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<document id="051A11F50B67EC30F65615360F6358E1" ID-CLB-Dataset="298421" ID-DOI="10.11646/zootaxa.5463.1.9" ID-GBIF-Dataset="4d1f32e1-5ee2-4869-b5bb-d90e5cfae15e" ID-ISSN="1175-5326" ID-Zenodo-Dep="11610836" ID-ZooBank="2D36452E-1238-4DEF-A6A9-8370227FF31F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718181404149" checkinUser="plazi" docAuthor="Vilkamaa, Pekka" docDate="2024" docId="03EEA42FFFE02477FF4CD1BB8032FDDA" docLanguage="en" docName="zootaxa.5463.1.9.pdf" docOrigin="Zootaxa 5463 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5463.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Claustropyga alveata Vilkamaa 2024, sp. n." docType="treatment" docVersion="5" lastPageNumber="141" masterDocId="FFD7DC57FFE12473FFDBD011812AFFC3" masterDocTitle="Description of two new species of Claustropyga Hippa, Vilkamaa &amp; Mohrig (Diptera, Sciaridae) from the northern Palaearctic" masterLastPageNumber="144" masterPageNumber="137" pageNumber="138" updateTime="1718217206606" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5253067B398BEF14DBB043B986148815">Description of two new species of Claustropyga Hippa, Vilkamaa &amp; Mohrig (Diptera, Sciaridae) from the northern Palaearctic</mods:title>
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<mods:date id="B37DE0ED59216A6E1BED5CE4512CCA5D">2024</mods:date>
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<treatment id="03EEA42FFFE02477FF4CD1BB8032FDDA" ID-DOI="http://doi.org/10.5281/zenodo.11611049" ID-GBIF-Taxon="232735749" ID-Zenodo-Dep="11611049" LSID="urn:lsid:plazi:treatment:03EEA42FFFE02477FF4CD1BB8032FDDA" httpUri="http://treatment.plazi.org/id/03EEA42FFFE02477FF4CD1BB8032FDDA" lastPageId="4" lastPageNumber="141" pageId="1" pageNumber="138">
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<paragraph id="8BF81539FFE02472FF4CD1BB80CAFE06" blockId="1.[151,480,426,489]" box="[151,480,426,453]" pageId="1" pageNumber="138">
<heading id="D0B0A255FFE02472FF4CD1BB80CAFE06" bold="true" box="[151,480,426,453]" fontSize="11" level="1" pageId="1" pageNumber="138" reason="1">
<emphasis id="B933C92BFFE02472FF4CD1BB80CAFE06" bold="true" box="[151,480,426,453]" pageId="1" pageNumber="138">
<taxonomicName id="4C476EBAFFE02472FF4CD1BB80BCFE06" authority="Vilkamaa, 2024" authorityName="Vilkamaa" authorityYear="2024" box="[151,406,426,453]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="alveata" status="sp. nov.">
<emphasis id="B933C92BFFE02472FF4CD1BB80BCFE06" bold="true" box="[151,406,426,453]" italics="true" pageId="1" pageNumber="138">Claustropyga alveata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2007450FFE02472FE45D1BA80CAFE06" box="[414,480,427,453]" pageId="1" pageNumber="138" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8BF81539FFE02472FF4CD1DE8004FE2A" blockId="1.[151,480,426,489]" box="[151,302,463,489]" pageId="1" pageNumber="138">
<figureCitation id="137C09BCFFE02472FF4CD1DE81D1FE2A" box="[151,251,463,489]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1983,2007]" captionStartId-1="2.[151,250,1570,1594]" captionStartId-2="3.[151,250,1967,1991]" captionTargetBox-0="[409,1199,540,1883]" captionTargetBox-1="[171,1348,993,1547]" captionTargetBox-2="[311,1276,309,1943]" captionTargetId-0="figure-87@1.[328,1259,510,1957]" captionTargetId-1="figure-408@2.[171,1415,993,1547]" captionTargetId-2="figure-78@3.[311,1276,309,1943]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Claustropyga alveata sp. n. (A holotype, B paratype). A. Hypopygium, ventral. B. Hypopygium, dorsal. Scale 0.1 mm." captionText-1="FIGURE 2. Claustropyga alveata sp. n. (holotype). A. Gonostylus, dorsal. B. Apical part of fore tibia, prolateral. Scale for A 0.1 mm, for B 0.05 mm." captionText-2="FIGURE 3. Claustropyga alveata sp. n. (paratype). Medial structures of hypopygium, ventral. A. Focus on ventral portion of tegmen. B. Focus on dorsal portion of tegmen. C. Focus on apodeme of gonocoxa. Scale 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11610842" figureDoi-1="http://doi.org/10.5281/zenodo.11610854" figureDoi-2="http://doi.org/10.5281/zenodo.11610858" httpUri-0="https://zenodo.org/record/11610842/files/figure.png" httpUri-1="https://zenodo.org/record/11610854/files/figure.png" httpUri-2="https://zenodo.org/record/11610858/files/figure.png" pageId="1" pageNumber="138">Figs 13</figureCitation>
,
<figureCitation id="137C09BCFFE02472FEDDD1DE8004FE2A" box="[262,302,463,489]" captionStart="FIGURE 4" captionStartId="4.[151,250,1433,1457]" captionTargetBox="[158,1436,597,1402]" captionTargetId="figure-187@4.[151,1436,597,1409]" captionTargetPageId="4" captionText="FIGURE 4. Gonostylus, ventral. A. Claustropyga alveata sp. n. (holotype). B. C. caetrata (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610860" httpUri="https://zenodo.org/record/11610860/files/figure.png" pageId="1" pageNumber="138">4 A</figureCitation>
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<paragraph id="8BF81539FFE02472FF4CD7AE81ECF839" blockId="1.[151,1437,1983,2042]" pageId="1" pageNumber="138">
<emphasis id="B933C92BFFE02472FF4CD7AE8039F814" bold="true" box="[151,275,1983,2007]" pageId="1" pageNumber="138">FIGURE 1.</emphasis>
<taxonomicName id="4C476EBAFFE02472FEC2D7AE80DEF814" authority="Vilkamaa, 2024" authorityName="Vilkamaa" authorityYear="2024" box="[281,500,1983,2007]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="alveata" status="sp. nov.">
<emphasis id="B933C92BFFE02472FEC2D7AE80DEF814" box="[281,500,1983,2007]" italics="true" pageId="1" pageNumber="138">Claustropyga alveata</emphasis>
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<emphasis id="B933C92BFFE02472FE22D7AE8318F814" bold="true" box="[505,562,1983,2007]" pageId="1" pageNumber="138">
<taxonomicNameLabel id="A2007450FFE02472FE22D7AE8318F814" box="[505,562,1983,2007]" pageId="1" pageNumber="138" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(A holotype, B paratype).
<emphasis id="B933C92BFFE02472FC9AD7AE8270F814" bold="true" box="[833,858,1983,2007]" pageId="1" pageNumber="138">A.</emphasis>
Hypopygium, ventral.
<emphasis id="B933C92BFFE02472FB9ED7AE8576F814" bold="true" box="[1093,1116,1983,2007]" pageId="1" pageNumber="138">B.</emphasis>
Hypopygium, dorsal. Scale 0.1 mm.
</paragraph>
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<paragraph id="8BF81539FFE32471FF4CD0868514FF3A" blockId="2.[151,1437,150,969]" pageId="2" pageNumber="139">
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<typeStatus id="54FCAB9BFFE32471FF4CD08681D4FF72" box="[151,254,151,177]" pageId="2" pageNumber="139" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
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,
<location id="8E9843E2FFE32471FE3CD086832FFF72" LSID="urn:lsid:plazi:treatment:03EEA42FFFE02477FF4CD1BB8032FDDA:8E9843E2FFE32471FE3CD086832FFF72" box="[487,517,151,177]" country="Finland" county="Annjaloanji" municipality="Enontekio" name="Le" pageId="2" pageNumber="139">Le</location>
(Lapponia enontekiensis),
<collectingMunicipality id="6B9C8F43FFE32471FC92D0868295FF72" box="[841,959,151,177]" pageId="2" pageNumber="139">Enontekiö</collectingMunicipality>
,
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(Grid 76860:2806),
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<collectingDate id="EFBDCA11FFE32471FA8BD087802AFF16" pageId="2" pageNumber="139" value="2007-07-11" valueMax="2007-07-15" valueMin="2007-07-11">1115. VII.2007</collectingDate>
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,
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. Jussila (
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)
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.
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<typeStatus id="54FCAB9BFFE32471FDE8D0AA83B0FF17" box="[563,666,187,212]" pageId="2" pageNumber="139" type="paratype">Paratype</typeStatus>
</emphasis>
.
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,
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,
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,
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<collectingDate id="EFBDCA11FFE32471FF4CD0CE80AEFF3A" box="[151,388,223,249]" pageId="2" pageNumber="139" value="2009-07-14" valueMax="2009-08-21" valueMin="2009-07-14">14.VII21.VIII.2009</collectingDate>
</date>
,
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.
<materialsCitation id="3B2F1F64FFE32471FE7DD0CE8510FF3A" ID-GBIF-Occurrence="4876418304" box="[422,1082,223,249]" collectionCode="TMU" collectorName="E. Barstad &amp; Sample ID" country="Norway" pageId="2" pageNumber="139" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectorName id="26B270EFFFE32471FE7DD0CE8336FF3A" box="[422,540,223,249]" pageId="2" pageNumber="139">E. Barstad</collectorName>
,
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(
<collectorName id="26B270EFFFE32471FD5AD0CE822AFF3A" box="[641,768,223,249]" pageId="2" pageNumber="139">Sample ID</collectorName>
: TSZD-JKJ-103800;
<collectionCode id="ED568DFCFFE32471FC34D0CE8518FF3A" box="[1007,1074,223,249]" pageId="2" pageNumber="139">TMU</collectionCode>
)
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.
</paragraph>
<paragraph id="8BF81539FFE32471FF1CD112832EFC0A" blockId="2.[151,1437,150,969]" pageId="2" pageNumber="139">
Male.
<emphasis id="B933C92BFFE32471FED6D1128067FEDE" bold="true" box="[269,333,259,285]" pageId="2" pageNumber="139">Head</emphasis>
. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with 6 scattered strong setae. Clypeus with 1 seta. Maxillary palpus with 3 segments; 1
<superScript id="7C32B871FFE32471FBA0D13685ACFEF6" attach="left" box="[1147,1158,295,309]" fontSize="6" pageId="2" pageNumber="139">st</superScript>
segment longer than 2
<superScript id="7C32B871FFE32471FA57D13784B6FEF7" attach="none" box="[1420,1436,294,308]" fontSize="6" pageId="2" pageNumber="139">nd</superScript>
segment, 3
<superScript id="7C32B871FFE32471FED4D15B8037FE9B" attach="left" box="[271,285,330,344]" fontSize="6" pageId="2" pageNumber="139">rd</superScript>
segment shortest; 1
<superScript id="7C32B871FFE32471FE2CD15A8328FE9A" attach="left" box="[503,514,331,345]" fontSize="6" pageId="2" pageNumber="139">st</superScript>
segment with 1 sharp seta, with a large dorsal patch of sensilla. Body of 4
<superScript id="7C32B871FFE32471FAF0D15B8412FE9B" attach="left" box="[1323,1336,330,344]" fontSize="6" pageId="2" pageNumber="139">th</superScript>
antennal flagellomere 1.65x as long as wide, the neck shorter than wide, the longest setae as long as the width of flagellomere.
<emphasis id="B933C92BFFE32471FF4CD18281DAFE6E" bold="true" box="[151,240,403,429]" pageId="2" pageNumber="139">Thorax</emphasis>
. Brown, setae dark. Scutum with some long and strong laterals and long dorsocentrals. Scutellum with 26 long and strong setae and with some short setae. Anterior pronotum with 15 setae. Proepisternum with 34 setae.
<emphasis id="B933C92BFFE32471FF4CD1CB81F5FE37" bold="true" box="[151,223,474,500]" pageId="2" pageNumber="139">Wing.</emphasis>
Length
<quantity id="4CBFB8DCFFE32471FEE4D1CA80B3FE36" box="[319,409,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="139" unit="mm" value="2.1">2.1 mm</quantity>
. Width/length 0.45. R
<subScript id="17C3177CFFE32471FD49D1FC83B0FE38" attach="left" box="[658,666,493,507]" fontSize="6" pageId="2" pageNumber="139">1</subScript>
/R 0.650.95. stM longer or shorter than fork of M. bM longer than r-m. bM non-setose, r-m non-setose or with 3 dorsal setae. Halter pale brown.
<emphasis id="B933C92BFFE32471FBC0D1EE8579FDDA" bold="true" box="[1051,1107,511,537]" pageId="2" pageNumber="139">Legs</emphasis>
. Fore coxa yellow, mid- and hind coxae pale brown, coxal setae dark. Fore tibial organ with indistinct patch of few strong setae. Fore tibial spur slightly longer than tibial width.
<emphasis id="B933C92BFFE32471FDDCD2578356FDA3" bold="true" box="[519,636,582,608]" pageId="2" pageNumber="139">Abdomen</emphasis>
. Pale brown, setae long and strong.
<emphasis id="B933C92BFFE32471FBC9D2578584FDA3" bold="true" box="[1042,1198,582,608]" pageId="2" pageNumber="139">Hypopygium</emphasis>
(
<figureCitation id="137C09BCFFE32471FB66D25685D4FDA2" box="[1213,1278,583,609]" captionStart="FIGURE 1" captionStartId="1.[151,250,1983,2007]" captionTargetBox="[409,1199,540,1883]" captionTargetId="figure-87@1.[328,1259,510,1957]" captionTargetPageId="1" captionText="FIGURE 1. Claustropyga alveata sp. n. (A holotype, B paratype). A. Hypopygium, ventral. B. Hypopygium, dorsal. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610842" httpUri="https://zenodo.org/record/11610842/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
). Pale brown, as abdomen. Tergite IX long and wide, nearly covering cerci dorsally. Intergonocoxal area lobe-like produced with roundish medial notch; gonocoxa much longer than gonostylus, with moderately long setosity; with apicoventral elongated seta rather short, gonocoxal apodemes strongly sclerotized, with hook-formed basal extensions; extended caudad forming a V-shaped structure with its closed end pointing caudad (
<figureCitation id="137C09BCFFE32471FC30D2C68565FD32" box="[1003,1103,727,753]" captionStart="FIGURE 3" captionStartId="3.[151,250,1967,1991]" captionTargetBox="[311,1276,309,1943]" captionTargetId="figure-78@3.[311,1276,309,1943]" captionTargetPageId="3" captionText="FIGURE 3. Claustropyga alveata sp. n. (paratype). Medial structures of hypopygium, ventral. A. Focus on ventral portion of tegmen. B. Focus on dorsal portion of tegmen. C. Focus on apodeme of gonocoxa. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610858" httpUri="https://zenodo.org/record/11610858/files/figure.png" pageId="2" pageNumber="139">Fig. 3 C</figureCitation>
). Gonostylus (
<figureCitation id="137C09BCFFE32471FB2ED2C68476FD32" box="[1269,1372,727,753]" captionStart="FIGURE 2" captionStartId="2.[151,250,1570,1594]" captionTargetBox="[171,1348,993,1547]" captionTargetId="figure-408@2.[171,1415,993,1547]" captionTargetPageId="2" captionText="FIGURE 2. Claustropyga alveata sp. n. (holotype). A. Gonostylus, dorsal. B. Apical part of fore tibia, prolateral. Scale for A 0.1 mm, for B 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.11610854" httpUri="https://zenodo.org/record/11610854/files/figure.png" pageId="2" pageNumber="139">Figs 2 A</figureCitation>
,
<figureCitation id="137C09BCFFE32471FAB2D2C684BFFD32" box="[1385,1429,727,753]" captionStart="FIGURE 4" captionStartId="4.[151,250,1433,1457]" captionTargetBox="[158,1436,597,1402]" captionTargetId="figure-187@4.[151,1436,597,1409]" captionTargetPageId="4" captionText="FIGURE 4. Gonostylus, ventral. A. Claustropyga alveata sp. n. (holotype). B. C. caetrata (holotype). Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610860" httpUri="https://zenodo.org/record/11610860/files/figure.png" pageId="2" pageNumber="139">4 A</figureCitation>
) roundish, with short setosity; strongly excavated medially; with straight dorsomedial margin, excavation with central ridge on which 1 megaseta very close to apical tooth and 1 megaseta at middle position; with 1 subapical megaseta at ventral margin; with small pyramidal apical tooth. Tegmen (
<figureCitation id="137C09BCFFE32471FCA8D35282F7FC9E" box="[883,989,835,861]" captionStart="FIGURE 1" captionStartId="1.[151,250,1983,2007]" captionTargetBox="[409,1199,540,1883]" captionTargetId="figure-87@1.[328,1259,510,1957]" captionTargetPageId="1" captionText="FIGURE 1. Claustropyga alveata sp. n. (A holotype, B paratype). A. Hypopygium, ventral. B. Hypopygium, dorsal. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610842" httpUri="https://zenodo.org/record/11610842/files/figure.png" pageId="2" pageNumber="139">Figs 1 A</figureCitation>
,
<figureCitation id="137C09BCFFE32471FC37D3528511FC9E" box="[1004,1083,835,861]" captionStart="FIGURE 3" captionStartId="3.[151,250,1967,1991]" captionTargetBox="[311,1276,309,1943]" captionTargetId="figure-78@3.[311,1276,309,1943]" captionTargetPageId="3" captionText="FIGURE 3. Claustropyga alveata sp. n. (paratype). Medial structures of hypopygium, ventral. A. Focus on ventral portion of tegmen. B. Focus on dorsal portion of tegmen. C. Focus on apodeme of gonocoxa. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610858" httpUri="https://zenodo.org/record/11610858/files/figure.png" pageId="2" pageNumber="139">3 A, B</figureCitation>
) long, constricted subapically, weakly sclerotized laterally; apical part unsclerotized; with divergent ventromedial sclerotizations extended from middle to apical part; with strongly sclerotized V-formed apodeme; with small area of tiny needle-like aedeagal teeth. Aedeagal apodeme strong.
</paragraph>
<caption id="DF3845B1FFE32471FF4CD63380A4F99D" ID-DOI="http://doi.org/10.5281/zenodo.11610854" ID-Zenodo-Dep="11610854" httpUri="https://zenodo.org/record/11610854/files/figure.png" pageId="2" pageNumber="139" startId="2.[151,250,1570,1594]" targetBox="[171,1348,993,1547]" targetPageId="2" targetType="figure">
<paragraph id="8BF81539FFE32471FF4CD63380A4F99D" blockId="2.[151,1436,1570,1630]" pageId="2" pageNumber="139">
<emphasis id="B933C92BFFE32471FF4CD633803EF9F9" bold="true" box="[151,276,1570,1594]" pageId="2" pageNumber="139">FIGURE 2.</emphasis>
<taxonomicName id="4C476EBAFFE32471FEC0D63380DCF9F9" authority="Vilkamaa, 2024" authorityName="Vilkamaa" authorityYear="2024" box="[283,502,1570,1594]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="alveata" status="sp. nov.">
<emphasis id="B933C92BFFE32471FEC0D63380DCF9F9" box="[283,502,1570,1594]" italics="true" pageId="2" pageNumber="139">Claustropyga alveata</emphasis>
</taxonomicName>
<emphasis id="B933C92BFFE32471FE27D633831CF9F9" bold="true" box="[508,566,1570,1594]" pageId="2" pageNumber="139">
<taxonomicNameLabel id="A2007450FFE32471FE27D633831CF9F9" box="[508,566,1570,1594]" pageId="2" pageNumber="139" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype).
<emphasis id="B933C92BFFE32471FD6FD63383E7F9F9" bold="true" box="[692,717,1570,1594]" pageId="2" pageNumber="139">A.</emphasis>
Gonostylus, dorsal.
<emphasis id="B933C92BFFE32471FC7BD633829DF9F9" bold="true" box="[928,951,1570,1594]" pageId="2" pageNumber="139">B.</emphasis>
Apical part of fore tibia, prolateral. Scale for A 0.1 mm, for B 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BF81539FFE32471FF1CD6A180B5F909" blockId="2.[151,1437,1712,2027]" box="[199,415,1712,1739]" pageId="2" pageNumber="139">
<emphasis id="B933C92BFFE32471FF1CD6A18031F909" box="[199,283,1712,1738]" italics="true" pageId="2" pageNumber="139">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BF81539FFE32471FF1CD6C480D3F8D1" blockId="2.[151,1437,1712,2027]" pageId="2" pageNumber="139">
<emphasis id="B933C92BFFE32471FF1CD6C481D5F92C" box="[199,255,1749,1775]" italics="true" pageId="2" pageNumber="139">DNA</emphasis>
. Barcode COI, BIN BOLD ADL3758. Deposited in The Barcode of Life Data System (BOLD) (
<bibRefCitation id="EFD668C8FFE32471FF44D6E980D8F8D1" author="Rathasingham, S. &amp; Hebert, P. D. N." box="[159,498,1784,1810]" pageId="2" pageNumber="139" pagination="355 - 364" refId="ref3081" refString="Rathasingham, S. &amp; Hebert, P. D. N. (2007) BOLD: The Barcode of Lifa Data System (www. barcodinglife. org). Molecular ecology Note, 7, 355 - 364. https: // doi. org / 10.1111 / j. 1471 - 8286.2007.01678. x" type="journal article" year="2007">Rathasingham &amp; Hebert 2007</bibRefCitation>
)
</paragraph>
<paragraph id="8BF81539FFE32470FF1CD70C8570FEDE" blockId="2.[151,1437,1712,2027]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="140" pageId="2" pageNumber="139">
<emphasis id="B933C92BFFE32471FF1CD70C8068F8F5" box="[199,322,1821,1846]" italics="true" pageId="2" pageNumber="139">Discussion</emphasis>
.
<taxonomicName id="4C476EBAFFE32471FE88D70D8316F8F5" authority="Vilkamaa, 2024" authorityName="Vilkamaa" authorityYear="2024" box="[339,572,1820,1846]" class="Insecta" family="Sciaridae" genus="Claustopyga" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="alveata" status="sp. nov.">
<emphasis id="B933C92BFFE32471FE88D70D8316F8F5" box="[339,572,1820,1846]" italics="true" pageId="2" pageNumber="139">Claustopyga alveata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2007450FFE32471FD9DD70D83ADF8F4" box="[582,647,1820,1847]" pageId="2" pageNumber="139" rank="species">
<emphasis id="B933C92BFFE32471FD9DD70D83ABF8F5" bold="true" box="[582,641,1820,1846]" pageId="2" pageNumber="139">sp. n</emphasis>
.
</taxonomicNameLabel>
has unusual, strongly sclerotized, vertically superimposed structures on the medial part of the hypopygium (
<figureCitation id="137C09BCFFE32471FDAED751822BF899" box="[629,769,1856,1882]" captionStart="FIGURE 3" captionStartId="3.[151,250,1967,1991]" captionTargetBox="[311,1276,309,1943]" captionTargetId="figure-78@3.[311,1276,309,1943]" captionTargetPageId="3" captionText="FIGURE 3. Claustropyga alveata sp. n. (paratype). Medial structures of hypopygium, ventral. A. Focus on ventral portion of tegmen. B. Focus on dorsal portion of tegmen. C. Focus on apodeme of gonocoxa. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610858" httpUri="https://zenodo.org/record/11610858/files/figure.png" pageId="2" pageNumber="139">Fig. 3 AC</figureCitation>
); dorsally the V-shaped gonocoxal apodeme is extended caudally, ventral to this the apodeme of the tegmen is also V-shaped, and at the ventralmost position, two rod-like medial sclerotizations of the tegmen extend from the basal part to the apical part of the tegmen. Whether these ventral rods are united into the apodeme of tegmen, is not detectable in the preparations. Usually in
<taxonomicName id="4C476EBAFFE32471FAD8D7BD8444F805" authorityName="Billberg" authorityYear="1820" box="[1283,1390,1964,1990]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
, the apodeme of tegmen, mediad of the usually strongly sclerotized lateral parts (which are usually called apodemes of tegmen), is a transverse or apically smoothly curved variably sclerotized bridge. In general, the basal part of the gonocoxal apodeme consists of a transverse bridge uniting the left and right apodemes, with simple longer or shorter proximally orientated rods, not the hook-like structures present in
<emphasis id="B933C92BFFE22470FC07D0CE85FCFF3A" box="[988,1238,223,249]" italics="true" pageId="3" pageNumber="140">
<taxonomicName id="4C476EBAFFE22470FC07D0CE85F8FF3A" authorityName="Vilkamaa" authorityYear="2024" box="[988,1234,223,249]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alveata">Claustropyga alveata</taxonomicName>
.
</emphasis>
The combination of sclerotized structures in the hypopygium of this species is unique among
<taxonomicName id="4C476EBAFFE22470FC31D112857CFEDE" authorityName="Billberg" authorityYear="1820" box="[1002,1110,259,285]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
.
</paragraph>
<caption id="DF3845B1FFE22470FF4CD7BE855FF828" ID-DOI="http://doi.org/10.5281/zenodo.11610858" ID-Zenodo-Dep="11610858" httpUri="https://zenodo.org/record/11610858/files/figure.png" pageId="3" pageNumber="140" startId="3.[151,250,1967,1991]" targetBox="[311,1276,309,1943]" targetPageId="3" targetType="figure">
<paragraph id="8BF81539FFE22470FF4CD7BE855FF828" blockId="3.[151,1436,1967,2028]" pageId="3" pageNumber="140">
<emphasis id="B933C92BFFE22470FF4CD7BE8039F804" bold="true" box="[151,275,1967,1991]" pageId="3" pageNumber="140">FIGURE 3.</emphasis>
<taxonomicName id="4C476EBAFFE22470FEC0D7BE80DFF804" authority="Vilkamaa, 2024" authorityName="Vilkamaa" authorityYear="2024" box="[283,501,1967,1991]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="alveata" status="sp. nov.">
<emphasis id="B933C92BFFE22470FEC0D7BE80DFF804" box="[283,501,1967,1991]" italics="true" pageId="3" pageNumber="140">Claustropyga alveata</emphasis>
</taxonomicName>
<emphasis id="B933C92BFFE22470FE27D7BE831FF804" bold="true" box="[508,565,1967,1991]" pageId="3" pageNumber="140">
<taxonomicNameLabel id="A2007450FFE22470FE27D7BE831FF804" box="[508,565,1967,1991]" pageId="3" pageNumber="140" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype). Medial structures of hypopygium, ventral.
<emphasis id="B933C92BFFE22470FBBCD7BE8555F804" bold="true" box="[1127,1151,1967,1991]" pageId="3" pageNumber="140">A.</emphasis>
Focus on ventral portion of tegmen.
<emphasis id="B933C92BFFE22470FF35D7C2802CF828" bold="true" box="[238,262,2003,2027]" pageId="3" pageNumber="140">B.</emphasis>
Focus on dorsal portion of tegmen.
<emphasis id="B933C92BFFE22470FDAED7C283A7F828" bold="true" box="[629,653,2003,2027]" pageId="3" pageNumber="140">C.</emphasis>
Focus on apodeme of gonocoxa. Scale 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF81539FFE52477FF1CD0868196FE12" blockId="4.[151,1437,151,537]" pageId="4" pageNumber="141">
By its subapically constricted tegmen,
<taxonomicName id="4C476EBAFFE52477FD5ED0868244FF72" authorityName="Vilkamaa" authorityYear="2024" box="[645,878,151,177]" class="Insecta" family="Sciaridae" genus="Claustopyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="alveata">
<emphasis id="B933C92BFFE52477FD5ED0868244FF72" box="[645,878,151,177]" italics="true" pageId="4" pageNumber="141">Claustopyga alveata</emphasis>
</taxonomicName>
resembles only
<taxonomicName id="4C476EBAFFE52477FBF7D08681F8FF16" authority="Hippa &amp; Vilkamaa, 2016" authorityName="Hippa &amp; Vilkamaa" authorityYear="2016" class="Insecta" family="Sciaridae" genus="Claustopyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="stupenda">
<emphasis id="B933C92BFFE52477FBF7D086859EFF72" box="[1068,1204,151,177]" italics="true" pageId="4" pageNumber="141">C. stupenda</emphasis>
<bibRefCitation id="EFD668C8FFE52477FB65D08681F8FF16" author="Hippa, H. &amp; Vilkamaa, P." pageId="4" pageNumber="141" pagination="594 - 600" refId="ref2724" refString="Hippa, H. &amp; Vilkamaa, P. (2016) New species of Claustropyga Hippa, Vilkamaa &amp; Mohrig (Diptera, Sciaridae) from the Holarctic region. Zootaxa, 4088 (4), 594 - 600. https: // doi. org / 10.11646 / zootaxa. 4088.4.10" type="journal article" year="2016">Hippa &amp; Vilkamaa, 2016</bibRefCitation>
</taxonomicName>
, but the latter has two very long basal megasetae on its gonostylus instead of one short one and lacks the characteristic medial sclerotizations of
<taxonomicName id="4C476EBAFFE52477FD82D0CE83E5FF3A" authorityName="Vilkamaa" authorityYear="2024" box="[601,719,223,249]" class="Insecta" family="Sciaridae" genus="Claustopyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="alveata">
<emphasis id="B933C92BFFE52477FD82D0CE83E5FF3A" box="[601,719,223,249]" italics="true" pageId="4" pageNumber="141">C. alveata</emphasis>
</taxonomicName>
in its hypopygium. Moreover, the gonostylus of
<taxonomicName id="4C476EBAFFE52477FAD1D0CE8455FF3A" authorityName="Vilkamaa" authorityYear="2024" box="[1290,1407,223,249]" class="Insecta" family="Sciaridae" genus="Claustopyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="alveata">
<emphasis id="B933C92BFFE52477FAD1D0CE8455FF3A" box="[1290,1407,223,249]" italics="true" pageId="4" pageNumber="141">C. alveata</emphasis>
</taxonomicName>
is much smaller. In its small, globular gonostylus with a very small apical tooth,
<taxonomicName id="4C476EBAFFE52477FC25D11285C7FEDE" authorityName="Vilkamaa" authorityYear="2024" box="[1022,1261,259,285]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="alveata">
<emphasis id="B933C92BFFE52477FC25D11285C7FEDE" box="[1022,1261,259,285]" italics="true" pageId="4" pageNumber="141">Claustropyga alveata</emphasis>
</taxonomicName>
differs from all other species of the genus.
<taxonomicName id="4C476EBAFFE52477FE10D1368396FE82" authorityName="Vilkamaa" authorityYear="2024" box="[459,700,295,321]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="alveata">
<emphasis id="B933C92BFFE52477FE10D1368396FE82" box="[459,700,295,321]" italics="true" pageId="4" pageNumber="141">Claustropyga alveata</emphasis>
</taxonomicName>
has a long and unusually wide tergite IX, in dorsal aspect nearly covering the cerci (
<figureCitation id="137C09BCFFE52477FEAED15A80FDFEA6" box="[373,471,331,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1983,2007]" captionTargetBox="[409,1199,540,1883]" captionTargetId="figure-87@1.[328,1259,510,1957]" captionTargetPageId="1" captionText="FIGURE 1. Claustropyga alveata sp. n. (A holotype, B paratype). A. Hypopygium, ventral. B. Hypopygium, dorsal. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11610842" httpUri="https://zenodo.org/record/11610842/files/figure.png" pageId="4" pageNumber="141">Fig. 1 B</figureCitation>
). The length of tergite IX is variable amongst
<taxonomicName id="4C476EBAFFE52477FC35D15A857CFEA6" authorityName="Billberg" authorityYear="1820" box="[1006,1110,331,357]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
genera and amongst species of
<taxonomicName id="4C476EBAFFE52477FF6CD17E8064FE4A" authorityName="Hippa, Vilkamaa &amp; Mohrig" authorityYear="2003" box="[183,334,367,393]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B933C92BFFE52477FF6CD17E8064FE4A" box="[183,334,367,393]" italics="true" pageId="4" pageNumber="141">Claustropyga</emphasis>
</taxonomicName>
, with
<taxonomicName id="4C476EBAFFE52477FE43D17E833DFE4B" authority="Vilkamaa, 2024" authorityName="Vilkamaa" authorityYear="2024" box="[408,535,367,393]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="caetrata" status="sp. n">
<emphasis id="B933C92BFFE52477FE43D17E833DFE4B" box="[408,535,367,393]" italics="true" pageId="4" pageNumber="141">C. caetrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2007450FFE52477FDC4D17E837DFE4A" box="[543,599,367,393]" pageId="4" pageNumber="141" rank="species">sp. n</taxonomicNameLabel>
,
<taxonomicName id="4C476EBAFFE52477FDBED17E829DFE4A" authority="(Freeman, 1983)" baseAuthorityName="Freeman" baseAuthorityYear="1983" box="[613,951,367,393]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="abblanda">
<emphasis id="B933C92BFFE52477FDBED17E83DBFE4A" box="[613,753,367,393]" italics="true" pageId="4" pageNumber="141">C. abblanda</emphasis>
(
<bibRefCitation id="EFD668C8FFE52477FCDBD17E8284FE4A" author="Freeman, P." box="[768,942,367,393]" pageId="4" pageNumber="141" pagination="161 - 170" refId="ref2605" refString="Freeman, P. (1983) Revisionary notes on British Sciaridae (Diptera). Entomologist's Monthly Magazine, 119, 161 - 170." type="journal article" year="1983">Freeman, 1983</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C476EBAFFE52477FC1FD17E845CFE4A" authority="Hippa &amp; Vilkamaa, 2016" authorityName="Hippa &amp; Vilkamaa" authorityYear="2016" box="[964,1398,367,393]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="clavulata">
<emphasis id="B933C92BFFE52477FC1FD17E8564FE4A" box="[964,1102,367,393]" italics="true" pageId="4" pageNumber="141">C. clavulata</emphasis>
<bibRefCitation id="EFD668C8FFE52477FB8DD17E845CFE4A" author="Hippa, H. &amp; Vilkamaa, P." box="[1110,1398,367,393]" pageId="4" pageNumber="141" pagination="594 - 600" refId="ref2724" refString="Hippa, H. &amp; Vilkamaa, P. (2016) New species of Claustropyga Hippa, Vilkamaa &amp; Mohrig (Diptera, Sciaridae) from the Holarctic region. Zootaxa, 4088 (4), 594 - 600. https: // doi. org / 10.11646 / zootaxa. 4088.4.10" type="journal article" year="2016">Hippa &amp; Vilkamaa, 2016</bibRefCitation>
</taxonomicName>
, C.
<emphasis id="B933C92BFFE52477FF4CD1828038FE6E" box="[151,274,403,429]" italics="true" pageId="4" pageNumber="141">subcorticis</emphasis>
(
<bibRefCitation id="EFD668C8FFE52477FEF9D1828345FE6E" author="Mohrig, W. &amp; Mamaev, B. &amp; Krivosheina, N." box="[290,623,402,429]" pageId="4" pageNumber="141" pagination="429 - 434" refId="ref2970" refString="Mohrig, W., Mamaev, B. &amp; Krivosheina, N. (1985). Beitrag zur Kenntnis der Trauermucken der Sowjetunion (Diptera, Sciaridae). Teil IX. Neue Arten aus der Tundra nordlich des Polarkreises. Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere, 112, 429 - 434." type="journal article" year="1985">Mohrig &amp; Krivosheina, 1985</bibRefCitation>
) and
<taxonomicName id="4C476EBAFFE52477FD6AD1828583FE6E" authority="Hippa, Vilkamaa &amp; Mohrig, 2003" authorityName="Hippa, Vilkamaa &amp; Mohrig" authorityYear="2003" box="[689,1193,403,429]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="B933C92BFFE52477FD6AD182820BFE6E" box="[689,801,403,429]" italics="true" pageId="4" pageNumber="141">C. tumida</emphasis>
<bibRefCitation id="EFD668C8FFE52477FCF2D1828583FE6E" author="Hippa, H. &amp; Vilkamaa, P. &amp; Mohrig, W." box="[809,1193,403,429]" pageId="4" pageNumber="141" pagination="469 - 511" refId="ref2778" refString="Hippa, H., Vilkamaa, P. &amp; Mohrig, W. (2003) Phylogeny of Corynoptera Winnertz and related genera, with the description of Claustropyga gen. nov. (Diptera, Sciaridae). Studia dipterologica, 9, 469 - 511. [2002]" type="journal article" year="2003">Hippa, Vilkamaa &amp; Mohrig, 2003</bibRefCitation>
</taxonomicName>
having a long tergite IX.
</paragraph>
<paragraph id="8BF81539FFE52477FF1CD1CA8032FDDA" blockId="4.[151,1437,151,537]" pageId="4" pageNumber="141">
<emphasis id="B933C92BFFE52477FF1CD1CA8015FE36" box="[199,319,475,501]" italics="true" pageId="4" pageNumber="141">Etymology</emphasis>
. The name is a Latin adjective,
<taxonomicName id="4C476EBAFFE52477FD1ED1CA823DFE36" authorityName="Vilkamaa" authorityYear="2024" box="[709,791,475,501]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="alveata">
<emphasis id="B933C92BFFE52477FD1ED1CA823DFE36" box="[709,791,475,501]" italics="true" pageId="4" pageNumber="141">alveata</emphasis>
</taxonomicName>
, hollowed, referring to the roundish, deeply excavated gonostylus.
</paragraph>
</subSubSection>
</treatment>
</document>