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<document id="7462433F75A3B9C4443ED77B4A469CE2" ID-DOI="10.5852/ejt.2023.905.2325" ID-ISSN="2118-9773" ID-Zenodo-Dep="10127256" ID-ZooBank="160592A4-64B3-40C7-BA75-8CE075F873A5" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1699620584683" checkinUser="plazi" docAuthor="Gumovsky, Alex" docDate="2023" docId="0385E767FF92AC4F6E02F936FE56FC6C" docLanguage="en" docName="EJT.2023.905.1-83.pdf" docOrigin="European Journal of Taxonomy 905" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2325/10109" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Colpixys Waterston 1916" docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="FFBC9F1FFF9DAC5F6C5BFFECFFFEFFFB" masterDocTitle="Revision of Xiphentedon Risbec, 1957 and Colpixys Waterston, 1916 (Hymenoptera, Eulophidae), with descriptions of new species from the Afrotropics" masterLastPageNumber="83" masterPageNumber="1" pageNumber="16" updateTime="1703245466857" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F93FE2E0B6064C389C11C6AB9D2EAEEA">Revision of Xiphentedon Risbec, 1957 and Colpixys Waterston, 1916 (Hymenoptera, Eulophidae), with descriptions of new species from the Afrotropics</mods:title>
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<paragraph id="8B935671FF92AC506E02F936FC24F90F" blockId="15.[601,986,1753,1780]" box="[601,986,1753,1780]" pageId="15" pageNumber="16">
<heading id="D0DBE11DFF92AC506E02F936FC24F90F" box="[601,986,1753,1780]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
Genus
<taxonomicName id="4C2C2DF2FF92AC506EF7F935FC24F90F" ID-CoL="3SKK" authority="Waterston, 1916" authorityName="Waterston" authorityYear="1916" box="[684,986,1753,1780]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B9588A63FF92AC506EF7F935FCEAF90F" bold="true" box="[684,788,1753,1780]" italics="true" pageId="15" pageNumber="16">Colpixys</emphasis>
<bibRefCitation id="EFBD2B80FF92AC506F40F936FC24F90F" author="Waterston J." box="[795,986,1754,1780]" pageId="15" pageNumber="16" pagination="123 - 132" refId="ref33512" refString="Waterston J. 1916. Notes on African Chalcidoidea-V. Bulletin of Entomological Research 7: 123 - 132. https: // doi. org / 10.1017 / S 0007485300017442" type="journal article" year="1916">Waterston, 1916</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C33605FAFF92AC506CE6F8F0FDDEF883" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8B935671FF92AC506CE6F8F0FDD3F8CD" blockId="15.[189,557,1819,1846]" box="[189,557,1819,1846]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB3C715FFF92AC506CE6F8F0FDD3F8CD" box="[189,557,1819,1846]" pageId="15" pageNumber="16">
<treatmentCitation id="0A8D7060FF92AC506CE6F8F0FDD6F8CD" author="Waterston J." box="[189,552,1819,1846]" page="129" pageId="15" pageNumber="16" year="1916">
<taxonomicName id="4C2C2DF2FF92AC506CE6F8F0FDD6F8CD" ID-CoL="3SKK" authority="Waterston, 1916: 129" authorityName="Waterston" authorityPageNumber="129" authorityYear="1916" box="[189,552,1819,1846]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B9588A63FF92AC506CE6F8F0FEDAF8CD" box="[189,292,1820,1846]" italics="true" pageId="15" pageNumber="16">Colpixys</emphasis>
<bibRefCitation id="EFBD2B80FF92AC506D71F8F0FDD6F8CD" author="Waterston J." box="[298,552,1819,1846]" pageId="15" pageNumber="16" pagination="123 - 132" refId="ref33512" refString="Waterston J. 1916. Notes on African Chalcidoidea-V. Bulletin of Entomological Research 7: 123 - 132. https: // doi. org / 10.1017 / S 0007485300017442" type="journal article" year="1916">Waterston, 1916: 129</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B935671FF92AC506CE6F8B1FDDEF883" blockId="15.[189,544,1885,1912]" box="[189,544,1885,1912]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB3C715FFF92AC506CE6F8B1FDDEF883" box="[189,544,1885,1912]" pageId="15" pageNumber="16">
<emphasis id="B9588A63FF92AC506CE6F8B1FEC4F88C" box="[189,314,1885,1911]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C2C2DF2FF92AC506CE6F8B1FEDAF88C" ID-CoL="3SKK" authorityName="Waterston" authorityYear="1916" box="[189,292,1885,1911]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Colpixys</taxonomicName>
</emphasis>
<treatmentCitation id="0A8D7060FF92AC506D1AF8B1FDE2F883" author="Boucek Z." box="[321,540,1885,1912]" page="203" pageId="15" pageNumber="16" year="1972">
<bibRefCitation id="EFBD2B80FF92AC506D1AF8B1FDE2F883" author="Boucek Z." box="[321,540,1885,1912]" pageId="15" pageNumber="16" pagination="199 - 205" refId="ref32376" refString="Boucek Z. 1972. Descriptions of new eulophid parasites (Hym., Chalcidoidea) from Africa and the Canary Islands. Bulletin of Entomological Research 62: 199 - 205. https: // doi. org / 10.1017 / S 0007485300047647" type="journal article" year="1972">Bouček 1972: 203</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C33605FAFF92AC506CE6F873FBDEF813" pageId="15" pageNumber="16" type="type_taxon">
<paragraph id="8B935671FF92AC506CE6F873FEA6F842" blockId="15.[189,344,1951,1977]" box="[189,344,1951,1977]" pageId="15" pageNumber="16">
<heading id="D0DBE11DFF92AC506CE6F873FEA6F842" bold="true" box="[189,344,1951,1977]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B9588A63FF92AC506CE6F873FEA6F842" bold="true" box="[189,344,1951,1977]" pageId="15" pageNumber="16">
<typeStatus id="5497E8D3FF92AC506CE6F873FF04F842" box="[189,250,1951,1977]" pageId="15" pageNumber="16">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B935671FF92AC506CE6F822FBDEF813" blockId="15.[189,1056,1998,2025]" box="[189,1056,1998,2025]" pageId="15" pageNumber="16">
<taxonomicName id="4C2C2DF2FF92AC506CE6F822FDB3F813" ID-CoL="X8MX" authority="Waterston, 1916" authorityName="Waterston" authorityYear="1916" box="[189,589,1998,2025]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="necator">
<emphasis id="B9588A63FF92AC506CE6F822FE7BF813" box="[189,389,1998,2024]" italics="true" pageId="15" pageNumber="16">Colpixys necator</emphasis>
<bibRefCitation id="EFBD2B80FF92AC506DD0F823FDB3F813" author="Waterston J." box="[395,589,1998,2025]" pageId="15" pageNumber="16" pagination="123 - 132" refId="ref33512" refString="Waterston J. 1916. Notes on African Chalcidoidea-V. Bulletin of Entomological Research 7: 123 - 132. https: // doi. org / 10.1017 / S 0007485300017442" type="journal article" year="1916">Waterston, 1916</bibRefCitation>
</taxonomicName>
, by monotypy and original designation.
</paragraph>
</subSubSection>
<subSubSection id="C33605FAFF8DAC4F6CE6FEE4FD77FD69" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B935671FF8DAC4F6CE6FEE4FEC8FED9" blockId="16.[189,310,264,290]" box="[189,310,264,290]" pageId="16" pageNumber="17">
<heading id="D0DBE11DFF8DAC4F6CE6FEE4FEC8FED9" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9588A63FF8DAC4F6CE6FEE4FEC8FED9" bold="true" box="[189,310,264,290]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B935671FF8DAC4F6CE6FED4FD77FD69" blockId="16.[187,1401,311,658]" pageId="16" pageNumber="17">
Pronotum with lateral semicircular ridge under protruding shoulders (
<figureCitation id="13174AF4FF8DAC4F6848FEDBFB8DFEA9" box="[1043,1139,311,338]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[189,1399,627,1792]" captionTargetId="figure-264@6.[189,1399,627,1792]" captionTargetPageId="6" captionText="Fig. 2. AC. Entedon sparetus Walker, 1839, ♀ (reared from Gymnetron asellus (Gravenhorst, 1807)) (used for SEM). A. Head and mesosoma. B. Enlarged area in frame on Fig. A. C. Orifices enlarged. D. E. diotimus Walker, 1839, ♀, plica on lateral panel of pronotum (used for SEM). EF. Lateral panel of pronotum. E. Colpixys necator Waterson, 1916, ♀ (NHMUK). F. Xiphentedon neserorum sp. nov., ♀ (paratype, NHMUK). Abbreviations: lpl = lateral pronotal plica; or = orifices; psh = lateral pronotal shoulder." figureDoi="http://doi.org/10.5281/zenodo.10127260" httpUri="https://zenodo.org/record/10127260/files/figure.png" pageId="16" pageNumber="17">Fig. 2E</figureCitation>
); face with V-shaped frontal suture; propodeum with coriaceous median furrow delimited by margins which are subparallel or weakly diverging posteriorly and more or less straight, submedian areas usually reticulate: lighter anteriorly and coarser posteriorly, lateral sulcus of propodeum incomplete (
<figureCitation id="13174AF4FF8DAC4F681EFE4EFB5FFE47" box="[1093,1185,418,444]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[189,1399,535,1756]" captionTargetId="figure-212@4.[189,1399,535,1756]" captionTargetPageId="4" captionText="Fig. 1. Propodeum. A. Entedon sylvestris Szelényi, 1981, ♀ (used for SEM). B. Entedon crassiscapus Erdös, 1944, ♀ (used for SEM). C. Entedon magnificus (Girault &amp; Dodd, 1913), ♀ (SANC). D. Xiphentedon neserorum sp. nov., ♀ (used for SEM). E. Pediobius ropalidiae (Risbec, 1958), ♀ (paratype, MRAC). F. Colpixys gigas Bouček, 1972, ♀ (paratype, NHMUK). Abbreviations: lc = lateral channel on sides of median carina; lp = lateral plica; ls = lateral sulcus of propodeum; mc = median carina; mf = median furrow; msc = metascutellum; mstr = median strip; smc = submedian carinae; spt = spiracular tubercle on propodeum." figureDoi="http://doi.org/10.5281/zenodo.10127258" httpUri="https://zenodo.org/record/10127258/files/figure.png" pageId="16" pageNumber="17">Figs 1F</figureCitation>
,
<figureCitation id="13174AF4FF8DAC4F68EBFE4EFB2AFE47" box="[1200,1236,418,444]" captionStart="Fig" captionStartId="18.[190,233,1926,1952]" captionTargetBox="[189,1399,452,1882]" captionTargetId="figure-125@18.[189,1399,452,1882]" captionTargetPageId="18" captionText="Fig. 8. Colpixys necator Waterston, 1916 (SANC). AD. ♀, from Zimbabwe. EF. ♂, from RSA. AB, EF. Habitus in dorso-lateral (A, F), lateral (B) and dorsal (E) views. C. Propodeum. D. Head and anterior part of mesosoma in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10127274" httpUri="https://zenodo.org/record/10127274/files/figure.png" pageId="16" pageNumber="17">8C</figureCitation>
,
<figureCitation id="13174AF4FF8DAC4F68B8FE4EFAD2FE47" box="[1251,1324,418,444]" captionStart="Fig" captionStartId="20.[189,232,1845,1871]" captionTargetBox="[189,1399,265,1792]" captionTargetId="figure-55@20.[189,1399,265,1792]" captionTargetPageId="20" captionText="Fig. 9. A, CD. Colpixys necator Waterston, 1916 (NHMUK). B. C. eburnus sp. nov. (paratype, Ivory Coast; CBGP). EF. C. gigas Bouček, 1972 (paratype, Ghana; NHMUK). AB. Propodeum. CD, F. Axillula. E. Head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10127276" httpUri="https://zenodo.org/record/10127276/files/figure.png" pageId="16" pageNumber="17">9AB</figureCitation>
,
<figureCitation id="13174AF4FF8DAC4F6961FE4EFA94FE47" box="[1338,1386,418,444]" captionStart="Fig" captionStartId="24.[189,232,1755,1781]" captionTargetBox="[189,1399,265,1705]" captionTargetId="figure-55@24.[189,1399,265,1705]" captionTargetPageId="24" captionText="Fig. 11. Colpixys eburnus sp. nov., ♀, holotype (Ivory Coast; CBGP). AB. Wings under direct (A) and reflected (B, showing WIP) light. C, E. Habitus in lateral (C) and dorsal (E) views. D. Head in frontal view. F. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.10127280" httpUri="https://zenodo.org/record/10127280/files/figure.png" pageId="16" pageNumber="17">11F</figureCitation>
); prosternum evenly curved or at most with a carina (
<taxonomicName id="4C2C2DF2FF8DAC4F6F44FE2AFC5FFE1B" authority="Gumovsky, 2023" authorityName="Gumovsky" authorityYear="2023" box="[799,929,454,480]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="eburnus" status="sp. nov.">
<emphasis id="B9588A63FF8DAC4F6F44FE2AFCCDFE1B" box="[799,819,454,480]" italics="true" pageId="16" pageNumber="17">C</emphasis>
.
<emphasis id="B9588A63FF8DAC4F6F19FE2AFC5FFE1B" box="[834,929,454,480]" italics="true" pageId="16" pageNumber="17">eburnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B3718FF8DAC4F6FF3FE2AFBF9FE1B" box="[936,1031,454,480]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
); mesopectus poorly pubescent above mesofurcal pit; mesepisternum without epicnemial protrusion; axillula without a projection, at most with carinulae or plicae(
<figureCitation id="13174AF4FF8DAC4F6E4DFDE1FD4FFDDC" box="[534,689,525,551]" captionStart="Fig" captionStartId="20.[189,232,1845,1871]" captionTargetBox="[189,1399,265,1792]" captionTargetId="figure-55@20.[189,1399,265,1792]" captionTargetPageId="20" captionText="Fig. 9. A, CD. Colpixys necator Waterston, 1916 (NHMUK). B. C. eburnus sp. nov. (paratype, Ivory Coast; CBGP). EF. C. gigas Bouček, 1972 (paratype, Ghana; NHMUK). AB. Propodeum. CD, F. Axillula. E. Head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10127276" httpUri="https://zenodo.org/record/10127276/files/figure.png" pageId="16" pageNumber="17">Fig. 9CD, F</figureCitation>
); metascutellum long, semi-circular (but somewhat shorter in
<taxonomicName id="4C2C2DF2FF8DAC4F6CE0FDDDFEC3FDB0" authority="Gumovsky, 2023" authorityName="Gumovsky" authorityYear="2023" box="[187,317,561,587]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="eburnus" status="sp. nov.">
<emphasis id="B9588A63FF8DAC4F6CE0FDDDFF31FDB0" box="[187,207,561,587]" italics="true" pageId="16" pageNumber="17">C</emphasis>
.
<emphasis id="B9588A63FF8DAC4F6C85FDDDFEC3FDB0" box="[222,317,561,587]" italics="true" pageId="16" pageNumber="17">eburnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B3718FF8DAC4F6D19FDDDFE63FDB0" box="[322,413,561,587]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
); supracoxal flange with setae; gena not bulging; male antenna with three-segmented funicle and two-segmented clava (
<taxonomicName id="4C2C2DF2FF8DAC4F6E0CFDB9FD2AFD95" authorityName="Waterston" authorityYear="1916" box="[599,724,597,623]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="necator">
<emphasis id="B9588A63FF8DAC4F6E0CFDB9FD95FD94" box="[599,619,597,623]" italics="true" pageId="16" pageNumber="17">C</emphasis>
.
<emphasis id="B9588A63FF8DAC4F6E21FDB9FD2AFD95" box="[634,724,597,622]" italics="true" pageId="16" pageNumber="17">necator</emphasis>
</taxonomicName>
) or five-segmented funicle (clava not present,
<taxonomicName id="4C2C2DF2FF8DAC4F695AFDB9FA9BFD95" authorityName="Boucek" authorityYear="1972" box="[1281,1381,597,623]" class="Insecta" family="Eulophidae" genus="Colpixys" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B9588A63FF8DAC4F695AFDB9FA9BFD95" box="[1281,1381,597,623]" italics="true" pageId="16" pageNumber="17">C. gigas</emphasis>
</taxonomicName>
); male gaster without pale subbasal spot.
</paragraph>
</subSubSection>
<subSubSection id="C33605FAFF8DAC4F6CE6FD55FE4FFCDD" pageId="16" pageNumber="17" type="biology_ecology">
<paragraph id="8B935671FF8DAC4F6CE6FD55FEE2FD28" blockId="16.[189,284,697,723]" box="[189,284,697,723]" pageId="16" pageNumber="17">
<heading id="D0DBE11DFF8DAC4F6CE6FD55FEE2FD28" bold="true" box="[189,284,697,723]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9588A63FF8DAC4F6CE6FD55FEE2FD28" bold="true" box="[189,284,697,723]" pageId="16" pageNumber="17">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8B935671FF8DAC4F6CE6FD04FE4FFCDD" blockId="16.[186,1398,744,806]" pageId="16" pageNumber="17">
The available records concern parasitism of larvae of
<taxonomicName id="4C2C2DF2FF8DAC4F6F45FD04FC66FCF9" authorityName="Latreille" authorityYear="1802" box="[798,920,744,770]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Erotylidae</taxonomicName>
and
<taxonomicName id="4C2C2DF2FF8DAC4F6F90FD04FBA7FCF9" authorityName="Hope" authorityYear="1840" box="[971,1113,744,770]" class="Insecta" family="Languriidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Languriidae</taxonomicName>
(
<taxonomicName id="4C2C2DF2FF8DAC4F683EFD04FB14FCF9" authorityName="Linnaeus" authorityYear="1758" box="[1125,1258,744,770]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) (
<bibRefCitation id="EFBD2B80FF8DAC4F68A5FD05FF06FCDD" author="Waterston J." pageId="16" pageNumber="17" pagination="123 - 132" refId="ref33512" refString="Waterston J. 1916. Notes on African Chalcidoidea-V. Bulletin of Entomological Research 7: 123 - 132. https: // doi. org / 10.1017 / S 0007485300017442" type="journal article" year="1916">Waterston 1916</bibRefCitation>
;
<bibRefCitation id="EFBD2B80FF8DAC4F6D5FFCE0FE5AFCDD" author="Boucek Z." box="[260,420,780,806]" pageId="16" pageNumber="17" pagination="199 - 205" refId="ref32376" refString="Boucek Z. 1972. Descriptions of new eulophid parasites (Hym., Chalcidoidea) from Africa and the Canary Islands. Bulletin of Entomological Research 62: 199 - 205. https: // doi. org / 10.1017 / S 0007485300047647" type="journal article" year="1972">Bouček 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33605FAFF8DAC4F6CE6FCA2FE56FC6C" box="[189,424,846,919]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B935671FF8DAC4F6CE6FCA2FEA8FC93" blockId="16.[189,342,846,872]" box="[189,342,846,872]" pageId="16" pageNumber="17">
<heading id="D0DBE11DFF8DAC4F6CE6FCA2FEA8FC93" bold="true" box="[189,342,846,872]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9588A63FF8DAC4F6CE6FCA2FEA8FC93" bold="true" box="[189,342,846,872]" pageId="16" pageNumber="17">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B935671FF8DAC4F6CE6FC91FE56FC6C" blockId="16.[189,424,893,919]" box="[189,424,893,919]" pageId="16" pageNumber="17">Afrotropical region.</paragraph>
</subSubSection>
</treatment>
</document>