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<document id="F205DAFD0548CFC40F249777C6D4ECA8" ID-DOI="10.21248/contrib.entomol.71.1.001-028" ID-ISSN="0005-805X" ID-Zenodo-Dep="5743187" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638271018364" checkinUser="felipe" docAuthor="Assing, Volker &amp; Meybohm, Heinrich" docDate="2021" docId="03FA8790C85F5756B36EFD11383117E1" docLanguage="en" docName="ContribEntomol.71.1.1-28.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Cephennium varispinosum Assing &amp; Meybohm 2021, spec. nov." docType="treatment" docUuid="04C4304D-F84A-400B-8710-15407DE9BCCD" docUuidSource="ZooBank" docVersion="3" lastPageNumber="18" masterDocId="FFC3FFE8C84F5747B00BFFA239001320" masterDocTitle="The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="17" updateTime="1699437454083" updateUser="plazi">
<mods:mods id="FAA8C44BA99D7E08D05CD0EAC5731C22" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E75EE014E4383691F51054062F64BD35">
<mods:title id="1D53F1DE040E082A30623C38EED40E77">The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="62275CE07C60D3357A669CEA95803ACC" type="personal">
<mods:role id="B6F522732F07584FDBF169051A1BCD6F">
<mods:roleTerm id="9F2DB68329AC590046D0079EE16DE845">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E50385A6B2A0E0C2A648510DE6E94B4C">Assing, Volker</mods:namePart>
</mods:name>
<mods:name id="A37221AFC208A7B72E121CDEDC9E8B22" type="personal">
<mods:role id="9C180AD00C8283BBD01C5E6266C3CC01">
<mods:roleTerm id="A571F4B4949107C3ABC30AD22B40F5A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C29381C5057C4AB3B65A93F49FA9C06">Meybohm, Heinrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="9890DAE2D18519DDFF6579CDF6C1C7AE">text</mods:typeOfResource>
<mods:relatedItem id="476A3977057B2783680F96F15A1B605B" type="host">
<mods:titleInfo id="ABAAE63F13FC3F26C9EE50A492633B17">
<mods:title id="DCCAC4521AFA22E63A8B46D8AAE0B4C3">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="CBC9BC95065F7EB94DA47593A2D0510F">
<mods:date id="571BAD310AC94A52CD467A782AA10D34">2021</mods:date>
<mods:detail id="7BF95CCC5AAB2DAAB16DD8B891AEF4EB" type="pubDate">
<mods:number id="51E04C1B8548953ECE06E59C7B550F78">2021-06-30</mods:number>
</mods:detail>
<mods:detail id="934539A48F70DE83E87B5527563DE9CB" type="volume">
<mods:number id="14B30CD23E05E82C282713923D0CDA05">71</mods:number>
</mods:detail>
<mods:detail id="B1DBA6B23A9A5D2B1F58358ADBF797E9" type="issue">
<mods:number id="E65FC7E6B2C844ABFCBD4E7A32EDED05">1</mods:number>
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<mods:start id="8891C19905DA455529A135A6EBC2AA94">1</mods:start>
<mods:end id="FBF3DA8A3194B90BD9A1B7A8DFC8D00C">28</mods:end>
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<mods:classification id="4D356B96449BCABBE053F00FD6FEA336">journal article</mods:classification>
<mods:identifier id="8B87869C43B4FA884A007C86ADC43BAD" type="DOI">10.21248/contrib.entomol.71.1.001-028</mods:identifier>
<mods:identifier id="76724682B355FE440DD227E2EF4A237A" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="66C6F7D0A6C2F15DA3F8BDF00C10AC4F" type="Zenodo-Dep">5743187</mods:identifier>
<mods:identifier id="D24D805BF753100394493C1A6399B7D4" type="ZooBank">3325609F-6C1F-491F-8C48-C07149C0A0E6</mods:identifier>
</mods:mods>
<treatment id="03FA8790C85F5756B36EFD11383117E1" LSID="urn:lsid:zoobank.org:act:04C4304D-F84A-400B-8710-15407DE9BCCD" httpUri="http://treatment.plazi.org/id/03FA8790C85F5756B36EFD11383117E1" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C349650DC85F5757B36EFD113C7511CF" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BEC3686C85F5757B36EFD113DE611EE" blockId="16.[869,1397,691,783]" box="[869,1254,691,718]" pageId="16" pageNumber="17">
<heading id="D0A481EAC85F5757B36EFD113DE611EE" box="[869,1254,691,718]" fontSize="12" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="4C534D05C85F5757B36EFD113D7711EE" ID-CoL="8XWYD" authority="Assing &amp; Meybohm, 2021" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[869,1143,691,718]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="varispinosum" status="sp. nov.">
<emphasis id="B927EA94C85F5757B36EFD113D7711EE" box="[869,1143,691,718]" italics="true" pageId="16" pageNumber="17">Cephennium varispinosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457EFC85F5757B476FD113DE611EE" box="[1149,1254,691,718]" pageId="16" pageNumber="17" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BEC3686C85F5757B361FD793C7511CF" blockId="16.[869,1397,691,783]" box="[874,1397,731,751]" pageId="16" pageNumber="17">
<uri id="FFC23A84C85F5757B361FD793C7511CF" box="[874,1397,731,751]" pageId="16" pageNumber="17">
urn:lsid:zoobank.org:act:
<uuid id="FFF50C53C85F5757B41DFD793C7511CF" box="[1046,1397,731,751]" pageId="16" pageNumber="17">04C4304D-F84A-400B-8710-15407DE9BCCD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C349650DC85F5757B36EFD593D24102F" box="[869,1060,763,783]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEC3686C85F5757B36EFD593D24102F" blockId="16.[869,1397,691,783]" box="[869,1060,763,783]" pageId="16" pageNumber="17">
(
<figureCitation id="13682A03C85F5757B362FD593AAD102F" box="[873,941,763,783]" captionStart-0="Figs 1–8" captionStart-1="Figs 9–16" captionStartId-0="8.[189,229,1980,2002]" captionStartId-1="9.[151,191,1985,2007]" captionTargetBox-0="[189,1433,232,1964]" captionTargetBox-1="[153,1398,233,1971]" captionTargetId-0="figure-103@8.[189,1436,232,1970]" captionTargetId-1="figure-101@9.[151,1398,233,1971]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figs 1–8: Male habitus in dorsal view (1–6) and head and pronotum in lateral view (7–8) of Cephennium bodemeyeri (1), C. fractum (2–3), C. infractum (4), C. immutatum (5), C. ericinum (6–7), and C. varispinosum (8). Scale bars: 1–6: 1.0 mm; 7–8: 0.5 mm." captionText-1="Figs 9–16: Male habitus in dorsal view of Cephennium varispinosum (9), C. victrix (10), C. spinigerum (11), C. digitale (12), C. reductum (13), C. corruptum (14), C. marasicum (15), and C. dolucanum (16). Scale bar: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5743191" figureDoi-1="http://doi.org/10.5281/zenodo.5743193" httpUri-0="https://zenodo.org/record/5743191/files/figure.png" httpUri-1="https://zenodo.org/record/5743193/files/figure.png" pageId="16" pageNumber="17">Figs 8–9</figureCitation>
,
<figureCitation id="13682A03C85F5757B3BFFD593AE6102F" box="[948,998,763,783]" captionStart="Figs 36–47" captionStartId="12.[189,229,1983,2005]" captionTargetBox="[195,1436,243,1957]" captionTargetId="figure-132@12.[189,1436,233,1972]" captionTargetPageId="12" captionText="Figs 36–47: Aedeagus in ventral view (36, 38, 40, 42, 44), internal structures of aedeagus (37, 39, 41, 43, 45), and apical portion of aedeagus in ventral view (46–47) of Cephennium ericinum (36–37), C. varispinosum (38–39), C. victrix (40–41), C. spinigerum (42–43), C. digitale (44–45), C. fractum (46), and C. infractum (47). Scale bars: 36, 38, 40, 42, 44: 0.2 mm; 37, 39, 41, 43, 45–47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743199" httpUri="https://zenodo.org/record/5743199/files/figure.png" pageId="16" pageNumber="17">38–39</figureCitation>
,
<figureCitation id="13682A03C85F5757B3E6FD593D1F102F" box="[1005,1055,763,783]" captionStart="Map 2" captionStartId="16.[190,235,2036,2057]" captionTargetBox="[190,1437,1371,2023]" captionTargetId="figure-542@16.[190,1437,1371,2023]" captionTargetPageId="16" captionText="Map 2: Distributions of Cephennium fractum (black circles) and of the species of the C. ericinum group: C. digitale (white triangle); C. spinigerum (black triangle); C. victrix (white circles); C. varispinosum (black diamond); C. ericinum (white diamond)." figureDoi="http://doi.org/10.5281/zenodo.5743208" httpUri="https://zenodo.org/record/5743208/files/figure.png" pageId="16" pageNumber="17">Map 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C349650DC85F5757B334FC8B3ABD10C1" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BEC3686C85F5757B334FC8B3ABD10C1" blockId="16.[831,1437,809,994]" pageId="16" pageNumber="17">
<emphasis id="B927EA94C85F5757B334FC8B3D5D1061" bold="true" box="[831,1117,809,833]" pageId="16" pageNumber="17">Type material examined</emphasis>
:
<materialsCitation id="3B3B3CDBC85F5757B47FFC8B3D3810E1" collectingDate="2008-07-15" collectionCode="V" elevation="920" location="Akkus" pageId="16" pageNumber="17" specimenCount="2" stateProvince="Ordu" typeStatus="holotype">
<typeStatus id="54E88824C85F5757B47FFC8B3DD71061" box="[1140,1239,809,833]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: “TR [11] –
<collectingRegion id="4997F864C85F5757B334FCEB3A7A1041" box="[831,890,841,865]" country="Turkey" name="Ordu" pageId="16" pageNumber="17">Ordu</collectingRegion>
,
<quantity id="4CAB9B63C85F5757B384FCEB3AD21041" box="[911,978,841,865]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="17" unit="km" value="18.0">18 km</quantity>
NE
<location id="8E8C605DC85F5757B41DFCEB3D5D1041" LSID="urn:lsid:plazi:treatment:03FA8790C85F5756B36EFD11383117E1:8E8C605DC85F5757B41DFCEB3D5D1041" box="[1046,1117,841,865]" name="Akkus" pageId="16" pageNumber="17" stateProvince="Ordu">Akkuş</location>
, 40°56'03&quot;[N], 37°06'47&quot;[E],
<quantity id="4CAB9B63C85F5757B334FCCB3A8410A1" box="[831,900,873,897]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="16" pageNumber="17" unit="m" value="920.0">
<elevation id="007ED1B5C85F5757B334FCCB3A8410A1" box="[831,900,873,897]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="16" pageNumber="17" unit="m" value="920.0">920 m</elevation>
</quantity>
, mixed deciduous forest,
<date id="FFED1046C85F5757B4A1FCCB3C2A10A1" box="[1194,1322,873,897]" pageId="16" pageNumber="17" value="2008-07-15">
<collectingDate id="EFA9E9AEC85F5757B4A1FCCB3C2A10A1" box="[1194,1322,873,897]" pageId="16" pageNumber="17" value="2008-07-15">15.VII.2008</collectingDate>
</date>
,
<collectionCode id="ED42AE43C85F5757B531FCC83C4C10A1" box="[1338,1356,874,897]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="17" type="Museum">V</collectionCode>
. Assing /
<typeStatus id="54E88824C85F5757B35CFC2B3AC61081" box="[855,966,905,929]" pageId="16" pageNumber="17" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C534D05C85F5757B3F2FC2B3C191081" authority="Assing &amp; Meybohm, 2021" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[1017,1305,905,929]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="varispinosum" status="sp. nov.">
<emphasis id="B927EA94C85F5757B3F2FC2B3C191081" box="[1017,1305,905,929]" italics="true" pageId="16" pageNumber="17">Cephennium varispinosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457EFC85F5757B523FC283C611081" box="[1320,1377,906,929]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
, det.
<collectionCode id="ED42AE43C85F5757B334FC083A5110E1" box="[831,849,938,961]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="17" type="Museum">V</collectionCode>
. Assing 2020” (cAss)
</materialsCitation>
.
<typeStatus id="54E88824C85F5757B44FFC083DA110E2" box="[1092,1185,938,962]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
: same data as holotype (cAss).
</paragraph>
</subSubSection>
<subSubSection id="C349650DC85F5757B334FBAB3DFB17A1" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BEC3686C85F5757B334FBAB3DFB17A1" blockId="16.[831,1437,1033,1153]" pageId="16" pageNumber="17">
<emphasis id="B927EA94C85F5757B334FBAB3AB61701" bold="true" box="[831,950,1033,1057]" pageId="16" pageNumber="17">Etymology</emphasis>
: The specific epithet is an adjective composed of the adjective varius (various) and spinosum (spinose). It alludes to the presence of spines of various sizes and shapes in the internal sac of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C349650DC85F5756B334FB0B38371721" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEC3686C85F5757B334FB0B3C771601" blockId="16.[831,1436,1193,1313]" pageId="16" pageNumber="17">
<emphasis id="B927EA94C85F5757B334FB0B3AC117E1" bold="true" box="[831,961,1193,1217]" pageId="16" pageNumber="17">Description</emphasis>
: Body length
<quantity id="4CAB9B63C85F5757B457FB0B3DDD17E1" box="[1116,1245,1193,1217]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="16" pageNumber="17" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
; length of pronotum
<quantity id="4CAB9B63C85F5757B37CFB6B3AD717C1" box="[887,983,1225,1249]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="17" unit="mm" value="0.45">0.45 mm</quantity>
; width of pronotum
<quantity id="4CAB9B63C85F5757B4C3FB6B3C2917C1" box="[1224,1321,1225,1249]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="16" pageNumber="17" unit="mm" value="0.54">0.54 mm</quantity>
; length of antennae approximately
<quantity id="4CAB9B63C85F5757B459FB4B3DA61621" box="[1106,1190,1257,1281]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="mm" value="0.6">0.6 mm</quantity>
. Habitus as in
<figureCitation id="13682A03C85F5757B550FB4B3C981621" box="[1371,1432,1257,1281]" captionStart="Figs 9–16" captionStartId="9.[151,191,1985,2007]" captionTargetBox="[153,1398,233,1971]" captionTargetId="figure-101@9.[151,1398,233,1971]" captionTargetPageId="9" captionText="Figs 9–16: Male habitus in dorsal view of Cephennium varispinosum (9), C. victrix (10), C. spinigerum (11), C. digitale (12), C. reductum (13), C. corruptum (14), C. marasicum (15), and C. dolucanum (16). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743193" httpUri="https://zenodo.org/record/5743193/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
. Colouration: forebody reddish-brown; elytra reddish.
</paragraph>
<caption id="DF2C660EC85F5757B0B5F8563C371B07" ID-DOI="http://doi.org/10.5281/zenodo.5743208" ID-Zenodo-Dep="5743208" httpUri="https://zenodo.org/record/5743208/files/figure.png" pageId="16" pageNumber="17" startId="16.[190,235,2036,2057]" targetBox="[190,1437,1371,2023]" targetPageId="16">
<paragraph id="8BEC3686C85F5757B0B5F8563C9E1B2A" blockId="16.[190,1438,2036,2087]" box="[190,1438,2036,2058]" pageId="16" pageNumber="17">
<emphasis id="B927EA94C85F5757B0B5F85639FC1B29" bold="true" box="[190,252,2036,2057]" pageId="16" pageNumber="17">Map 2</emphasis>
: Distributions of
<taxonomicName id="4C534D05C85F5757B1AFF8563B6C1B29" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[420,620,2036,2057]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="fractum">
<emphasis id="B927EA94C85F5757B1AFF8563B6C1B29" box="[420,620,2036,2057]" italics="true" pageId="16" pageNumber="17">Cephennium fractum</emphasis>
</taxonomicName>
(black circles) and of the species of the
<taxonomicName id="4C534D05C85F5757B3ECF8563D551B2A" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[999,1109,2036,2058]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ericinum">
<emphasis id="B927EA94C85F5757B3ECF8563D551B2A" box="[999,1109,2036,2058]" italics="true" pageId="16" pageNumber="17">C. ericinum</emphasis>
</taxonomicName>
group:
<taxonomicName id="4C534D05C85F5757B495F8563DFE1B29" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[1182,1278,2036,2057]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="digitale">
<emphasis id="B927EA94C85F5757B495F8563DFE1B29" box="[1182,1278,2036,2057]" italics="true" pageId="16" pageNumber="17">C. digitale</emphasis>
</taxonomicName>
(white triangle);
</paragraph>
<paragraph id="8BEC3686C85F5757B0B5F7B03C371B07" blockId="16.[190,1438,2036,2087]" box="[190,1335,2065,2087]" pageId="16" pageNumber="17">
<taxonomicName id="4C534D05C85F5757B0B5F7B038401B07" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[190,320,2066,2087]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="spinigerum">
<emphasis id="B927EA94C85F5757B0B5F7B038401B07" box="[190,320,2066,2087]" italics="true" pageId="16" pageNumber="17">C. spinigerum</emphasis>
</taxonomicName>
(black triangle);
<taxonomicName id="4C534D05C85F5757B1E8F7B03B3B1B07" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[483,571,2066,2087]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="victrix">
<emphasis id="B927EA94C85F5757B1E8F7B03B3B1B07" box="[483,571,2066,2087]" italics="true" pageId="16" pageNumber="17">C. victrix</emphasis>
</taxonomicName>
(white circles);
<taxonomicName id="4C534D05C85F5757B2D9F7B03A6B1B07" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[722,875,2066,2087]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="varispinosum">
<emphasis id="B927EA94C85F5757B2D9F7B03A6B1B07" box="[722,875,2066,2087]" italics="true" pageId="16" pageNumber="17">C. varispinosum</emphasis>
</taxonomicName>
(black diamond);
<taxonomicName id="4C534D05C85F5757B410F7B03D891B07" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[1051,1161,2066,2087]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ericinum">
<emphasis id="B927EA94C85F5757B410F7B03D891B07" box="[1051,1161,2066,2087]" italics="true" pageId="16" pageNumber="17">C. ericinum</emphasis>
</taxonomicName>
(white diamond).
</paragraph>
</caption>
<paragraph id="8BEC3686C85E5756B09CFF4838351121" blockId="17.[151,757,233,769]" pageId="17" pageNumber="18">
Eyes reduced to small rudiments without ommatidia (
<figureCitation id="13682A03C85E5756B094FEAB39DC1201" box="[159,220,265,289]" captionStart="Figs 1–8" captionStartId="8.[189,229,1980,2002]" captionTargetBox="[189,1433,232,1964]" captionTargetId="figure-103@8.[189,1436,232,1970]" captionTargetPageId="8" captionText="Figs 1–8: Male habitus in dorsal view (1–6) and head and pronotum in lateral view (7–8) of Cephennium bodemeyeri (1), C. fractum (2–3), C. infractum (4), C. immutatum (5), C. ericinum (6–7), and C. varispinosum (8). Scale bars: 1–6: 1.0 mm; 7–8: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743191" httpUri="https://zenodo.org/record/5743191/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
). Antennomeres VIII–X strongly transverse. Pronotum rather large in relation to elytra and as broad as elytra, 1.2 times as broad as long, broadest in anterior half, and strongly convex in cross-section; lateral margins straight in posterior half. Elytra without tomentose impressions anteriorly; humeral carina distinct, approximately as long as the combined length of the basal 3–4 antennomeres.
</paragraph>
<paragraph id="8BEC3686C85E5756B09CFDA83BBB1021" blockId="17.[151,757,233,769]" pageId="17" pageNumber="18">
: protibia subapically not curved, but somewhat flattened on inner side; aedeagus (
<figureCitation id="13682A03C85E5756B1D2FD8B3B4F1161" box="[473,591,553,577]" captionStart="Figs 36–47" captionStartId="12.[189,229,1983,2005]" captionTargetBox="[195,1436,243,1957]" captionTargetId="figure-132@12.[189,1436,233,1972]" captionTargetPageId="12" captionText="Figs 36–47: Aedeagus in ventral view (36, 38, 40, 42, 44), internal structures of aedeagus (37, 39, 41, 43, 45), and apical portion of aedeagus in ventral view (46–47) of Cephennium ericinum (36–37), C. varispinosum (38–39), C. victrix (40–41), C. spinigerum (42–43), C. digitale (44–45), C. fractum (46), and C. infractum (47). Scale bars: 36, 38, 40, 42, 44: 0.2 mm; 37, 39, 41, 43, 45–47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743199" httpUri="https://zenodo.org/record/5743199/files/figure.png" pageId="17" pageNumber="18">Figs 38–39</figureCitation>
)
<quantity id="4CAB9B63C85E5756B257FD8B3BBA1161" box="[604,698,553,577]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="17" pageNumber="18" unit="mm" value="0.38">0.38 mm</quantity>
long; ventral process gradually narrowed apicad in ventral view; internal sac with approximately eight weakly sclerotized and rather small spines of various sizes and shapes ventrally, and with a pair of larger and strongly curved spines dorsally; parameres very thin and indistinct, not sinuate, extending to apical third of ventral process.
</paragraph>
<paragraph id="8BEC3686C85E5756B09CFC8B38371721" blockId="17.[151,756,809,1025]" pageId="17" pageNumber="18">
<emphasis id="B927EA94C85E5756B09CFC8B38691061" bold="true" box="[151,361,809,833]" pageId="17" pageNumber="18">Comparative notes</emphasis>
: Aside from its aedeagus,
<taxonomicName id="4C534D05C85E5756B28CFC8B39DD1041" authorityName="Assing &amp; Meybohm" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="varispinosum">
<emphasis id="B927EA94C85E5756B28CFC8B39DD1041" italics="true" pageId="17" pageNumber="18">C. varispinosum</emphasis>
</taxonomicName>
is characterized among the Turkish
<taxonomicName id="4C534D05C85E5756B265FCEB39E210A1" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B927EA94C85E5756B265FCEB3BF41041" box="[622,756,841,865]" italics="true" pageId="17" pageNumber="18">Cephennium</emphasis>
species
</taxonomicName>
by a rather large pronotum, strongly reduced eyes, and strongly transverse antennomeres VIII–X. It is additionally separated from the geographically close
<taxonomicName id="4C534D05C85E5756B09CFC6B381210C1" authorityName="Assing &amp; Meybohm" authorityYear="2021" box="[151,274,969,993]" class="Insecta" family="Staphylinidae" genus="Cephennium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ericinum">
<emphasis id="B927EA94C85E5756B09CFC6B381210C1" box="[151,274,969,993]" italics="true" pageId="17" pageNumber="18">C. ericinum</emphasis>
</taxonomicName>
by much smaller size and weakly modified male protibiae.
</paragraph>
</subSubSection>
<subSubSection id="C349650DC85E5756B09CFB8B383117E1" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BEC3686C85E5756B09CFB8B383117E1" blockId="17.[151,756,1065,1217]" pageId="17" pageNumber="18">
<emphasis id="B927EA94C85E5756B09CFB8B3B091761" bold="true" box="[151,521,1065,1089]" pageId="17" pageNumber="18">Distribution and natural history</emphasis>
: The
<typeStatus id="54E88824C85E5756B25BFB883B7E1762" box="[592,638,1066,1090]" pageId="17" pageNumber="18">type</typeStatus>
locality is situated to the northeast of Akkuş,
<collectingRegion id="4997F864C85E5756B200FBEB3BA91741" box="[523,681,1097,1121]" country="Turkey" name="Ordu" pageId="17" pageNumber="18">Ordu province</collectingRegion>
, North
<collectingCountry id="F3447616C85E5756B09CFBCB39F217A1" box="[151,242,1129,1153]" name="Turkey" pageId="17" pageNumber="18">Anatolia</collectingCountry>
(
<figureCitation id="13682A03C85E5756B10FFBCB384B17A1" box="[260,331,1129,1153]" captionStart="Map 2" captionStartId="16.[190,235,2036,2057]" captionTargetBox="[190,1437,1371,2023]" captionTargetId="figure-542@16.[190,1437,1371,2023]" captionTargetPageId="16" captionText="Map 2: Distributions of Cephennium fractum (black circles) and of the species of the C. ericinum group: C. digitale (white triangle); C. spinigerum (black triangle); C. victrix (white circles); C. varispinosum (black diamond); C. ericinum (white diamond)." figureDoi="http://doi.org/10.5281/zenodo.5743208" httpUri="https://zenodo.org/record/5743208/files/figure.png" pageId="17" pageNumber="18">Map 2</figureCitation>
). The specimens were sifted from leaf litter in a mixed forest with predominant
<taxonomicName id="4C534D05C85E5756B246FB283B8A1781" box="[589,650,1162,1185]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B927EA94C85E5756B246FB283B8A1781" box="[589,650,1162,1185]" italics="true" pageId="17" pageNumber="18">Fagus</emphasis>
</taxonomicName>
at an altitude of
<specimenCount id="9D55FD0FC85E5756B0E3FB0B383117E1" box="[232,305,1193,1217]" pageId="17" pageNumber="18" type="generic">
<quantity id="4CAB9B63C85E5756B0E3FB0B382C17E1" box="[232,300,1193,1217]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="17" pageNumber="18" unit="m" value="620.0">620 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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