collectorName>
, T. P., at lights, L N 200889 639300 #91343, INB0004080803 (
<collectionCode box="[859,935,439,465]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="15" pageNumber="593">INBio</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="2004-09-16" collectionCode="INBio" collectorName="R. V. S. Rio Piro &amp; Y. Cardenas &amp; M. Moraga &amp; D. Briceno &amp; Gamboa" country="Costa Rica" elevation="200" location="Finca Catalino" municipality="Golfito" pageId="15" pageNumber="593" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis bold="true" box="[955,1101,439,465]" pageId="15" pageNumber="593">
<collectingRegion box="[955,1096,439,465]" country="Costa Rica" name="Puntarenas" pageId="15" pageNumber="593">Puntarenas</collectingRegion>
:
</emphasis>
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, 
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, 
<collectingMunicipality box="[1348,1432,439,465]" pageId="15" pageNumber="593">Golfito</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E:202D2B94AE2FFFC4FF0DFE655634FE7D" box="[151,321,475,501]" country="Costa Rica" municipality="Golfito" name="Finca Catalino" pageId="15" pageNumber="593" stateProvince="Puntarenas">Finca Catalino</location>
, 
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<elevation box="[334,401,475,501]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="593" unit="m" value="200.0">200m</elevation>
</quantity>
, 
<date box="[414,546,475,501]" pageId="15" pageNumber="593" value="2004-09-16">
<collectingDate box="[414,546,475,501]" pageId="15" pageNumber="593" value="2004-09-16">16/IX/2004</collectingDate>
</date>
, 
<collectorName box="[559,699,475,502]" pageId="15" pageNumber="593">Y. Cardenas</collectorName>
, 
<collectorName box="[712,844,476,502]" pageId="15" pageNumber="593">M. Moraga</collectorName>
, 
<collectorName box="[857,983,475,502]" pageId="15" pageNumber="593">D. Briceño</collectorName>
&amp; B. 
<collectorName box="[1058,1156,475,501]" pageId="15" pageNumber="593">Gamboa</collectorName>
, at lights, L_S_264550_ 535590 #78215, INB0003880887 (
<collectionCode box="[543,619,511,537]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="15" pageNumber="593">INBio</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1981-01-05" collectingDateMax="1981-01-11" collectingDateMin="1981-01-05" collectionCode="INBio" collectorName="D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" county="Sirena" location="Osa Penin." municipality="Corcovado Nat. Pk." pageId="15" pageNumber="593" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount box="[638,673,511,537]" pageId="15" pageNumber="593" type="female">1♀</specimenCount>
, 
<collectingCounty box="[687,761,511,537]" pageId="15" pageNumber="593">Sirena</collectingCounty>
, 
<collectingMunicipality box="[773,996,511,538]" pageId="15" pageNumber="593">Corcovado Nat. Pk.</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E:202D2B94AE2FFFC4FC75FE41531EFD91" box="[1007,1131,511,537]" country="Costa Rica" county="Sirena" municipality="Corcovado Nat. Pk." name="Osa Penin." pageId="15" pageNumber="593" stateProvince="Puntarenas">Osa Penin.</location>
, 
<date box="[1142,1281,511,538]" pageId="15" pageNumber="593" value="1981-01-05" valueMax="1981-01-11" valueMin="1981-01-05">
<collectingDate box="[1142,1281,511,538]" pageId="15" pageNumber="593" value="1981-01-05" valueMax="1981-01-11" valueMin="1981-01-05">5–11/I/1981</collectingDate>
</date>
, 
<collectorName box="[1292,1436,512,538]" pageId="15" pageNumber="593">D. H. Janzen</collectorName>
&amp; 
<collectorName box="[182,344,547,574]" pageId="15" pageNumber="593">W. Hallwachs</collectorName>
, CRI001 714731 (
<collectionCode box="[559,635,547,573]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="15" pageNumber="593">INBio</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1992-04" collectionCode="JEE" collectorName="P. N. Corcovado &amp; G. Rodriguez &amp; Costa Rica &amp; Fernandes" country="Costa Rica" elevation="50" location="Est. Sirena" pageId="15" pageNumber="593" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount box="[657,692,547,573]" pageId="15" pageNumber="593" type="female">1♀</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E:202D2B94AE2FFFC4FD5FFD9A5431FDB5" box="[709,836,547,574]" country="Costa Rica" name="Est. Sirena" pageId="15" pageNumber="593" stateProvince="Puntarenas">Est. Sirena</location>
, 
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<elevation box="[851,945,547,573]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="15" pageNumber="593" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</elevation>
</quantity>
, 
<collectorName box="[959,1152,547,574]" pageId="15" pageNumber="593">P. N. Corcovado</collectorName>
, 
<date box="[1166,1268,547,573]" bridgedPair="-" pageId="15" pageNumber="593" value="1992-04">
<collectingDate box="[1166,1268,547,573]" bridgedPair="-" pageId="15" pageNumber="593" value="1992-04">IV–1992</collectingDate>
</date>
, 
<collectorName box="[1282,1437,547,573]" pageId="15" pageNumber="593">G. Rodriguez</collectorName>
(
<collectorName box="[159,286,583,609]" pageId="15" pageNumber="593">Costa Rica</collectorName>
, INBIO CRI000, 545535; sp 195, 
<collectorName box="[676,791,583,609]" pageId="15" pageNumber="593">Fernandes</collectorName>
JAM) (
<collectionCode box="[880,931,583,609]" pageId="15" pageNumber="593">JEE</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1979-01-02" collectionCode="DOE" collectorName="Henk Wolda" country="Panama" location="Miramar" longitude="-82.25" pageId="15" pageNumber="593" specimenCount="1" specimenCount-female="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<emphasis bold="true" box="[952,1275,583,609]" pageId="15" pageNumber="593">
<collectingCountry box="[952,1076,583,609]" name="Panama" pageId="15" pageNumber="593">PANAMA</collectingCountry>
, 
<collectingRegion box="[1089,1269,583,609]" country="Panama" name="Bocas del Toro" pageId="15" pageNumber="593">Bocas del Toro</collectingRegion>
:
</emphasis>
<specimenCount box="[1283,1318,583,609]" pageId="15" pageNumber="593" type="female">1♀</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E:202D2B94AE2FFFC4FAAEFDF952EDFDE9" box="[1332,1432,583,609]" country="Panama" longitude="-82.25" name="Miramar" pageId="15" pageNumber="593" stateProvince="Bocas del Toro">Miramar</location>
, 9ºN; 
<geoCoordinate box="[202,302,619,646]" degrees="82" direction="west" minutes="15" orientation="longitude" pageId="15" pageNumber="593" precision="925" value="-82.25">82º15’W</geoCoordinate>
, 
<date box="[314,412,619,645]" pageId="15" pageNumber="593" value="1979-01-02">
<collectingDate box="[314,412,619,645]" pageId="15" pageNumber="593" value="1979-01-02">2/I/1979</collectingDate>
</date>
, 
<collectorName box="[423,562,619,645]" pageId="15" pageNumber="593">Henk Wolda</collectorName>
(
<collectionCode box="[577,641,619,645]" pageId="15" pageNumber="593">DOE</collectionCode>
)
</materialsCitation>
; 
<materialsCitation box="[660,1348,619,648]" collectingDate="1979-01-28" collectionCode="DOE" collectorName="Henk Wolda" country="Panama" latitude="9.0" location="Miramar" longLatPrecision="0" longitude="-82.25" pageId="15" pageNumber="593" specimenCount="1" stateProvince="Bocas del Toro" typeStatus="paratype">
1♁, 
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2FFFC4FF0DFF2953C0F81E:202D2B94AE2FFFC4FD5FFDD5545CFD0D" box="[709,809,619,645]" country="Panama" latitude="9.0" longLatPrecision="0" longitude="-82.25" name="Miramar" pageId="15" pageNumber="593" stateProvince="Bocas del Toro">Miramar</location>
, 
<geoCoordinate box="[820,865,619,645]" degrees="9" direction="north" orientation="latitude" pageId="15" pageNumber="593" precision="55555" value="9.0">9°N</geoCoordinate>
<geoCoordinate box="[872,975,619,646]" degrees="82" direction="west" minutes="15" orientation="longitude" pageId="15" pageNumber="593" precision="925" value="-82.25">82°15’W</geoCoordinate>
, 
<date box="[987,1110,619,645]" pageId="15" pageNumber="593" value="1979-01-28">
<collectingDate box="[987,1110,619,645]" pageId="15" pageNumber="593" value="1979-01-28">28–I–1979</collectingDate>
</date>
, 
<collectorName box="[1122,1261,619,645]" pageId="15" pageNumber="593">Henk Wolda</collectorName>
(
<collectionCode box="[1277,1341,619,645]" pageId="15" pageNumber="593">DOE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="593" type="description">
<paragraph blockId="15.[151,1437,259,1942]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[199,374,655,681]" pageId="15" pageNumber="593">Measurements</emphasis>
(n= 8): antennomeres length: 1st: 
<quantity box="[799,958,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="8.8" metricValueMin="8.2" pageId="15" pageNumber="593" unit="mm" value="0.85" valueMax="0.88" valueMin="0.82">0.82–0.88mm</quantity>
; 2nd: 
<quantity box="[1038,1197,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" metricValueMax="1.57" metricValueMin="1.47" pageId="15" pageNumber="593" unit="mm" value="1.52" valueMax="1.57" valueMin="1.47">1.47–1.57mm</quantity>
; 3rd: 
<quantity box="[1273,1432,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" metricValueMax="2.02" metricValueMin="1.52" pageId="15" pageNumber="593" unit="mm" value="1.77" valueMax="2.02" valueMin="1.52">1.52–2.02mm</quantity>
; 4th: 
<quantity box="[206,364,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="3.995" metricValueMax="4.26" metricValueMin="3.73" pageId="15" pageNumber="593" unit="mm" value="3.995" valueMax="4.26" valueMin="3.73">3.73–4.26mm</quantity>
; 5th: 
<quantity box="[434,593,691,718]" metricMagnitude="-3" metricUnit="m" metricValue="4.095" metricValueMax="4.35" metricValueMin="3.84" pageId="15" pageNumber="593" unit="mm" value="4.095" valueMax="4.35" valueMin="3.84">3.84–4.35mm</quantity>
; head length: 
<quantity box="[760,920,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.79" metricValueMin="1.4" pageId="15" pageNumber="593" unit="mm" value="1.595" valueMax="1.79" valueMin="1.4">1.40–1.79mm</quantity>
; head width: 
<quantity box="[1080,1239,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.685" metricValueMax="3.1" metricValueMin="2.27" pageId="15" pageNumber="593" unit="mm" value="2.685" valueMax="3.1" valueMin="2.27">2.27–3.10mm</quantity>
; pronotal length: 
<quantity box="[151,310,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="3.24" metricValueMax="3.36" metricValueMin="3.12" pageId="15" pageNumber="593" unit="mm" value="3.24" valueMax="3.36" valueMin="3.12">3.12–3.36mm</quantity>
; pronotal width: 
<quantity box="[508,682,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.68" metricValueMax="10.16" metricValueMin="7.2" pageId="15" pageNumber="593" unit="mm" value="8.68" valueMax="10.16" valueMin="7.2">7.20–10.16mm</quantity>
; scutellum length: 
<quantity box="[901,1060,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="5.88" metricValueMax="6.22" metricValueMin="5.54" pageId="15" pageNumber="593" unit="mm" value="5.88" valueMax="6.22" valueMin="5.54">5.54–6.22mm</quantity>
; scutellum width: 
<quantity box="[1273,1432,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.46" metricValueMax="4.69" metricValueMin="4.23" pageId="15" pageNumber="593" unit="mm" value="4.46" valueMax="4.69" valueMin="4.23">4.23–4.69mm</quantity>
; abdominal width: 
<quantity box="[355,528,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="10.08" metricValueMin="7.02" pageId="15" pageNumber="593" unit="mm" value="8.55" valueMax="10.08" valueMin="7.02">7.02–10.08mm</quantity>
; total length: 
<quantity box="[681,868,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" metricValueMax="1.738" metricValueMin="1.222" pageId="15" pageNumber="593" unit="mm" value="14.8" valueMax="17.38" valueMin="12.22">12.22–17.38mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="593" type="diagnosis">
<paragraph blockId="15.[151,1437,259,1942]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[199,324,799,825]" pageId="15" pageNumber="593">Diagnosis:</emphasis>
body dorsally green (
<figureCitation box="[568,663,799,825]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Antennae light-brown (
<figureCitation box="[943,1038,799,825]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Anterolateral margin of pronotum yellow (
<figureCitation box="[247,351,835,861]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Black minute longitudinal callosity adjacent to humeral angles (
<figureCitation box="[1127,1231,835,861]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Humeral angles concolorous with dorsum, straight, short, less than the width of an eye (
<figureCitation box="[974,1074,871,897]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Last connexival segment with only posterior margin black (
<figureCitation box="[495,600,907,933]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Ventral surface: green with median area yellow, without dark stripes, intersegmental areas of the abdomen slightly darker than the rest (
<figureCitation box="[892,985,943,969]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 F</figureCitation>
). Anterior bifurcation of the metasternal process with arms very short, wide, swollen, apices rounded (
<figureCitation box="[838,930,979,1005]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 F</figureCitation>
). Male genitalia: pygophore trapezoidal (
<figureCitation captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 A, B</figureCitation>
). Superior process of the genital cup subtriangular, surface rugose, slightly concave (
<figureCitation box="[1172,1301,1015,1042]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C, D</figureCitation>
). Parameres with anterior lobe long and subtriangular (
<figureCitation box="[627,757,1051,1078]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C, D</figureCitation>
); dorsal margin shallowly excavated (
<figureCitation box="[1185,1315,1051,1078]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C, D</figureCitation>
); posterior lobe short and triangular (
<figureCitation box="[439,567,1087,1114]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C, D</figureCitation>
). Proctiger laterally strongly excavated, separated by feeble low carina (
<figureCitation captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C</figureCitation>
); posterior face rounded to slightly elliptic (
<figureCitation box="[661,755,1123,1150]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C</figureCitation>
). Ventral rim with median notch shallow and wide (
<figureCitation box="[1328,1421,1123,1150]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 B</figureCitation>
); expansions slightly developed (
<figureCitation box="[505,600,1159,1186]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 B</figureCitation>
). Female genitalia (
<figureCitation box="[822,917,1159,1186]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 E</figureCitation>
): valvifers VIII punctured with wide and deep V-shaped excavation; inner margin overlapping each other.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="593" type="description">
<paragraph blockId="15.[151,1437,259,1942]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[199,419,1231,1257]" pageId="15" pageNumber="593">Description: head:</emphasis>
clypeus and jugae, slightly ridged. Bucculae subtriangular, completely harboring first labial segment. 
<emphasis bold="true" box="[257,355,1267,1293]" pageId="15" pageNumber="593">Thorax:</emphasis>
dorsal surface: pronotum with punctures concolorous, sparse, mostly present on posterior ¾ (
<figureCitation captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Anterior margin of pronotum with row of sparse punctures (
<figureCitation box="[884,980,1303,1329]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Scutellum with punctures concolorous, sparse on anterior half, grouped much tighter on posterior half (
<figureCitation box="[860,955,1339,1365]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Corium with concolorous punctures (
<figureCitation captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
); membrane transparent, unpigmented (
<figureCitation box="[637,731,1375,1401]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). Ventral surface: prothorax–mesothorax irregularly punctured, punctures concolorous or dark (
<figureCitation box="[512,607,1411,1437]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 F</figureCitation>
). Peritreme ruga-like, reaching 4/5 of the distance between ostiole of the scent gland and lateral margin of metapleuron (
<figureCitation box="[680,772,1447,1473]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 F</figureCitation>
). Metasternal process with anterior arms slightly divergent, anterior bifurcation receiving only part of fourth rostral segment (
<figureCitation box="[875,965,1483,1509]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 F</figureCitation>
). 
<emphasis bold="true" box="[984,1110,1483,1509]" pageId="15" pageNumber="593">Abdomen:</emphasis>
dorsal surface: posterolateral angles of connexival segments with apices black (
<figureCitation box="[714,810,1519,1545]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
). 
<emphasis bold="true" box="[831,1011,1519,1545]" pageId="15" pageNumber="593">Male genitalia:</emphasis>
dorsal rim with brown large spot that extends to dorsal surface (
<figureCitation box="[445,541,1555,1582]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 A</figureCitation>
); shallowly concave (
<figureCitation box="[785,881,1555,1582]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 A</figureCitation>
). Posterolateral angles well developed (
<figureCitation box="[1327,1423,1555,1582]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 A</figureCitation>
); rounded, slightly swollen (
<figureCitation box="[446,540,1591,1618]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 A</figureCitation>
). Ventral rim with expansions short, rounded, slightly swollen (
<figureCitation box="[1238,1329,1591,1618]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 B</figureCitation>
). 
<emphasis bold="true" pageId="15" pageNumber="593">Female genitalia</emphasis>
(
<figureCitation box="[272,370,1627,1654]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 E</figureCitation>
): valvifers VIII convex reaching base of valvifers IX, excavation exposing part of valvulae IX. Laterotergites VIII acute, outer lateral margin arched, sinuous, free distal spinose part about 1/4 of its length. Valvifers IX gently punctured, trapezoidal. Laterotergites IX acute clearly surpassing sclerite uniting laterotergites VIII.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="593" type="diagnosis">
<paragraph blockId="15.[151,1437,259,1942]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[199,466,1771,1797]" pageId="15" pageNumber="593">Differential diagnosis:</emphasis>
among the species in this article, 
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[858,1091,1771,1797]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="593" phylum="Arthropoda" rank="species" species="holochlorata" status="sp. n.">
<emphasis box="[858,1091,1771,1797]" italics="true" pageId="15" pageNumber="593">Edessa holochlorata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1100,1165,1771,1797]" pageId="15" pageNumber="593">
<taxonomicNameLabel box="[1100,1165,1771,1797]" pageId="15" pageNumber="593" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
stands out for having a completely green corium (
<figureCitation box="[449,545,1807,1833]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">Fig. 9 C</figureCitation>
), unlike the brown corium of all others (
<figureCitation box="[1004,1179,1807,1834]" captionStart="FIGURE 8" captionStartId="17.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@17.[203,1385,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. A–H. Dorsal and ventral surface: A–D, Edessa bituberculata sp. n.; B–E, Edessa bivenulata sp. n.; C–F, Edessa enargocelida sp. n.; G–H, Edessa fuscolimbata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906776" httpUri="https://zenodo.org/record/7906776/files/figure.png" pageId="15" pageNumber="593">Figs. 8 A, B, C</figureCitation>
; 
<figureCitation box="[1191,1267,1807,1833]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. A–F. Dorsal and ventral surface: A–D, Edessa fuscopunctata sp. n.; B–E, Edessa helvoalata sp. n.; C–F, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="15" pageNumber="593">9 A, B</figureCitation>
). Furthermore, male and female genitalia also present distinctive characteristics like the shape of the parameres and overlapping valvifers VIII (
<figureCitation box="[319,446,1879,1906]" captionStart="FIGURE 7" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[195,1393,179,1917]" captionTargetId="figure-20@16.[195,1393,179,1917]" captionTargetPageId="16" captionText="FIGURE 7.A–E.Edessa holochlorata sp. n.:A–D, pygophore in dorsal, ventral, posterior andposterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906772" httpUri="https://zenodo.org/record/7906772/files/figure.png" pageId="15" pageNumber="593">Fig. 7 C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,1205,1915,1942]" pageId="15" pageNumber="593" type="distribution">
<paragraph blockId="15.[151,1437,259,1942]" box="[199,1205,1915,1942]" pageId="15" pageNumber="593">
<emphasis bold="true" box="[199,345,1915,1941]" pageId="15" pageNumber="593">Distribution</emphasis>
(
<figureCitation box="[361,467,1915,1941]" captionStart="FIGURE 10" captionStartId="19.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@19.[203,1385,181,1919]" captionTargetPageId="19" captionText="FIGURE 10. A–C. Distribution map of the species: A, Edessa bituberculata sp. n., Edessa fuscolimbata sp. n., Edessa fuscopunctatasp.n.;B,Edessabivenulatasp.n.,Edessaenargocelidasp.n.,Edessaholochloratasp.n.;C,Edessahelvoalatasp.n." figureDoi="http://doi.org/10.5281/zenodo.7906784" httpUri="https://zenodo.org/record/7906784/files/figure.png" pageId="15" pageNumber="593">Fig. 10 B</figureCitation>
): 
<collectingCountry box="[492,658,1915,1941]" name="Costa Rica" pageId="15" pageNumber="593">COSTA RICA</collectingCountry>
: 
<collectingRegion box="[669,747,1915,1941]" country="Costa Rica" name="Limon" pageId="15" pageNumber="593">Limón</collectingRegion>
, 
<collectingRegion box="[758,885,1916,1942]" country="Costa Rica" name="Puntarenas" pageId="15" pageNumber="593">Puntarenas</collectingRegion>
; 
<collectingCountry box="[895,1018,1915,1941]" name="Panama" pageId="15" pageNumber="593">PANAMA</collectingCountry>
: 
<collectingRegion box="[1029,1201,1915,1942]" country="Panama" name="Bocas del Toro" pageId="15" pageNumber="593">Bocas del Toro</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>