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<document id="21AE2668589332A6890903E4E48BACEC" ID-CLB-Dataset="31389" ID-DOI="10.11646/zootaxa.4083.3.7" ID-GBIF-Dataset="48173856-c4e6-4b27-bb6d-c23b1f8d01e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="1053910" ID-ZooBank="C999EBFD-4EAF-44E1-B7E9-95C9C63E556B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510926234376" checkinUser="plazi" docAuthor="Schmidt, Joachim, Hoffmann, Hannes &amp; Michalik, Peter" docDate="2016" docId="03C387EA101B3610FF2BDC47FC611B3D" docLanguage="en" docName="zootaxa.4083.3.7.pdf" docOrigin="Zootaxa 4083 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus eoanophthalmus Schmidt &amp; Hoffmann &amp; Michalik 2016, sp. n." docType="treatment" docVersion="7" lastPageNumber="439" masterDocId="FFFAFF92101A3618FFBCD872FF9E1E30" masterDocTitle="Blind life in the Baltic amber forests: description of an eyeless species of the ground beetle genus Trechus Clairville, 1806 (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="443" masterPageNumber="431" pageNumber="432" updateTime="1698500895377" updateUser="ExternalLinkService">
<mods:mods id="00F10AC1947E160DFA506EA07CF6EBC0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C2FF1C804CAA50D385F24B08321ABC7D">Blind life in the Baltic amber forests: description of an eyeless species of the ground beetle genus Trechus Clairville, 1806 (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="8A4948714140E62FCCAD4A140BF77F61">Schmidt, Joachim</mods:namePart>
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<mods:namePart id="954E58FE219631C538C40169ABC888F4">Hoffmann, Hannes</mods:namePart>
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<mods:namePart id="30DFAC07DBD3119A834890D0DB9993E8">Michalik, Peter</mods:namePart>
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<mods:title id="F1970B4B5020E94010ACCED818FE9767">Zootaxa</mods:title>
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<mods:date id="9DC666ABC7B298D6E60DB9E036CF8D8F">2016</mods:date>
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<mods:number id="3C49FD8D681D3CC355D34A1B6A29DF2E">2016-02-22</mods:number>
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<mods:number id="71473E9CFA5ED092328070EB9DB4060F">4083</mods:number>
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<mods:number id="4B6B50B5735BACA18A17137DC80B9496">3</mods:number>
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<mods:classification id="31795CF048D7D2B82653D16E9D6638A9">journal article</mods:classification>
<mods:identifier id="3C177DCD1947A7F5AB582A7EC48487BD" type="CLB-Dataset">31389</mods:identifier>
<mods:identifier id="EB2F6A24512162D1C770268ABF33DDF9" type="DOI">10.11646/zootaxa.4083.3.7</mods:identifier>
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<treatment id="03C387EA101B3610FF2BDC47FC611B3D" ID-DOI="http://doi.org/10.5281/zenodo.6067615" ID-GBIF-Taxon="137453387" ID-Zenodo-Dep="6067615" LSID="urn:lsid:plazi:treatment:03C387EA101B3610FF2BDC47FC611B3D" httpUri="http://treatment.plazi.org/id/03C387EA101B3610FF2BDC47FC611B3D" lastPageId="8" lastPageNumber="439" pageId="1" pageNumber="432">
<subSubSection id="C3706577101B3619FF2BDC47FE931A40" pageId="1" pageNumber="432" type="nomenclature">
<paragraph id="8BD536FC101B3619FF2BDC47FD8A1A7F" blockId="1.[151,532,1077,1136]" box="[151,532,1077,1103]" pageId="1" pageNumber="432">
<heading id="D09D8190101B3619FF2BDC47FD8A1A7F" bold="true" box="[151,532,1077,1103]" fontSize="11" level="1" pageId="1" pageNumber="432" reason="1">
<emphasis id="B91EEAEE101B3619FF2BDC47FD8A1A7F" bold="true" box="[151,532,1077,1103]" pageId="1" pageNumber="432">
<taxonomicName id="4C6A4D7F101B3619FF2BDC47FE541A7F" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[151,458,1077,1103]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE101B3619FF2BDC47FE541A7F" bold="true" box="[151,458,1077,1103]" italics="true" pageId="1" pageNumber="432">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D5795101B3619FE6EDC47FD8A1A7F" box="[466,532,1077,1103]" pageId="1" pageNumber="432" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD536FC101B3619FF2BDC25FE931A40" blockId="1.[151,532,1077,1136]" box="[151,269,1111,1136]" pageId="1" pageNumber="432">
<figureCitation id="13512A79101B3619FF2BDC25FE931A40" box="[151,269,1111,1136]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 4" captionStart-2="FIGURES 5 – 7" captionStart-3="FIGURES 8 – 12" captionStart-4="FIGURE 13" captionStartId-0="2.[151,264,1987,2010]" captionStartId-1="3.[151,264,1861,1884]" captionStartId-2="4.[151,264,1886,1909]" captionStartId-3="5.[151,264,1732,1755]" captionStartId-4="6.[151,250,1959,1981]" captionTargetBox-0="[151,1436,193,1966]" captionTargetBox-1="[179,1402,193,1839]" captionTargetBox-2="[151,1436,193,1866]" captionTargetBox-3="[151,1436,193,1711]" captionTargetBox-4="[251,1324,193,1935]" captionTargetId-0="figure@2.[151,1436,193,1966]" captionTargetId-1="figure@3.[179,1402,193,1839]" captionTargetId-2="figure@4.[151,1436,193,1866]" captionTargetId-3="figure@5.[151,1436,193,1711]" captionTargetId-4="figure@6.[251,1324,193,1935]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURES 1 – 2. Trechus eoanophthalmus sp. n., holotype. Fig. 1, left lateral aspect. Fig. 2, right lateral aspect." captionText-1="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." captionText-2="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." captionText-3="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." captionText-4="FIGURE 13. Trechus eoanophthalmus sp. n., reconstruction of the external shape in dorsal view. Since the labrum is only hardly visible in the fossil specimen, the assumed shape is indicated by a dotted line." httpUri-0="https://zenodo.org/record/1053912/files/figure.png" httpUri-1="https://zenodo.org/record/1053914/files/figure.png" httpUri-2="https://zenodo.org/record/1053916/files/figure.png" httpUri-3="https://zenodo.org/record/1053918/files/figure.png" httpUri-4="https://zenodo.org/record/1053920/files/figure.png" pageId="1" pageNumber="432">Figs. 1–13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3706577101B3619FF2BDCEDFABC192D" pageId="1" pageNumber="432" type="materials_examined">
<paragraph id="8BD536FC101B3619FF2BDCEDFCFA1B30" blockId="1.[151,1437,1183,2001]" pageId="1" pageNumber="432">
<typeStatus id="54D1885E101B3619FF2BDCEDFE9D1A88" box="[151,259,1183,1208]" pageId="1" pageNumber="432" type="holotype">Holotype</typeStatus>
. Male in Baltic amber; size of piece approximately 17.0 x 15.6 x 8.6 mm (irregularly cut,
<figureCitation id="13512A79101B3619FB57DCEDFAB31A88" box="[1259,1325,1183,1208]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="1" pageNumber="432">Fig. 7</figureCitation>
), No.
<quantity id="4C929B19101B3619FACEDCEDFF331AED" metricMagnitude="1" metricUnit="m" metricValue="1.1480799999999998" pageId="1" pageNumber="432" unit="in" value="452.0">452 in</quantity>
the collection F. Eichmann, Hannover,
<collectingCountry id="F37D766C101B3619FD06DCB6FCBC1AEC" box="[698,802,1220,1244]" name="Germany" pageId="1" pageNumber="432">Germany</collectingCountry>
(likely to be later deposited at the Geologisch- Paläontologisches Museum, University of
<collectingRegion id="49AEF81E101B3619FDC8DC95FD7E1B30" box="[628,736,1255,1280]" country="Germany" name="Hamburg" pageId="1" pageNumber="432">Hamburg</collectingRegion>
,
<collectingCountry id="F37D766C101B3619FD57DC9AFCC71B30" box="[747,857,1256,1280]" name="Germany" pageId="1" pageNumber="432">Germany</collectingCountry>
).
</paragraph>
<paragraph id="8BD536FC101B3619FF7BDD7EFAC51BE8" blockId="1.[151,1437,1183,2001]" pageId="1" pageNumber="432">
<emphasis id="B91EEAEE101B3619FF7BDD7EFE231B15" bold="true" box="[199,445,1292,1317]" pageId="1" pageNumber="432">Preservation status.</emphasis>
The specimen is markedly shrunken with its exoskeleton, which is completely dissociated from the inclusion wall. The latter represents the negative imprint of the beetle body and is visible in most portions. Detached parts of the original beetle exoskeleton, e.g., the distorted right elytron, which moves freely in the cavity of the amber piece, and the shrunken legs, are visible under optical microscope and could be imaged in detail using micro-CT (
<figureCitation id="13512A79101B3619FDA8DDEEFD191B84" box="[532,647,1436,1461]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="1" pageNumber="432">Figs 8–11</figureCitation>
). This particular condition of preservation is probably the result of the embedding of an immature and thus teneral specimen whose exoskeleton collapsed after the resin was cured.
</paragraph>
<paragraph id="8BD536FC101B3619FF7BDD97FE5918C8" blockId="1.[151,1437,1183,2001]" pageId="1" pageNumber="432">
Most parts of the negative imprint of the beetle body were visible in detail using a light microscope (see, e.g., microsculpture of head,
<figureCitation id="13512A79101B3619FE17DE75FE6E1810" box="[427,496,1543,1568]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1861,1884]" captionTargetBox="[179,1402,193,1839]" captionTargetId="figure@3.[179,1402,193,1839]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="1" pageNumber="432">Fig. 3</figureCitation>
, protibial groove,
<figureCitation id="13512A79101B3619FD7EDE75FC981810" box="[706,774,1543,1568]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="1" pageNumber="432">Fig. 6</figureCitation>
). However, since the amber piece was polished only from two sides lateral to the inclusion (
<figureCitation id="13512A79101B3619FDB1DE5EFDD11874" box="[525,591,1580,1605]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="1" pageNumber="432">Fig. 7</figureCitation>
) a view directly from dorsad and ventrad of the beetle is not possible due to refractions. The mouth parts are surrounded by dirt particles and therefore, the basal portion of mandibles, maxillae and mentum could not be examined using light microscopy (only the apical part of mandibles and apical segments of the palpi were visible,
<figureCitation id="13512A79101B3619FE7FDEE5FD8A1880" box="[451,532,1687,1712]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1861,1884]" captionTargetBox="[179,1402,193,1839]" captionTargetId="figure@3.[179,1402,193,1839]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="1" pageNumber="432">Figs. 3</figureCitation>
). In addition, the results of micro-CT imaging showed that the imprint of the inclusion was destroyed on the ventral surface of head (
<figureCitation id="13512A79101B3619FCB9DECEFCCA18E4" box="[773,852,1724,1749]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="1" pageNumber="432">Fig. 11</figureCitation>
) and therefore, diagnostic characters of the mentum could not be reconstructed.
</paragraph>
<paragraph id="8BD536FC101B3619FF7BDF76FABC192D" blockId="1.[151,1437,1183,2001]" box="[199,1314,1796,1821]" pageId="1" pageNumber="432">
<emphasis id="B91EEAEE101B3619FF7BDF76FEEF192D" bold="true" box="[199,369,1796,1821]" pageId="1" pageNumber="432">Syninclusions.</emphasis>
Apart from several dirt particles, no syninclusions are preserved in the amber piece.
</paragraph>
</subSubSection>
<subSubSection id="C3706577101B361FFF7BDF55FB671860" lastPageId="7" lastPageNumber="438" pageId="1" pageNumber="432" type="description">
<paragraph id="8BD536FC101B3619FF7BDF55FDCB1970" blockId="1.[151,1437,1183,2001]" box="[199,597,1831,1856]" pageId="1" pageNumber="432">
<emphasis id="B91EEAEE101B3619FF7BDF55FEC71970" bold="true" box="[199,345,1831,1856]" pageId="1" pageNumber="432">Description.</emphasis>
Body length: 4.4 mm.
</paragraph>
<paragraph id="8BD536FC101B3619FF7BDF3EFD7D1955" blockId="1.[151,1437,1183,2001]" box="[199,739,1868,1893]" pageId="1" pageNumber="432">Colour: Colour of body surface undeterminable.</paragraph>
<paragraph id="8BD536FC101B3619FF7BDF1DFB2719E1" blockId="1.[151,1437,1183,2001]" pageId="1" pageNumber="432">
Microsculpture: Clypeus and frons with slightly engraved markedly transverse meshes, neck with larger and less transverse meshes (
<figureCitation id="13512A79101B3619FE11DFE6FE6B199C" box="[429,501,1940,1965]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1861,1884]" captionTargetBox="[179,1402,193,1839]" captionTargetId="figure@3.[179,1402,193,1839]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="1" pageNumber="432">Fig. 3</figureCitation>
). Discs of pronotum and elytra with very fine and weakly impressed transverse meshes (visible under magnification&gt; 150 x), appearing polished under lower magnifications.
</paragraph>
<caption id="DF1566741018361AFF2BDFB1FB6D19EA" box="[151,1267,1987,2010]" httpUri="https://zenodo.org/record/1053912/files/figure.png" pageId="2" pageNumber="433" targetBox="[151,1436,193,1966]" targetPageId="2">
<paragraph id="8BD536FC1018361AFF2BDFB1FB6D19EA" blockId="2.[151,1267,1987,2010]" box="[151,1267,1987,2010]" pageId="2" pageNumber="433">
<emphasis id="B91EEAEE1018361AFF2BDFB1FEA419E9" bold="true" box="[151,314,1987,2010]" pageId="2" pageNumber="433">FIGURES 1–2.</emphasis>
<taxonomicName id="4C6A4D7F1018361AFEFCDFB6FDA319EA" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[320,573,1988,2010]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE1018361AFEFCDFB6FDA319EA" box="[320,573,1988,2010]" italics="true" pageId="2" pageNumber="433">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57951018361AFDF8DFB6FDE819E9" box="[580,630,1988,2009]" pageId="2" pageNumber="433" rank="species">
<emphasis id="B91EEAEE1018361AFDF8DFB6FDE819E9" bold="true" box="[580,630,1988,2009]" pageId="2" pageNumber="433">sp.n.</emphasis>
</taxonomicNameLabel>
, holotype. Fig. 1, left lateral aspect. Fig. 2, right lateral aspect.
</paragraph>
</caption>
<caption id="DF1566741019361BFF2BDF37FDFE19E6" httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="3" pageNumber="434" targetBox="[179,1402,193,1839]" targetPageId="3">
<paragraph id="8BD536FC1019361BFF2BDF37FDFE19E6" blockId="3.[151,1436,1861,2006]" pageId="3" pageNumber="434">
<emphasis id="B91EEAEE1019361BFF2BDF37FEA5196B" bold="true" box="[151,315,1861,1884]" pageId="3" pageNumber="434">FIGURES 3–4.</emphasis>
<taxonomicName id="4C6A4D7F1019361BFEFDDF34FDA0196C" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[321,574,1862,1884]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE1019361BFEFDDF34FDA0196C" box="[321,574,1862,1884]" italics="true" pageId="3" pageNumber="434">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57951019361BFDF9DF34FDE6196B" box="[581,632,1862,1883]" pageId="3" pageNumber="434" rank="species">
<emphasis id="B91EEAEE1019361BFDF9DF34FDE6196B" bold="true" box="[581,632,1862,1883]" pageId="3" pageNumber="434">sp.n.</emphasis>
</taxonomicNameLabel>
, holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae.
</paragraph>
</caption>
<caption id="DF156674101E361CFF2BDF2CFD181982" httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="4" pageNumber="435" targetBox="[151,1436,193,1866]" targetPageId="4">
<paragraph id="8BD536FC101E361CFF2BDF2CFD181982" blockId="4.[151,1436,1886,1970]" pageId="4" pageNumber="435">
<emphasis id="B91EEAEE101E361CFF2BDF2CFEA51944" bold="true" box="[151,315,1886,1909]" pageId="4" pageNumber="435">FIGURES 5–7.</emphasis>
<taxonomicName id="4C6A4D7F101E361CFEFFDF2DFDDE1945" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[323,576,1887,1909]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE101E361CFEFFDF2DFDDE1945" box="[323,576,1887,1909]" italics="true" pageId="4" pageNumber="435">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D5795101E361CFDF4DF2DFDE51944" box="[584,635,1887,1908]" pageId="4" pageNumber="435" rank="species">
<emphasis id="B91EEAEE101E361CFDF4DF2DFDE51944" bold="true" box="[584,635,1887,1908]" pageId="4" pageNumber="435">sp.n.</emphasis>
</taxonomicNameLabel>
, holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg= protibial dorsolateral groove.
</paragraph>
</caption>
<caption id="DF156674101F361DFF2BDEB6FD1219A2" httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="5" pageNumber="436" targetBox="[151,1436,193,1711]" targetPageId="5">
<paragraph id="8BD536FC101F361DFF2BDEB6FD1219A2" blockId="5.[151,1436,1732,1938]" pageId="5" pageNumber="436">
<emphasis id="B91EEAEE101F361DFF2BDEB6FED818EA" bold="true" box="[151,326,1732,1755]" pageId="5" pageNumber="436">FIGURES 8–12.</emphasis>
<taxonomicName id="4C6A4D7F101F361DFEF0DEB6FDD618EA" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[332,584,1732,1754]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE101F361DFEF0DEB6FDD618EA" box="[332,584,1732,1754]" italics="true" pageId="5" pageNumber="436">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D5795101F361DFDF2DEB7FD1E18EA" box="[590,640,1733,1754]" pageId="5" pageNumber="436" rank="species">
<emphasis id="B91EEAEE101F361DFDF2DEB7FD1E18EA" bold="true" box="[590,640,1733,1754]" pageId="5" pageNumber="436">sp.n.</emphasis>
</taxonomicNameLabel>
, holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria.
</paragraph>
</caption>
<caption id="DF156674101C361EFF2BDFD5FC6F19EB" httpUri="https://zenodo.org/record/1053920/files/figure.png" pageId="6" pageNumber="437" targetBox="[251,1324,193,1935]" targetPageId="6">
<paragraph id="8BD536FC101C361EFF2BDFD5FC6F19EB" blockId="6.[151,1436,1959,2011]" pageId="6" pageNumber="437">
<emphasis id="B91EEAEE101C361EFF2BDFD5FEBC198C" bold="true" box="[151,290,1959,1981]" pageId="6" pageNumber="437">FIGURE 13.</emphasis>
<taxonomicName id="4C6A4D7F101C361EFE97DFD5FDB4198D" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[299,554,1959,1981]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="437" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE101C361EFE97DFD5FDB4198D" box="[299,554,1959,1981]" italics="true" pageId="6" pageNumber="437">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D5795101C361EFD8FDFD5FDF0198C" box="[563,622,1959,1980]" pageId="6" pageNumber="437" rank="species">
<emphasis id="B91EEAEE101C361EFD8FDFD5FDF0198C" bold="true" box="[563,622,1959,1980]" pageId="6" pageNumber="437">sp. n.</emphasis>
</taxonomicNameLabel>
, reconstruction of the external shape in dorsal view. Since the labrum is only hardly visible in the fossil specimen, the assumed shape is indicated by a dotted line.
</paragraph>
</caption>
<paragraph id="8BD536FC101D361FFF7BD8E5FB531FE0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="438">
Head: Length 1.09 mm, short and robust, notably broad near base (
<figureCitation id="13512A79101D361FFC6FD8E5FB841E80" box="[979,1050,151,176]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">Fig. 8</figureCitation>
). Mandibles of normal length for
<taxonomicName id="4C6A4D7F101D361FFF2BD8CEFF711EE5" authorityName="Clairville" authorityYear="1806" box="[151,239,188,213]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE101D361FFF2BD8CEFF711EE5" box="[151,239,188,213]" italics="true" pageId="7" pageNumber="438">Trechus</emphasis>
</taxonomicName>
(mandibular dentition and chaetotaxy of the mandibular groove could not be reconstructed). Labrum with three pairs of setae (shape of the labral apical margin could not be displayed). Clypeus with two pairs of setae in normal position. Shape of apical segments of maxillary and labial palpi as in
<taxonomicName id="4C6A4D7F101D361FFBEBD976FB311F2D" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[1111,1199,260,285]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91EEAEE101D361FFBEBD976FB311F2D" box="[1111,1199,260,285]" italics="true" pageId="7" pageNumber="438">Trechus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D5795101D361FFB01D977FACD1F2D" box="[1213,1363,260,285]" pageId="7" pageNumber="438" sensu="stricto">sensu stricto</taxonomicNameLabel>
. Eyes completely reduced (
<figureCitation id="13512A79101D361FFE3BD955FE701F70" box="[391,494,295,320]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 4" captionStartId-0="2.[151,264,1987,2010]" captionStartId-1="3.[151,264,1861,1884]" captionTargetBox-0="[151,1436,193,1966]" captionTargetBox-1="[179,1402,193,1839]" captionTargetId-0="figure@2.[151,1436,193,1966]" captionTargetId-1="figure@3.[179,1402,193,1839]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 2. Trechus eoanophthalmus sp. n., holotype. Fig. 1, left lateral aspect. Fig. 2, right lateral aspect." captionText-1="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri-0="https://zenodo.org/record/1053912/files/figure.png" httpUri-1="https://zenodo.org/record/1053914/files/figure.png" pageId="7" pageNumber="438">Figs 1–4</figureCitation>
,
<figureCitation id="13512A79101D361FFE42D95AFD941F70" box="[510,522,296,320]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">8</figureCitation>
).Tempora, frons, orbital and supraorbital areas smooth, markedly convex, with supraorbital furrows deep, complete, and almost uniformly bent on disc (
<figureCitation id="13512A79101D361FFC72D93EFBAB1F54" box="[974,1077,332,357]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1861,1884]" captionTargetBox="[179,1402,193,1839]" captionTargetId="figure@3.[179,1402,193,1839]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="7" pageNumber="438">Figs 3–4</figureCitation>
); frons with a slight but distinct transverse impression (
<figureCitation id="13512A79101D361FFE1CD91DFE721FB8" box="[416,492,367,392]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1861,1884]" captionTargetBox="[179,1402,193,1839]" captionTargetId="figure@3.[179,1402,193,1839]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="7" pageNumber="438">Figs 3</figureCitation>
,
<figureCitation id="13512A79101D361FFE41D902FD921FB8" box="[509,524,368,392]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">8</figureCitation>
); two supraorbital setae present and in normal position for
<taxonomicName id="4C6A4D7F101D361FFB7BD91DFA801FB8" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[1223,1310,367,392]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE101D361FFB7BD91DFA801FB8" box="[1223,1310,367,392]" italics="true" pageId="7" pageNumber="438">Trechus</emphasis>
</taxonomicName>
. Antennae moderately short, with scapus rather robust (length 0.17 mm; length/width = 1.31; 1.3times broader than pedicellus), and with pedicellus and third antennomere of the same length (0.16 mm) and shape.
</paragraph>
<paragraph id="8BD536FC101D361FFF7BD9AEFF701DB0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="438">
Prothorax: Pronotum markedly transverse (width/length = 1.54), length 0.82 mm, 1.39 times broader than head, broadest in middle, with sides evenly rounded in anterior 5/6 and straight before the small but well developed laterobasal angles; basal margin 1.25 times broader than apical margin. Disk markedly convex, smooth (
<figureCitation id="13512A79101D361FFAFBDA56FA111C0C" box="[1351,1423,548,573]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1861,1884]" captionTargetBox="[179,1402,193,1839]" captionTargetId="figure@3.[179,1402,193,1839]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Trechus eoanophthalmus sp. n., holotype. Fig. 3, head (view at an angle from right above), showing markedly transverse micro-meshes on frons, and moderately transverse meshes on neck. For better visualization, the outline of the right mandible is indicated by a dotted line. Fig. 4, head and pronotum, left lateral aspect. Abbreviations: fti = frontal transverse impression; lbp = labial palpus; md = mandible; mx = maxilla; mxp = maxillary palpus; pbs = pronotal basolateral seta; pls = pronotal lateral seta; sos = supraorbital setae." httpUri="https://zenodo.org/record/1053914/files/figure.png" pageId="7" pageNumber="438">Fig. 4</figureCitation>
). Anterior margin almost straight, very finely beaded in outer third, smooth in middle, with apical angles very slightly protruded, rounded. Posterior margin not beaded, moderately convex in middle, slightly incised before outer quarters; laterobasal angles small, not shifted anteriad, almost rectangular, not protruded laterally. Median longitudinal impression distinct, somewhat deeper near base, disappearing near apex; anterior transverse impression very fine, smooth; posterior transverse impression deep and linear, smooth, laterobasal foveae indistinct; pronotal base convex between laterobasal foveae and side margin. Lateral gutter moderately narrow throughout, very slightly widened towards base, smooth. Both lateral and laterobasal setae present, with the lateral seta located at apical third of pronotum. Proepisternum and prosternal process glabrous and smooth, latter without margin.
</paragraph>
<paragraph id="8BD536FC101D361FFF7BDBFEFED21B48" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="438">
Pterothorax: Elytra markedly convex on disc (
<figureCitation id="13512A79101D361FFD6ADBFEFCDD1D94" box="[726,835,908,933]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1987,2010]" captionTargetBox="[151,1436,193,1966]" captionTargetId="figure@2.[151,1436,193,1966]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Trechus eoanophthalmus sp. n., holotype. Fig. 1, left lateral aspect. Fig. 2, right lateral aspect." httpUri="https://zenodo.org/record/1053912/files/figure.png" pageId="7" pageNumber="438">Figs. 1–2</figureCitation>
,
<figureCitation id="13512A79101D361FFCECDBFEFCF11D94" box="[848,879,908,932]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">10</figureCitation>
), in dorsal view short-ovate (
<figureCitation id="13512A79101D361FFB03DBFEFA9B1D94" box="[1215,1285,908,933]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">Fig. 9</figureCitation>
), length 2.49 mm, length/width = 1.21, widest near their mid-length, moderately wider than pronotum (width of elytra/width of pronotum = 1.20), glabrous beside normal setation. Humeral angles fully rounded, basal groove absent. Parascutellar stria and parascutellar seta present. First stria complete, second stria disappearing near apex, third and fourth striae distinctly finer impressed than inner striae, disappearing near base and apex, fifth and sixth striae indistinct, seventh and eighth striae absent; inner striae finely punctated apart from the apical fifth of elytra (
<figureCitation id="13512A79101D361FFAD8DC4DFF391A4D" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="7" pageNumber="438">Figs. 5</figureCitation>
,
<figureCitation id="13512A79101D361FFF08DC16FF4C1A4C" box="[180,210,1124,1148]" captionStart="FIGURE 13" captionStartId="6.[151,250,1959,1981]" captionTargetBox="[251,1324,193,1935]" captionTargetId="figure@6.[251,1324,193,1935]" captionTargetPageId="6" captionText="FIGURE 13. Trechus eoanophthalmus sp. n., reconstruction of the external shape in dorsal view. Since the labrum is only hardly visible in the fossil specimen, the assumed shape is indicated by a dotted line." httpUri="https://zenodo.org/record/1053920/files/figure.png" pageId="7" pageNumber="438">13</figureCitation>
). Each elytron with two discal setae in third stria, and with preapical seta, situated at apical eighth of elytra, almost equally as close to the apical margin as to the suture of elytra (
<figureCitation id="13512A79101D361FFC20DCF5FC7B1A90" box="[924,997,1159,1184]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="7" pageNumber="438">Figs 5</figureCitation>
,
<figureCitation id="13512A79101D361FFC4CDCFAFB911A90" box="[1008,1039,1160,1184]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">12</figureCitation>
). Recurrent stria extending towards the reduced fifth stria anteriorly, moderately long, almost straight, with front end situated slightly anterior of the level of the preapical seta (
<figureCitation id="13512A79101D361FFE77DCBDFDBE1AD8" box="[459,544,1231,1256]" captionStart="FIGURES 8 – 12" captionStartId="5.[151,264,1732,1755]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@5.[151,1436,193,1711]" captionTargetPageId="5" captionText="FIGURES 8 – 12. Trechus eoanophthalmus sp. n., holotype; MicroCT images. Fig. 8, negative imprint of head, dorsal view. Fig. 9, negative imprint of elytra and pronotum, dorsal view, and deformed right elytron and abdominal tergites in the cavity of the inclusion. Fig. 10, negative imprint of the inclusion with deformed remains of the beetle body in the cavity of the inclusion, left lateral view. Fig. 11, same, ventral view. Fig. 12, apical portion of elytra (negative imprint), caudal view (the insertions of the setae of the apical triangle are marked by white circles). Abbreviations: as-p = abdominal sternites (deformed positive); at-p = abdominal tergites (deformed positive); cl-p = clypeus (deformed positive); el-p = right elytron (deformed positive); fti = frontal transverse impression; rs = recurrent stria." httpUri="https://zenodo.org/record/1053918/files/figure.png" pageId="7" pageNumber="438">Fig. 12</figureCitation>
). Setae of umbilicate humeral series close to the elytral margin, with posterior three setae equidistant from each other, and with first seta slightly more distant from second (
<figureCitation id="13512A79101D361FFB0CDC86FB611B3D" box="[1200,1279,1268,1293]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="7" pageNumber="438">Figs. 5</figureCitation>
,
<figureCitation id="13512A79101D361FFAB0DC86FAB41B3C" box="[1292,1322,1268,1292]" captionStart="FIGURE 13" captionStartId="6.[151,250,1959,1981]" captionTargetBox="[251,1324,193,1935]" captionTargetId="figure@6.[251,1324,193,1935]" captionTargetPageId="6" captionText="FIGURE 13. Trechus eoanophthalmus sp. n., reconstruction of the external shape in dorsal view. Since the labrum is only hardly visible in the fossil specimen, the assumed shape is indicated by a dotted line." httpUri="https://zenodo.org/record/1053920/files/figure.png" pageId="7" pageNumber="438">13</figureCitation>
). The two setae of the medial group of the umbilicate series far removed from the preapical group (
<figureCitation id="13512A79101D361FFB26DD65FB751B00" box="[1178,1259,1303,1328]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="7" pageNumber="438">Figs. 5</figureCitation>
,
<figureCitation id="13512A79101D361FFB46DD6AFA861B00" box="[1274,1304,1304,1328]" captionStart="FIGURE 13" captionStartId="6.[151,250,1959,1981]" captionTargetBox="[251,1324,193,1935]" captionTargetId="figure@6.[251,1324,193,1935]" captionTargetPageId="6" captionText="FIGURE 13. Trechus eoanophthalmus sp. n., reconstruction of the external shape in dorsal view. Since the labrum is only hardly visible in the fossil specimen, the assumed shape is indicated by a dotted line." httpUri="https://zenodo.org/record/1053920/files/figure.png" pageId="7" pageNumber="438">13</figureCitation>
). Epipleura long, nearly reaching the apex. Metepisternum short, glabrous and smooth, with outer margin 1.2 times longer than anterior margin.
</paragraph>
<paragraph id="8BD536FC101D361FFF7BDDF6FD561BF0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="438">Abdomen: Abdominal sternites V–VII and anal sternite each with one (male) pair of setae near apical margin; surfaces smooth, without hairs or micropunctures.</paragraph>
<paragraph id="8BD536FC101D361FFF7BDDBFFD3F181D" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="438">
Legs: Moderately short, robust, all femora unmodified; protibia straight and moderately dilated towards apex, glabrous on their anterior surface, with a short and very fine longitudinal groove on external surface (
<figureCitation id="13512A79101D361FFAB8DD9DFAD61838" box="[1284,1352,1519,1544]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="7" pageNumber="438">Fig. 6</figureCitation>
). Basal two protarsomeres moderately dilated (
<figureCitation id="13512A79101D361FFDF2DE66FD0D181D" box="[590,659,1556,1581]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[151,1436,193,1866]" captionTargetId="figure@4.[151,1436,193,1866]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Trechus eoanophthalmus sp. n., holotype. Fig. 5, left elytron (the insertions of the setae are marked by white circles). Fig. 6, left foreleg and middle leg, dorsal view. Fig. 7, general view of the fossil with contours of the amber piece. Abbreviation: ptg = protibial dorsolateral groove." httpUri="https://zenodo.org/record/1053916/files/figure.png" pageId="7" pageNumber="438">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BD536FC101D361FFF7BDE45FB671860" blockId="7.[151,1437,151,2013]" box="[199,1273,1591,1616]" pageId="7" pageNumber="438">Male genitalia: Could not be located within the collapsed abdominal portion of the exoskeleton.</paragraph>
</subSubSection>
<subSubSection id="C3706577101D3610FF7BDE2EFB801EC8" lastPageId="8" lastPageNumber="439" pageId="7" pageNumber="438" type="etymology">
<paragraph id="8BD536FC101D361FFF7BDE2EFD8C188D" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="438">
<emphasis id="B91EEAEE101D361FFF7BDE2EFED01845" bold="true" box="[199,334,1628,1653]" pageId="7" pageNumber="438">Etymology.</emphasis>
The name of the new species is a combination of “eo“, derived from the Eocene epoch when the beetle was fossilized, and “anophthalmus” (= eyeless), derived from a combination of the Greek words “a(n)” = without, and “ophthalmos” = eye.
</paragraph>
<paragraph id="8BD536FC101D3610FF7BDEB5FB801EC8" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,1293]" lastPageId="8" lastPageNumber="439" pageId="7" pageNumber="438">
<emphasis id="B91EEAEE101D361FFF7BDEB5FEEE18D0" bold="true" box="[199,368,1735,1760]" pageId="7" pageNumber="438">Relationships.</emphasis>
<taxonomicName id="4C6A4D7F101D361FFEC4DEB5FD0E18D0" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[376,656,1735,1760]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE101D361FFEC4DEB5FD0E18D0" box="[376,656,1735,1760]" italics="true" pageId="7" pageNumber="438">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D5795101D361FFD25DEB5FD4418D0" box="[665,730,1735,1760]" pageId="7" pageNumber="438" rank="species">
<emphasis id="B91EEAEE101D361FFD25DEB5FD4418D0" bold="true" box="[665,730,1735,1760]" pageId="7" pageNumber="438">sp. n.</emphasis>
</taxonomicNameLabel>
shares all diagnostic features with the previously described
<taxonomicName id="4C6A4D7F101D361FFA3BDEBBFF6E1935" authorityName="Schmidt &amp; Faille" authorityYear="2015" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="B91EEAEE101D361FFA3BDEBBFF6E1935" italics="true" pageId="7" pageNumber="438">T. balticus</emphasis>
</taxonomicName>
placing the new fossil species either in the main
<taxonomicName id="4C6A4D7F101D361FFCA5DE9EFCEF1935" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[793,881,1772,1797]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE101D361FFCA5DE9EFCEF1935" box="[793,881,1772,1797]" italics="true" pageId="7" pageNumber="438">Trechus</emphasis>
</taxonomicName>
clade sensu
<bibRefCitation id="EFFB4B0D101D361FFC43DE9EFB4F1934" author="Faille, A. &amp; Casale, A. &amp; Balke, M. &amp; Ribera, I." box="[1023,1233,1772,1797]" journalOrPublisher="BMC Evolutionary Biology" pageId="7" pageNumber="438" pagination="248" part="13" publicationUrl="http://dx.doi.org/10.1186/1471-2148-13-248" refString="Faille, A., Casale, A., Balke, M. &amp; Ribera, I. (2013) A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae). BMC Evolutionary Biology, 13, 248. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 13 - 248" title="A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae)" type="journal article" year="2013">
Faille
<emphasis id="B91EEAEE101D361FFBFADE9FFBE11935" box="[1094,1151,1772,1797]" italics="true" pageId="7" pageNumber="438">et al.</emphasis>
(2013)
</bibRefCitation>
or in the Nearctic lineage
<taxonomicName id="4C6A4D7F101D361FFF53DF7DFDBB1918" authority="Jeannel, 1927" authorityName="Jeannel" authorityYear="1927" box="[239,549,1807,1832]" class="Insecta" family="Carabidae" genus="Microtrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE101D361FFF53DF7DFE1A1918" box="[239,388,1807,1832]" italics="true" pageId="7" pageNumber="438">Microtrechus</emphasis>
<bibRefCitation id="EFFB4B0D101D361FFE37DF7DFDBB1918" author="Jeannel, R." box="[395,549,1807,1832]" journalOrPublisher="L'Abeille" pageId="7" pageNumber="438" pagination="1 - 592" part="33" refString="Jeannel, R. (1927) Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Deuxieme livraison). L'Abeille, 33, 1 - 592." title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Deuxieme livraison)" type="journal article" year="1927">Jeannel, 1927</bibRefCitation>
</taxonomicName>
(see discussion in
<bibRefCitation id="EFFB4B0D101D361FFD47DF7DFB9F1918" author="Schmidt, J. &amp; Faille, A." box="[763,1025,1807,1832]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="438" pagination="539 - 550" part="4028" publicationUrl="http://dx.doi.org/10.11646/zootaxa.4028.4.5" refString="Schmidt, J. &amp; Faille, A. (2015) Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini). Zootaxa, 4028 (4), 539 - 550. http: // dx. doi. org / 10.11646 / zootaxa. 4028.4.5" title="Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini)" type="journal article" year="2015">Schmidt &amp; Faille 2015</bibRefCitation>
): shape of mandibles normal; orbital area glabrous, with deep furrows semicircularly separating well developed supraorbital region from frons and neck; two supraorbital setae present; apical segment of maxillary palpus as long as the penultimate, at base almost as broad as apex of penultimate segment, and at apex regularly conically pointed; pronotum and elytra glabrous; pronotum with posterior transverse impression and laterobasal foveae linear and sharply limited towards the convex disc, with one anterolateral and one laterobasal setae each side; elytra with basal groove absent, and with two discal setae; recurrent sutural stria forming a deep carina directed towards the end of the fifth stria; apical triangle present, setae of humeral group situated close to the elytral margin, setae of medial group well separated from the preapical group; apical portion of protibia glabrous on external surface.
</paragraph>
</subSubSection>
<subSubSection id="C370657710123610FF7BD976FCE01DDD" pageId="8" pageNumber="439" type="reference_group">
<paragraph id="8BD536FC10123610FF7BD976FE361CDF" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="439">
<taxonomicName id="4C6A4D7F10123610FF7BD976FE7E1F2D" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[199,480,260,285]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FF7BD976FE7E1F2D" box="[199,480,260,285]" italics="true" pageId="8" pageNumber="439">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FE54D976FDB71F2D" box="[488,553,260,285]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FE54D976FDB71F2D" bold="true" box="[488,553,260,285]" pageId="8" pageNumber="439">sp. n.</emphasis>
</taxonomicNameLabel>
is distinguished from
<taxonomicName id="4C6A4D7F10123610FC94D976FC231F2D" authorityName="Jeannel" authorityYear="1927" box="[808,957,260,285]" class="Insecta" family="Carabidae" genus="Microtrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE10123610FC94D976FC231F2D" box="[808,957,260,285]" italics="true" pageId="8" pageNumber="439">Microtrechus</emphasis>
</taxonomicName>
by two dilated male protarsomeres. Thus, the new species is most likely a representative of the main
<taxonomicName id="4C6A4D7F10123610FCEED955FC341F70" ID-CoL="8KZB9" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[850,938,295,320]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE10123610FCEED955FC341F70" box="[850,938,295,320]" italics="true" pageId="8" pageNumber="439">Trechus</emphasis>
</taxonomicName>
clade sensu
<bibRefCitation id="EFFB4B0D10123610FBFED955FABF1F70" author="Faille, A. &amp; Casale, A. &amp; Balke, M. &amp; Ribera, I." box="[1090,1313,295,320]" journalOrPublisher="BMC Evolutionary Biology" pageId="8" pageNumber="439" pagination="248" part="13" publicationUrl="http://dx.doi.org/10.1186/1471-2148-13-248" refString="Faille, A., Casale, A., Balke, M. &amp; Ribera, I. (2013) A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae). BMC Evolutionary Biology, 13, 248. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 13 - 248" title="A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae)" type="journal article" year="2013">
Faille
<emphasis id="B91EEAEE10123610FB31D95BFB571F70" box="[1165,1225,295,320]" italics="true" pageId="8" pageNumber="439">et al.</emphasis>
(2013)
</bibRefCitation>
. However, phylogenetic studies have shown that the genus
<taxonomicName id="4C6A4D7F10123610FD7DD93EFC871F55" ID-CoL="8KZB9" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[705,793,332,357]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE10123610FD7DD93EFC871F55" box="[705,793,332,357]" italics="true" pageId="8" pageNumber="439">Trechus</emphasis>
</taxonomicName>
sensu auctorum is a non-monophyletic assemblage (see
<bibRefCitation id="EFFB4B0D10123610FF2BD91DFEC71FB8" author="Faille, A. &amp; Ribera, I &amp; Deharveng, L. &amp; Bourdeau, C. &amp; Garnery, L. &amp; Queinnec, E. &amp; Deuve, T." box="[151,345,367,392]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="8" pageNumber="439" pagination="97 - 105" part="54" publicationUrl="http://dx.doi.org/10.1016/j.ympev.2009.10.008" refString="Faille, A., Ribera, I., Deharveng, L., Bourdeau, C., Garnery, L., Queinnec, E. &amp; Deuve, T. (2010) A molecular phylogeny shows the single origin of the Pyrenean subterranean Trechini ground beetles (Coleoptera: Carabidae). Molecular Phylogenetics and Evolution, 54, 97 - 105. http: // dx. doi. org / 10.1016 / j. ympev. 2009.10.008" title="A molecular phylogeny shows the single origin of the Pyrenean subterranean Trechini ground beetles (Coleoptera: Carabidae)" type="journal article" year="2010">
Faille
<emphasis id="B91EEAEE10123610FF62D903FE891FB8" box="[222,279,367,392]" italics="true" pageId="8" pageNumber="439">et al.</emphasis>
2010
</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FED9D902FE001FB8" author="Faille, A. &amp; Casale, A. &amp; Ribera, I." box="[357,414,368,392]" journalOrPublisher="Zoologica Scripta 40" pageId="8" pageNumber="439" pagination="282 - 295" part="3" publicationUrl="http://dx.doi.org/10.1111/j.1463-6409.2010.00467.x" refString="Faille, A., Casale, A. &amp; Ribera, I. (2011) Phylogenetic relationships of west Mediterranean troglobitic Trechini ground beetles (Coleoptera: Carabidae). Zoologica Scripta 40 (3), 282 - 295. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.2010.00467. x" title="Phylogenetic relationships of west Mediterranean troglobitic Trechini ground beetles (Coleoptera: Carabidae)" type="journal article" year="2011">2011</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FE16D902FE791FB8" author="Faille, A. &amp; Casale, A. &amp; Balke, M. &amp; Ribera, I." box="[426,487,368,392]" journalOrPublisher="BMC Evolutionary Biology" pageId="8" pageNumber="439" pagination="248" part="13" publicationUrl="http://dx.doi.org/10.1186/1471-2148-13-248" refString="Faille, A., Casale, A., Balke, M. &amp; Ribera, I. (2013) A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae). BMC Evolutionary Biology, 13, 248. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 13 - 248" title="A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae)" type="journal article" year="2013">2013</bibRefCitation>
). In addition, the phylogenetic positions of many of the non-European lineages need still to be investigated. Like in the previously described Baltic amber fossil
<taxonomicName id="4C6A4D7F10123610FC56D9E7FBC11F9D" ID-CoL="7CNYX" authorityName="Schmidt &amp; Faille" authorityYear="2015" box="[1002,1119,404,429]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="B91EEAEE10123610FC56D9E7FBC11F9D" box="[1002,1119,404,429]" italics="true" pageId="8" pageNumber="439">T. balticus</emphasis>
</taxonomicName>
, phylogenetically important genital characters could not be revealed in
<taxonomicName id="4C6A4D7F10123610FD39D9CBFCC21FE0" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[645,860,439,464]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FD39D9CBFCC21FE0" box="[645,860,439,464]" italics="true" pageId="8" pageNumber="439">T. eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FCD9D9C5FC381FE0" box="[869,934,439,464]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FCD9D9C5FC381FE0" bold="true" box="[869,934,439,464]" pageId="8" pageNumber="439">sp. n.</emphasis>
</taxonomicNameLabel>
due to the poor preservation of the internal parts of its body. Consequently, the systematic position of
<taxonomicName id="4C6A4D7F10123610FC94D9AFFC631FC5" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[808,1021,476,501]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FC94D9AFFC631FC5" box="[808,1021,476,501]" italics="true" pageId="8" pageNumber="439">T. eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FBB8D9AEFBA21FC5" box="[1028,1084,476,501]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FBB8D9AEFBA21FC5" bold="true" box="[1028,1084,476,501]" pageId="8" pageNumber="439">sp.n.</emphasis>
</taxonomicNameLabel>
within the mega-diverse genus
<taxonomicName id="4C6A4D7F10123610FF2BD98DFF711C28" ID-CoL="8KZB9" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[151,239,511,536]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE10123610FF2BD98DFF711C28" box="[151,239,511,536]" italics="true" pageId="8" pageNumber="439">Trechus</emphasis>
</taxonomicName>
remains dubious. Nevertheless, it seems very likely that the Eocene fossil species,
<taxonomicName id="4C6A4D7F10123610FB73DA73FAD61C28" ID-CoL="7CNYX" authorityName="Schmidt &amp; Faille" authorityYear="2015" box="[1231,1352,511,536]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="B91EEAEE10123610FB73DA73FAD61C28" box="[1231,1352,511,536]" italics="true" pageId="8" pageNumber="439">T. balticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A4D7F10123610FA3BDA73FECE1C0D" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FA3BDA73FECE1C0D" italics="true" pageId="8" pageNumber="439">T. eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FEE5DA56FE0F1C0D" box="[345,401,548,573]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FEE5DA56FE0F1C0D" bold="true" box="[345,401,548,573]" pageId="8" pageNumber="439">sp.n.</emphasis>
</taxonomicNameLabel>
, are closely related and may thus represent members of the same (extinct?) branch within
<taxonomicName id="4C6A4D7F10123610FF2BDA35FF711C50" ID-CoL="8KZB9" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[151,239,583,608]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B91EEAEE10123610FF2BDA35FF711C50" box="[151,239,583,608]" italics="true" pageId="8" pageNumber="439">Trechus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FF49DA3AFEF81C50" box="[245,358,583,608]" pageId="8" pageNumber="439" sensu="lato">sensu lato</taxonomicNameLabel>
. This hypothesis is based on two derived character states, which are potential synapomorphies of the two fossil species: i) microsculpture of pronotum and elytra consist of very fine very weakly impressed transverse meshes; ii) elytral striation markedly reduced laterally and apically. However, since these character states are also present in some of the extant
<taxonomicName id="4C6A4D7F10123610FD34DAC6FD7E1CFD" ID-CoL="8KZB9" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[648,736,692,717]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAEE10123610FD34DAC6FD7E1CFD" box="[648,736,692,717]" italics="true" pageId="8" pageNumber="439">Trechus</emphasis>
</taxonomicName>
species, this hypothesis needs further testing based on a more extensive fossil data set.
</paragraph>
<paragraph id="8BD536FC10123610FF7BDA8EFCE01DDD" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="439">
A particular character of the fossil
<taxonomicName id="4C6A4D7F10123610FDE9DA8FFCB51D25" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[597,811,764,789]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FDE9DA8FFCB51D25" box="[597,811,764,789]" italics="true" pageId="8" pageNumber="439">T. eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FC8FDA8EFCF51D25" box="[819,875,764,789]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FC8FDA8EFCF51D25" bold="true" box="[819,875,764,789]" pageId="8" pageNumber="439">sp.n.</emphasis>
</taxonomicNameLabel>
is the complete reduction of the compound eyes. Within the extant world fauna of
<taxonomicName id="4C6A4D7F10123610FD9EDB6DFD1E1D08" ID-CoL="8KTXW" authorityName="Bonelli" authorityYear="1810" box="[546,640,799,824]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
a huge number of anophthalmous and microphthalmous species are described, and their diversity is particularly rich in the southern parts of the Western Palaearctic (e.g.,
<bibRefCitation id="EFFB4B0D10123610FABFDB36FA061D6C" author="Jeannel, R." box="[1283,1432,836,861]" journalOrPublisher="L'Abeille" pageId="8" pageNumber="439" pagination="1 - 808" part="35" refString="Jeannel, R. (1928) Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Troisieme livraison). L'Abeille, 35, 1 - 808." title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Troisieme livraison)" type="journal article" year="1928">Jeannel 1928</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FF2BDB15FE311DB0" author="Casale, A. &amp; Laneyrie, R." box="[151,431,871,896]" journalOrPublisher="Memoires de Biospeologie" pageId="8" pageNumber="439" pagination="1 - 226" part="9" refString="Casale, A. &amp; Laneyrie, R. (1982) Trechodinae et Trechinae du Monde - tableau des sous-familles, tribus, series phyletiques, genres, et catalogue general des especes. Memoires de Biospeologie, 9, 1 - 226." title="Trechodinae et Trechinae du Monde - tableau des sous-familles, tribus, series phyletiques, genres, et catalogue general des especes" type="journal article" year="1982">Casale &amp; Laneyrie 1982</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FE07DB15FD151DB0" author="Casale, A. &amp; Vigna Taglianti, A. &amp; Juberthie, C." box="[443,651,871,896]" editor="Juberthie, C" journalOrPublisher="Societe Internationale de Biospeologie, pp. 1047 - 1081" pageId="8" pageNumber="439" refString="Casale, A., Vigna Taglianti, A. &amp; Juberthie, C. (1998) Coleoptera Carabidae. In: Juberthie, C. &amp; Decu, V. (Eds.), Encyclopaedia Biospeologica, Volume 2. Societe Internationale de Biospeologie, pp. 1047 - 1081." title="Coleoptera Carabidae" type="book" volumeTitle="Encyclopaedia Biospeologica, Volume 2" year="1998">
Casale
<emphasis id="B91EEAEE10123610FDB2DB1BFDD91DB0" box="[526,583,871,896]" italics="true" pageId="8" pageNumber="439">et al.</emphasis>
1998
</bibRefCitation>
). It has been shown, however, that different degrees of eye reduction evolved several times independently at least within
<taxonomicName id="4C6A4D7F10123610FD6BDBFEFCB11D95" ID-CoL="8KZB9" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[727,815,908,933]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B91EEAEE10123610FD6BDBFEFCB11D95" box="[727,815,908,933]" italics="true" pageId="8" pageNumber="439">Trechus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FC89DBFFFC3A1D95" box="[821,932,908,933]" pageId="8" pageNumber="439" sensu="lato">sensu lato</taxonomicNameLabel>
(
<bibRefCitation id="EFFB4B0D10123610FC0EDBFEFBA01D94" author="Casale, A." box="[946,1086,908,933]" journalOrPublisher="Contributions to Natural History" pageId="8" pageNumber="439" pagination="1 - 16" part="16" refString="Casale, A. (2011) Two new subterranean, microphthalmous trechine beetles from the Mediterranean area, and a synonymic note (Coleoptera: Carabidae, Trechini). Contributions to Natural History, 16, 1 - 16." title="Two new subterranean, microphthalmous trechine beetles from the Mediterranean area, and a synonymic note (Coleoptera: Carabidae, Trechini)" type="journal article" year="2011">Casale 2011</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FBF5DBFEFA901D94" author="Faille, A. &amp; Ribera, I &amp; Deharveng, L. &amp; Bourdeau, C. &amp; Garnery, L. &amp; Queinnec, E. &amp; Deuve, T." box="[1097,1294,908,933]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="8" pageNumber="439" pagination="97 - 105" part="54" publicationUrl="http://dx.doi.org/10.1016/j.ympev.2009.10.008" refString="Faille, A., Ribera, I., Deharveng, L., Bourdeau, C., Garnery, L., Queinnec, E. &amp; Deuve, T. (2010) A molecular phylogeny shows the single origin of the Pyrenean subterranean Trechini ground beetles (Coleoptera: Carabidae). Molecular Phylogenetics and Evolution, 54, 97 - 105. http: // dx. doi. org / 10.1016 / j. ympev. 2009.10.008" title="A molecular phylogeny shows the single origin of the Pyrenean subterranean Trechini ground beetles (Coleoptera: Carabidae)" type="journal article" year="2010">Faille et. al. 2010</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FAA5DBFEFACD1D94" author="Faille, A. &amp; Casale, A. &amp; Ribera, I." box="[1305,1363,908,932]" journalOrPublisher="Zoologica Scripta 40" pageId="8" pageNumber="439" pagination="282 - 295" part="3" publicationUrl="http://dx.doi.org/10.1111/j.1463-6409.2010.00467.x" refString="Faille, A., Casale, A. &amp; Ribera, I. (2011) Phylogenetic relationships of west Mediterranean troglobitic Trechini ground beetles (Coleoptera: Carabidae). Zoologica Scripta 40 (3), 282 - 295. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.2010.00467. x" title="Phylogenetic relationships of west Mediterranean troglobitic Trechini ground beetles (Coleoptera: Carabidae)" type="journal article" year="2011">2011</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FAE1DBFEFA061D94" author="Faille, A. &amp; Casale, A. &amp; Balke, M. &amp; Ribera, I." box="[1373,1432,908,932]" journalOrPublisher="BMC Evolutionary Biology" pageId="8" pageNumber="439" pagination="248" part="13" publicationUrl="http://dx.doi.org/10.1186/1471-2148-13-248" refString="Faille, A., Casale, A., Balke, M. &amp; Ribera, I. (2013) A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae). BMC Evolutionary Biology, 13, 248. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 13 - 248" title="A molecular phylogeny of alpine subterranean Trechini (Coleoptera: Carabidae)" type="journal article" year="2013">2013</bibRefCitation>
,
<bibRefCitation id="EFFB4B0D10123610FF2BDBC2FF4A1DF8" author="Faille, A. &amp; Andujar, C. &amp; Fadrique, F. &amp; Ribera, I." box="[151,212,944,968]" journalOrPublisher="Journal of Biogeography" pageId="8" pageNumber="439" pagination="1979 - 1990" part="41" publicationUrl="http://dx.doi.org/10.1111/jbi.12349" refString="Faille, A., Andujar, C., Fadrique, F. &amp; Ribera, I. (2014) Late Miocene origin of an Ibero-Maghrebian clade of ground beetles with multiple colonizations of the subterranean environment. Journal of Biogeography, 41, 1979 - 1990. http: // dx. doi. org / 10.1111 / jbi. 12349" title="Late Miocene origin of an Ibero-Maghrebian clade of ground beetles with multiple colonizations of the subterranean environment" type="journal article" year="2014">2014</bibRefCitation>
). Therefore, “eyelessness” as present in the fossil
<taxonomicName id="4C6A4D7F10123610FCA9DBC3FC721DF8" ID-CoL="57ZKM" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[789,1004,943,968]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FCA9DBC3FC721DF8" box="[789,1004,943,968]" italics="true" pageId="8" pageNumber="439">T. eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FC49DBDDFBB31DF8" box="[1013,1069,943,968]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FC49DBDDFBB31DF8" bold="true" box="[1013,1069,943,968]" pageId="8" pageNumber="439">sp.n.</emphasis>
</taxonomicNameLabel>
is likely not an indication for a close relationship with any of the extant eyeless
<taxonomicName id="4C6A4D7F10123610FD0ADBA6FC8A1DDD" ID-CoL="8KTXW" authorityName="Bonelli" authorityYear="1810" box="[694,788,980,1005]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
lineages.
</paragraph>
</subSubSection>
<subSubSection id="C370657710123610FF7BDB85FC611B3D" pageId="8" pageNumber="439" type="diagnosis">
<paragraph id="8BD536FC10123610FF7BDB85FC611B3D" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="439">
<emphasis id="B91EEAEE10123610FF7BDB85FE531A20" bold="true" box="[199,461,1015,1040]" pageId="8" pageNumber="439">Differential diagnosis.</emphasis>
The Baltic amber fossil species
<taxonomicName id="4C6A4D7F10123610FC8CDB85FC791A20" authorityName="Schmidt &amp; Faille" authorityYear="2015" box="[816,999,1015,1040]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="B91EEAEE10123610FC8CDB85FC791A20" box="[816,999,1015,1040]" italics="true" pageId="8" pageNumber="439">Trechus balticus</emphasis>
</taxonomicName>
is the only trechine beetle known from Paleogene deposits to date (
<bibRefCitation id="EFFB4B0D10123610FE70DC6EFD4F1A05" author="Schmidt, J. &amp; Faille, A." box="[460,721,1052,1077]" journalOrPublisher="Zootaxa" pageId="8" pageNumber="439" pagination="539 - 550" part="4028" publicationUrl="http://dx.doi.org/10.11646/zootaxa.4028.4.5" refString="Schmidt, J. &amp; Faille, A. (2015) Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini). Zootaxa, 4028 (4), 539 - 550. http: // dx. doi. org / 10.11646 / zootaxa. 4028.4.5" title="Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini)" type="journal article" year="2015">Schmidt &amp; Faille 2015</bibRefCitation>
).
<taxonomicName id="4C6A4D7F10123610FD5FDC6EFC641A05" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[739,1018,1052,1077]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FD5FDC6EFC641A05" box="[739,1018,1052,1077]" italics="true" pageId="8" pageNumber="439">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FBBDDC6EFBA71A05" box="[1025,1081,1052,1077]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FBBDDC6EFBA71A05" bold="true" box="[1025,1081,1052,1077]" pageId="8" pageNumber="439">sp.n.</emphasis>
</taxonomicNameLabel>
is easily distinguished from the latter by the completely reduced eyes, larger and more robust body, broader lateral gutter of pronotum, and the presence of a fine groove on external surface of the protibia.
<taxonomicName id="4C6A4D7F10123610FCE7DC16FABD1A4D" authority="Motschulsky, 1856" authorityName="Motschulsky" authorityYear="1856" box="[859,1315,1124,1149]" class="Insecta" family="Carabidae" genus="Trechoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B91EEAEE10123610FCE7DC16FBDC1A4C" box="[859,1090,1124,1149]" italics="true" pageId="8" pageNumber="439">Trechoides fasciatus</emphasis>
Motschulsky, 1856
</taxonomicName>
is another Baltic amber fossil with uncertain taxonomic position within
<taxonomicName id="4C6A4D7F10123610FCD3DCF5FC7D1A90" authorityName="" authorityYear="1802" box="[879,995,1159,1184]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
, and its
<typeStatus id="54D1885E10123610FBF3DCF5FB2F1A90" box="[1103,1201,1159,1184]" pageId="8" pageNumber="439" type="holotype">holotype</typeStatus>
specimen was very probably lost (
<bibRefCitation id="EFFB4B0D10123610FE84DCDEFDA01AF5" author="Schmidt, J. &amp; Faille, A." box="[312,574,1196,1221]" journalOrPublisher="Zootaxa" pageId="8" pageNumber="439" pagination="539 - 550" part="4028" publicationUrl="http://dx.doi.org/10.11646/zootaxa.4028.4.5" refString="Schmidt, J. &amp; Faille, A. (2015) Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini). Zootaxa, 4028 (4), 539 - 550. http: // dx. doi. org / 10.11646 / zootaxa. 4028.4.5" title="Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini)" type="journal article" year="2015">Schmidt &amp; Faille 2015</bibRefCitation>
). Comparison has thus to be based on the few characters given in the original description.
<taxonomicName id="4C6A4D7F10123610FE99DCBDFDDE1AD8" authority="Schmidt &amp; Hoffmann &amp; Michalik, 2016" authorityName="Schmidt &amp; Hoffmann &amp; Michalik" authorityYear="2016" box="[293,576,1231,1256]" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="439" phylum="Arthropoda" rank="species" species="eoanophthalmus" status="sp. nov.">
<emphasis id="B91EEAEE10123610FE99DCBDFDDE1AD8" box="[293,576,1231,1256]" italics="true" pageId="8" pageNumber="439">Trechus eoanophthalmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D579510123610FDF7DCBDFD1C1AD8" box="[587,642,1231,1256]" pageId="8" pageNumber="439" rank="species">
<emphasis id="B91EEAEE10123610FDF7DCBDFD1C1AD8" bold="true" box="[587,642,1231,1256]" pageId="8" pageNumber="439">sp.n.</emphasis>
</taxonomicNameLabel>
is distinguished from the latter taxon by the much larger body size (almost three times longer), the reduced eyes, and the apically rounded elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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