257 lines
28 KiB
XML
257 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.999.52831" ID-GBIF-Dataset="11e8fa4e-ab97-47a5-a8d1-a9748ce5382c" ID-PMC="PMC7723878" ID-Pensoft-Pub="1313-2970-999-147" ID-Pensoft-UUID="69960D48D282536EAC45F3D50DD62BF3" ID-PubMed="33343215" ID-ZooBank="A1116A782C1141FBA8B315AB7369A942" ModsDocID="1313-2970-999-147" checkinTime="1607107216301" checkinUser="pensoft" docAuthor="Kato, Daichi, Watanabe, Kozo & Kolcsar, Levente-Peter" docDate="2020" docId="5A28DC98C7A25BB299CC80712337C2EE" docLanguage="en" docName="ZooKeys 999: 147-163" docOrigin="ZooKeys 999" docSource="http://dx.doi.org/10.3897/zookeys.999.52831" docTitle="Ulomorpha nigricolor Alexander 1924" docType="treatment" docVersion="4" id="69960D48D282536EAC45F3D50DD62BF3" lastPageNumber="147" masterDocId="69960D48D282536EAC45F3D50DD62BF3" masterDocTitle="Synopsis of the genus Ulomorpha Osten Sacken, 1869 (Diptera, Limoniidae) in Japan" masterLastPageNumber="163" masterPageNumber="147" pageNumber="147" updateTime="1668169993919" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Synopsis of the genus Ulomorpha Osten Sacken, 1869 (Diptera, Limoniidae) in Japan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kato, Daichi</mods:namePart>
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<mods:affiliation>Echigo-Matsunoyama Museum of Natural Sciences, ' Kyororo', 1712 - 2 Matsunoyama, Tokamachi, 942 - 1411, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Watanabe, Kozo</mods:namePart>
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<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7062-595X</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kolcsar, Levente-Peter</mods:namePart>
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<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7784-2386</mods:nameIdentifier>
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<mods:nameIdentifier type="email">kolcsar.peter@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>999</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>147</mods:start>
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<mods:end>163</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.999.52831</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.999.52831</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-999-147</mods:identifier>
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<mods:identifier type="ZooBank">A1116A782C1141FBA8B315AB7369A942</mods:identifier>
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<mods:identifier type="Pensoft-UUID">69960D48D282536EAC45F3D50DD62BF3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="170678111" LSID="urn:lsid:plazi:treatment:5A28DC98C7A25BB299CC80712337C2EE" httpUri="http://treatment.plazi.org/id/5A28DC98C7A25BB299CC80712337C2EE" lastPageNumber="147" pageId="0" pageNumber="147">
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<subSubSection pageId="0" pageNumber="147" type="nomenclature">
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<paragraph pageId="0" pageNumber="147">
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<taxonomicName LSID="5A28DC98-C7A2-5BB2-99CC-80712337C2EE" authority="Alexander, 1924" authorityName="Alexander" authorityYear="1924" class="Insecta" family="Limoniidae" genus="Ulomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulomorpha nigricolor" order="Diptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="nigricolor">Ulomorpha nigricolor Alexander, 1924</taxonomicName>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ulomorpha. A U. amamiana Kato & Kolcsar, sp. nov. (orange), U. nigricolor Alexander, 1924 (blue), U. polytricha Alexander, 1930 (green) B U. longipenis Kato & Kolcsar, sp. nov. (red)." figureDoi="10.3897/zookeys.999.52831.figure5" httpUri="https://binary.pensoft.net/fig/481464" pageId="0" pageNumber="147">, 5A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="reference_group">
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<paragraph pageId="0" pageNumber="147">
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<taxonomicName authorityName="Alexander" authorityYear="1924" class="Insecta" family="Limoniidae" genus="Ulomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulomorpha nigricolor" order="Diptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="nigricolor">Ulomorpha nigricolor</taxonomicName>
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<bibRefCitation DOI="https://doi.org/10.1080/00222932508633181" author="Alexander, CP" journalOrPublisher="Memoirs, Cornell University Agricultural Experiment Station" pageId="0" pageNumber="147" refId="B4" refString="Alexander, CP, 1924. New or little-known Tipulidae (Diptera). XXVI. Palaearctic species. Annals and Magazine of Natural History (Series 9) 15(9): 65-81. https://doi.org/10.1080/00222932508633181" title="New or little-known Tipulidae (Diptera). XXVI. Palaearctic species. Annals and Magazine of Natural History (Series 9) 15 (9): 65 - 81." url="https://doi.org/10.1080/00222932508633181" year="1924">Alexander 1924</bibRefCitation>
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: 75 (type locality: Japan, Honshu I., Gunma or Fukushima, Lake Oze-numa);
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<bibRefCitation author="Alexander, CP" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="147" pagination="71 - 83" refId="B8" refString="Alexander, CP, 1953a. The insect fauna of Mt. Ishizuchi and Omogo Valley, Iyo, Japan. The Tipulidae (Diptera). Transactions of the Shikoku Entomological Society 3: 71 - 83" title="The insect fauna of Mt. Ishizuchi and Omogo Valley, Iyo, Japan. The Tipulidae (Diptera)." volume="3" year="1953 a">Alexander 1953a</bibRefCitation>
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: 82;
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<bibRefCitation author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="147" pagination="21 - 75" refId="B9" refString="Alexander, CP, 1953b. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 b">Alexander 1953b</bibRefCitation>
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: 68;
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<bibRefCitation author="Nakamura, T" editor="Nakamura, T" journalOrPublisher="Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza). Touka Shobo, Fukuoka" pageId="0" pageNumber="147" pagination="9 - 53" refId="B18" refString="Nakamura, T, 2014. Family Limoniidae. In: Nakamura, T, Saigusa, T, Suwa, M, Eds., Catalogue of the Insects of Japan. Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza). Touka Shobo, Fukuoka: 9 - 53" title="Family Limoniidae." volumeTitle="Catalogue of the Insects of Japan." year="2014">Nakamura 2014</bibRefCitation>
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: 20;
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<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza). Touka Shobo, Fukuoka" pageId="0" pageNumber="147" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B19" refString="Oosterbroek, P, 2020. Catalogue of the Craneflies of the World (Diptera, Tipuloidea: Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae). https://ccw.naturalis.nl/index.php" title="Catalogue of the Craneflies of the World (Diptera, Tipuloidea: Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae)." url="https://ccw.naturalis.nl/index.php" year="2020">Oosterbroek 2020</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="materials_examined">
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<paragraph pageId="0" pageNumber="147">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="147">Holotype</emphasis>
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♂. Japan, Honshu, Lake Ozenuma, on boundary between Iwashiro-no-kuni and Kotsuke-no-kuni (between Fukuoka and Gunma); altitude 5460 feet;
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<geoCoordinate degrees="36" direction="north" minutes="55.62" orientation="latitude" precision="9" value="36.927">36°55.62'N</geoCoordinate>
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,
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<geoCoordinate degrees="139" direction="east" minutes="18.23" orientation="longitude" precision="9" value="139.30383">139°18.23'E</geoCoordinate>
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(rough coordinate); 26 Jul. 1923; T. Esaki leg. (USNM).
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">
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<emphasis italics="true" pageId="0" pageNumber="147">Non-types</emphasis>
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.
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</emphasis>
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Japan: [Honshu] • 1♀; Aomori, Nishimeya-mura; alt. 275 m;
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<geoCoordinate degrees="40" direction="north" minutes="31.39" orientation="latitude" precision="9" value="40.523167">40°31.39'N</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="13.93" orientation="longitude" precision="9" value="140.23216">140°13.93'E</geoCoordinate>
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; 23. Jun. 2012; D. Kato leg. (BLKU) • 1♂; same data as previous except 24. Jun. 2012 • 2♂, 1♀; Aomori, Nishimeya-mura, Kawaratai, Shirakami Nature Observation Garden; alt. 225 m;
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<geoCoordinate degrees="40" direction="north" minutes="31.13" orientation="latitude" precision="9" value="40.518833">40°31.13'N</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="12.89" orientation="longitude" precision="9" value="140.21483">140°12.89'E</geoCoordinate>
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; 21 Jun. 2013; D. Kato leg. (BLKU) • 1♂; Aomori, Nakadomari-machi,
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<normalizedToken originalValue="Ôsawanai">Osawanai</normalizedToken>
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,
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<normalizedToken originalValue="Ôsawanai">Osawanai</normalizedToken>
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Pond; alt. 35 m;
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<geoCoordinate degrees="40" direction="north" minutes="56.78" orientation="latitude" precision="9" value="40.946335">40°56.78'N</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="27.74" orientation="longitude" precision="9" value="140.46233">140°27.74'E</geoCoordinate>
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; 15 May 2014; D. Kato leg. (BLKU) • 1♂; same data as previous except 24 May 2014 • 1♂; Aomori, Towada-shi, Okuse, Tsutanuma Path; alt. 470 m;
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<geoCoordinate degrees="40" direction="north" minutes="35.45" orientation="latitude" precision="9" value="40.59083">40°35.45'N</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="57.42" orientation="longitude" precision="9" value="140.957">140°57.42'E</geoCoordinate>
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; 23 May 2014; D. Kato leg. (BLKU) • 1♀; Iwate, Hachimantai-shi, near
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<normalizedToken originalValue="Tȏshichi">Tȏshichi</normalizedToken>
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Spa; alt. 1340 m;
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<geoCoordinate degrees="39" direction="north" minutes="56.55" orientation="latitude" precision="9" value="39.9425">39°56.55'N</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="52.08" orientation="longitude" precision="9" value="140.868">140°52.08'E</geoCoordinate>
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; 28 Aug. 2014; D. Kato leg. (BLKU) • 1♂; Niigata, Echigo, Iwafune, Mt Zao;
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<geoCoordinate degrees="38" direction="north" minutes="4.64" orientation="latitude" precision="9" value="38.07733">38°4.64'N</geoCoordinate>
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,
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<geoCoordinate degrees="139" direction="east" minutes="28.57" orientation="longitude" precision="9" value="139.47617">139°28.57'E</geoCoordinate>
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(rough coordinate, altitude unknown); 3 May 1955; K. Baba leg. (USNM) • 1♀; Gifu, Mino, Sakauchi;
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<geoCoordinate degrees="35" direction="north" minutes="36.41" orientation="latitude" precision="9" value="35.606834">35°36.41'N</geoCoordinate>
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,
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<geoCoordinate degrees="136" direction="east" minutes="22.96" orientation="longitude" precision="9" value="136.38266">136°22.96'E</geoCoordinate>
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(rough coordinate, altitude unknown); 9 Jun. 1957; Mishima leg. (USNM) • 5♂, 4♀; Okayama, Maniwa-shi, Hiruzen-Shimotokuyama; alt. 780 m;
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<geoCoordinate degrees="35" direction="north" minutes="19.76" orientation="latitude" precision="9" value="35.329334">35°19.76'N</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="35.84" orientation="longitude" precision="9" value="133.59734">133°35.84'E</geoCoordinate>
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; 17 May 2015; D. Kato leg. (BLKU) • 2♂, 2♀; Okayama, Maniwa-shi, Hiruzen-Kamifukuda, Nawashirodani-gawa River; alt. 600 m;
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<geoCoordinate degrees="35" direction="north" minutes="19.19" orientation="latitude" precision="9" value="35.31983">35°19.19'N</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="36.49" orientation="longitude" precision="9" value="133.60817">133°36.49'E</geoCoordinate>
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; 29 Apr. 2016; D. Kato leg. (BLKU) • 1♂; Hiroshima,
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<normalizedToken originalValue="Akiôta-chô">Akiota-cho</normalizedToken>
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,
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<normalizedToken originalValue="Yokogô">Yokogo</normalizedToken>
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; alt. 890 m;
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<geoCoordinate degrees="34" direction="north" minutes="35.65" orientation="latitude" precision="9" value="34.594166">34°35.65'N</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="8.7" orientation="longitude" precision="92" value="132.145">132°8.7'E</geoCoordinate>
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; 18 May 2015; D. Kato leg. (BLKU). [Shikoku] • 1♂; Ehime,
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<normalizedToken originalValue="Kumakôgen-chô">Kumakogen-cho</normalizedToken>
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, Nishidani, near Prefectural road 328; alt. 1430 m; 33°34'N, 132°56.2'E; 17 Jun. 2019; D. Kato leg. (BLKU) • 2♂, 2♀; same data as previous except alt. 1387 m;
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<geoCoordinate degrees="33" direction="north" minutes="33.89" orientation="latitude" precision="9" value="33.564835">33°33.89'N</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="56.1" orientation="longitude" precision="92" value="132.935">132°56.1'E</geoCoordinate>
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; L.-P.
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<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
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leg. (CKLP) • 4♂; Ehime,
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<normalizedToken originalValue="Saijô-shi">Saijo-shi</normalizedToken>
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, Nishinokawa-Tei, Mt Ishizuchi-san; alt. 1530 m;
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<geoCoordinate degrees="33" direction="north" minutes="45.29" orientation="latitude" precision="9" value="33.754833">33°45.29'N</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="9.19" orientation="longitude" precision="9" value="133.15317">133°9.19'E</geoCoordinate>
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; 16 Jun. 2019; D. Kato leg. (BLKU) • 2♂; same data as previous except alt. 1480 m;
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<geoCoordinate degrees="33" direction="north" minutes="45.3" orientation="latitude" precision="92" value="33.755">33°45.3'N</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="9.23" orientation="longitude" precision="9" value="133.15384">133°9.23'E</geoCoordinate>
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; 16 May 2019; L.-P.
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<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
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leg. (CKLP) • 1♂, same data as previous except 16 Jun. 2019. [Kyushu] • 3♂; Miyazaki,
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<normalizedToken originalValue="Takachiho-chô">Takachiho-cho</normalizedToken>
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, Gokasho, Mt Sobo-san, near Kitadani trailhead; alt. 1150 m;
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<geoCoordinate degrees="32" direction="north" minutes="49.36" orientation="latitude" precision="9" value="32.822666">32°49.36'N</geoCoordinate>
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,
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<geoCoordinate degrees="131" direction="east" minutes="19.64" orientation="longitude" precision="9" value="131.32733">131°19.64'E</geoCoordinate>
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; 22 May 2019; D. Kato leg. (BLKU) • 5♂, 1♀; same data as previous except alt. 1182 m;
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<geoCoordinate degrees="32" direction="north" minutes="49.36" orientation="latitude" precision="9" value="32.822666">32°49.36'N</geoCoordinate>
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,
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<geoCoordinate degrees="131" direction="east" minutes="19.64" orientation="longitude" precision="9" value="131.32733">131°19.64'E</geoCoordinate>
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; L.-P.
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<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
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leg. (CKLP).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="diagnosis">
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<paragraph pageId="0" pageNumber="147">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="147">Body blackish. Vertex sparsely pruinose, anterior part largely polished. Flagellomeres oval on basal 2-4 segments; ventral sides with pubescences on basal 2-6 segments. Prescutum polished, with narrow longitudinal line of pruinosity at middle. Wing brownish tinged, with oval, dark-brown stigma and dark spot or seam each on Rs origin, anterior part of cord, crossvein m-cu, and outer end of cell d; dark spot at tip of A2 sometimes present. Halter yellowish. Interbase with outer lobe as long as medial lobe in dorsal view; medial lobe medial to base of outer lobe about 3.5 times as long as wide. Aedeagus with rod-shaped distal part, twice as long as wide and weakly curved ventrally.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="description">
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<paragraph pageId="0" pageNumber="147">Description.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">Male</emphasis>
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. Body length 5.5-10.1 mm, wing length 5.3-10.0 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">
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<emphasis italics="true" pageId="0" pageNumber="147">Head</emphasis>
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:
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</emphasis>
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subnitidous black, sparsely dusted with gray pruinosity; vertex with brighter gray pruinosity at anterior end; anterior part of vertex largely shiny except anterior end. Rostrum and mouthparts dark brown. Antenna with scape and pedicel dark brown; flagellum dusky yellow to dark brown; basal 2-4 flagellomeres oval; basal 2-6 segments covered with pubescences ventrally.
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">
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<emphasis italics="true" pageId="0" pageNumber="147">Thorax</emphasis>
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:
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</emphasis>
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subnitidous black, sparsely dusted with brownish pruinosity; polished and pruinosity absent on prescutum and most of scutal lobe; postpronotum sometimes yellowish; prescutum with narrow longitudinal line of pruinosity at middle in whole length, sometimes anterior part of this line indistinct. Wing (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3B, C</figureCitation>
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) tinged with brown; basal and costal regions proximal to cord yellowish; stigma oval, dark brown, faint in outline; veins dark brown, yellowish on Sc and basal part of wing; dark, wide seam on fork of Rs to crossvein r-m; narrow, dark seam on each of crossvein m-cu and outer end of cell d, but one on latter sometimes indistinct; dark small spot at Rs origin and sometimes with additional one at tip of A2 (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3C</figureCitation>
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), this anal spot usually present in specimens from southern part of Japan. Halter yellow to pale yellow. Legs mainly yellow to dusky yellow; fore coxa dark on basal 1/2, sometimes entirely so; mid and hind coxae yellow to dark brown, sometimes dark at bases in case of yellowish coxae; femora narrowly dark at tips, dark area on fore femur sometimes occupying distal 1/3; tibiae narrowly dark at tips; tarsi weakly dark from tip of tarsomere 1 to apical segment.
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">
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<emphasis italics="true" pageId="0" pageNumber="147">Abdomen</emphasis>
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:
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</emphasis>
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subnitidous dark brown to black, sparsely covered with brownish pruinosity.
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="147">Male terminalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3D-H</figureCitation>
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): caudal margin of tergite 9 almost straight or weakly convex at middle, with shallow U-shaped notch at center. Outer gonostylus in dorsal view (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3E</figureCitation>
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) relatively wide on distal part, tip narrowed and curved anteriorly, concaved margin with indistinct teeth. Interbase with outer lobe as long as medial lobe in dorsal view (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3F</figureCitation>
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); outer lobe wide at base, inner basal end situated near middle of medial lobe (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3H</figureCitation>
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); medial lobe strongly narrowed and rod-shaped on distal half, distal part medial to base of outer lobe about 3.5 times as long as wide (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3H</figureCitation>
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); Aedeagus with rod-shaped distal part, twice as long as wide and tip weakly curved ventrally (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3G</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">Female</emphasis>
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. Body length 6.3-9.6 mm, wing length 5.4-8.2 mm. Generally resembling male.
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</paragraph>
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="147">Ovipositor</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3I</figureCitation>
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): dark brown; yellow on cercus, hypogynial valve, and distal 1/4-1/3 of tergite 10; cercus 2-2.5 times as long as tergite 10.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="distribution">
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<paragraph pageId="0" pageNumber="147">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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Japan (Hokkaido, Honshu, Shikoku, and Kyushu islands) (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ulomorpha. A U. amamiana Kato & Kolcsar, sp. nov. (orange), U. nigricolor Alexander, 1924 (blue), U. polytricha Alexander, 1930 (green) B U. longipenis Kato & Kolcsar, sp. nov. (red)." figureDoi="10.3897/zookeys.999.52831.figure5" httpUri="https://binary.pensoft.net/fig/481464" pageId="0" pageNumber="147">5A</figureCitation>
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) and North Korea.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="remarks">
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<paragraph pageId="0" pageNumber="147">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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This species is easily distinguished from the other species of the genus by the following combination of characters: thorax and abdomen excluding legs dark brown to black; wing with distinct dark areas each at origin of Rs, on fork of Rs to crossvein r-m, and outer end of cell d (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ulomorpha nigricolor Alexander, 1924. A habitus, male B, C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view I female ovipositor, lateral view. Abbreviations: cr - cercus; hv - hypogynial valve; s - sternite; t - tergite. Scale bars: 3 mm (A-C); 0.1 mm (D-H); 0.3 mm (I)." figureDoi="10.3897/zookeys.999.52831.figure3" httpUri="https://binary.pensoft.net/fig/481462" pageId="0" pageNumber="147">3B, C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |