178 lines
17 KiB
XML
178 lines
17 KiB
XML
<document id="9BFF4AA5DCBA701C2D9B6C116CFC1209" ID-CLB-Dataset="28831" ID-DOI="http://dx.doi.org/10.3897/zookeys.796.21877" ID-GBIF-Dataset="0476ce18-4e9c-4ee6-8d6e-99d1146cdbf4" ID-PMC="PMC6256130" ID-Pensoft-Pub="1313-2970-796-215" ID-PubMed="30505157" ID-ZooBank="CC96882F71234A2D83457C5E36F817C1" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-796-215" ModsDocOrigin="ZooKeys 796" ModsDocTitle="Review of the subgenus Plumiger of Myrmecophyes, with description of a new species (Heteroptera, Miridae, Halticini)" checkinTime="1542332380955" checkinUser="pensoft" docAuthor="Konstantinov, Fedor V. & Simov, Nikolay" docDate="2018" docId="A79985823EEC3F3DDA0B402A2C359538" docLanguage="en" docName="ZooKeys 796: 215-239" docOrigin="ZooKeys 796" docSource="http://dx.doi.org/10.3897/zookeys.796.21877" docTitle="Myrmecophyes armeniacus Drapolyuk 1989" docType="treatment" docVersion="7" lastPageNumber="226" masterDocId="6D0C0038FFD8FFBFFFD8867C7F0D2327" masterDocTitle="Review of the subgenus Plumiger of Myrmecophyes, with description of a new species (Heteroptera, Miridae, Halticini)" masterLastPageNumber="239" masterPageNumber="215" originalUpdateDomain="Boston" originalUpdateTime="1542332380955" originalUpdateUser="pensoft" pageNumber="225" updateTime="1701382271246" updateUser="ExternalLinkService">
|
||
<mods:mods id="CD3AC4CADB5D95D286769685E22C5C75" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E67B04935823EC94867D6D518E1F5D59">
|
||
<mods:title id="7DE4F284D6A83DED565BB821D427BBBC">Review of the subgenus Plumiger of Myrmecophyes, with description of a new species (Heteroptera, Miridae, Halticini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="59A263CE907809713A767FD7BBDD36CC" type="personal">
|
||
<mods:role id="6BA5B9F0F8915FE7B2CDE39425CE4642">
|
||
<mods:roleTerm id="2B3FEF2EEEAA883389B264CED9E1198A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CEB56B24FCC6C36204BA305DB02606B7">Konstantinov, Fedor V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="3B4C59F407D3166EC374FDAAC6E6DF72" type="personal">
|
||
<mods:role id="1E8647F3725C7998F7BC586E73206F5D">
|
||
<mods:roleTerm id="876A72C0CE842E698BB28C9D5ACD9CB2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="83259826E2D846C479BCF77BF28D4107">Simov, Nikolay</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B4630A48F3B7988D130333F86AA6D579">text</mods:typeOfResource>
|
||
<mods:relatedItem id="991877E4F94D72DA316373B8F1717AB9" type="host">
|
||
<mods:titleInfo id="36AD26B7A8676AAB59C759871396BAB4">
|
||
<mods:title id="30C3554EFAA57E9BA446D509ECC09ECC">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E2F318C830C5A214D79ED2DFCDA25AE4">
|
||
<mods:date id="F2555259200E79EF85D60FADF676DAB2">2018</mods:date>
|
||
<mods:detail id="EF9F4793FA1CB5872017FF9C075AF40E" type="volume">
|
||
<mods:number id="0F32FFCAA48CF8A7E8198758D21CDE49">796</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B069C80B2450BBEF5DAA369A4B73D3B0" unit="page">
|
||
<mods:start id="69E9267B96A9E27D4726CFC273AB6FC4">215</mods:start>
|
||
<mods:end id="94EC1532B00404FDBBF62B1A313A71DA">239</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="DDA493C1BF4E9B7BA85FEFAD41D50923">
|
||
<mods:url id="333D7E07C264408B1A3F2EE698EEB450">http://dx.doi.org/10.3897/zookeys.796.21877</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="8E241E112EC241A8467AAB58BCB40C49">journal article</mods:classification>
|
||
<mods:identifier id="67871E18AF9491DD9F37765BF50386C4" type="DOI">http://dx.doi.org/10.3897/zookeys.796.21877</mods:identifier>
|
||
<mods:identifier id="694256D940C7FD1583409B6B69D95A99" type="Pensoft-Pub">1313-2970-796-215</mods:identifier>
|
||
<mods:identifier id="D1AA75DFBCC32BB00702C372A1403526" type="ZooBank">CC96882F71234A2D83457C5E36F817C1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="A79985823EEC3F3DDA0B402A2C359538" ID-GBIF-Taxon="149667423" LSID="urn:lsid:plazi:treatment:A79985823EEC3F3DDA0B402A2C359538" httpUri="http://treatment.plazi.org/id/A79985823EEC3F3DDA0B402A2C359538" lastPageId="11" lastPageNumber="226" pageId="10" pageNumber="225">
|
||
<subSubSection id="CC3279062E1045916EB3B458549BFCFB" pageId="10" pageNumber="225" type="nomenclature">
|
||
<paragraph id="5B0009A7F6AD260B0B2AFF4542CFE3E4" pageId="10" pageNumber="225">
|
||
<taxonomicName id="E5AB79617DF5D441200AB1E65D0A5FAE" ID-CoL="45B7Y" authority="Drapolyuk, 1989" authorityName="Drapolyuk" authorityYear="1989" class="Insecta" family="Miridae" genus="Myrmecophyes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmecophyes armeniacus" order="Hemiptera" pageId="10" pageNumber="225" phylum="Arthropoda" rank="species" species="armeniacus">
|
||
<pageBreakToken id="ABDE3B53F1E991A3C70D87BCE72B39CE" pageId="10" pageNumber="225" start="start">Myrmecophyes</pageBreakToken>
|
||
armeniacus Drapolyuk, 1989
|
||
</taxonomicName>
|
||
Figs 5, 8, 14, 16, 22, 23, 25, 26, 30-33, 37, 38, 51, 55, 59, 64
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="27840435B1B6278F446E0B97DB7F32B9" pageId="10" pageNumber="225" type="reference_group">
|
||
<paragraph id="11E511AF370F06FC97DF61495008CB46" pageId="10" pageNumber="225">
|
||
<taxonomicName id="36A2FB4BE7C01B0D90473DE201F74D24" class="Insecta" family="Miridae" genus="Myrmecophyes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmecophyes (Plumiger) armeniacus" order="Hemiptera" pageId="10" pageNumber="225" phylum="Arthropoda" rank="species" species="armeniacus" subGenus="Plumiger">Myrmecophyes (Plumiger) armeniacus</taxonomicName>
|
||
Dapolyuk, 1989: 125; figs 51, 55.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D074B2C76C19DA4E6609E3BE80511F87" pageId="10" pageNumber="225" type="materials_examined">
|
||
<paragraph id="EE91732481DD73DB3D5AD89B6FB604FB" pageId="10" pageNumber="225">Material examined.</paragraph>
|
||
<paragraph id="ED3F87666C7F30E5B8428B93CDEA2825" pageId="10" pageNumber="225">
|
||
Holotype: TURKEY: Kars: Kars,
|
||
<geoCoordinate id="91497532AF0BEAD605BD88C0A0DD841B" direction="north" orientation="latitude" precision="1" value="40.58333">40.58333°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D3BD79229B7784A2051E36773088C958" direction="east" orientation="longitude" precision="1" value="43.06666">43.06666°E</geoCoordinate>
|
||
, 04 Jun 1915, Olsufiev, 1♀ (AMNH_PBI 00261681) (ZISP). Paratypes: ARMENIA: Shirak: Gyumri [Leninakan],
|
||
<geoCoordinate id="67045C2127D199F1B66FBEE7592D4977" direction="north" orientation="latitude" precision="1" value="40.78333">40.78333°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0CDBE24E7DA330E2D8DFEF6CD086C022" direction="east" orientation="longitude" precision="1" value="43.83333">43.83333°E</geoCoordinate>
|
||
, 16 Jul 1940, Esterberg, 1♀ (AMNH_PBI 00261680) (ZISP).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1D74C7FD0C3280EA38A87AEEA75EE916" pageId="10" pageNumber="225" type="materials_examined">
|
||
<paragraph id="26FBA707C10891D599DDBD0C8C3B6499" pageId="10" pageNumber="225">Other specimens examined:</paragraph>
|
||
<paragraph id="C688AAB35CB42877567E06FE76A84F34" pageId="10" pageNumber="225">
|
||
ARMENIA: Gegharkunik: N coast of Sevan Lake, Artanish - Shorzha Rd,
|
||
<geoCoordinate id="7B090DB4A9FACC3DB351CC7E5473B368" direction="north" orientation="latitude" precision="1" value="40.50109">40.50109°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="67F1A2513EF76A2EF282652C56470716" direction="east" orientation="longitude" precision="5" value="45.3282">45.3282°E</geoCoordinate>
|
||
, 1962 m, 14 Jun 2017, F. Konstantinov & N. Simov, (
|
||
<taxonomicName id="228418227088656B241F970F1FB74B4C" family="Poaceae" lsidName="Heteroptera" pageId="10" pageNumber="225" rank="family">Poaceae</taxonomicName>
|
||
), 15♂ (AMNH_PBI 00343264-AMNH_PBI 00343277, AMNH_PBI 00343331), 70♀ (AMNH_PBI 00343249-AMNH_PBI 00343263, AMNH_PBI 00343232-AMNH_PBI 00343248, AMNH_PBI 00343196-AMNH_PBI 00343231, AMNH_PBI 00343330, AMNH_PBI 00343376) (ZISP); 16♂, 49♀ (SOFM). N coast of Sevan Lake, Karmir Pass, 4 km ENE of Aghberk,
|
||
<geoCoordinate id="0C97054DBF26EC66A940C244646941E1" direction="north" orientation="latitude" precision="1" value="40.56328">40.56328°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="85388848CB980AA3FB9922146B2BC583" direction="east" orientation="longitude" precision="1" value="45.29903">45.29903°E</geoCoordinate>
|
||
, 2190 m, 15 Jun 2017, F. Konstantinov & N. Simov, (
|
||
<taxonomicName id="16A3748BCEF993BE37F5EA9C118BD7AB" family="Poaceae" lsidName="Heteroptera" pageId="10" pageNumber="225" rank="family">Poaceae</taxonomicName>
|
||
), 2♂ (AMNH_PBI 00343351, AMNH_PBI 00343352), 1♀ (AMNH_PBI 00343327) (ZISP); 6♂, 9♀ (SOFM). W coast of Sevan Lake, 2 km ESE of Semyonovka,
|
||
<geoCoordinate id="B95E71D54DC430953E13F1D825EB7AB4" direction="north" orientation="latitude" precision="1" value="40.65074">40.65074°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B47654161626E83091E835C4597AB20A" direction="east" orientation="longitude" precision="1" value="44.92336">44.92336°E</geoCoordinate>
|
||
, 2123 m, 15 Jun 2017, F. Konstantinov & N. Simov, (
|
||
<taxonomicName id="DC23ED258E4A8D1147F58DCF65DD4E97" family="Poaceae" lsidName="Heteroptera" pageId="10" pageNumber="225" rank="family">Poaceae</taxonomicName>
|
||
), 24♂ (AMNH_PBI 00343278-AMNH_PBI 00343285, AMNH_PBI 00343288-AMNH_PBI 00343301, AMNH_PBI 00343332, AMNH_PBI 00343334), 25♀ (AMNH_PBI 00343302-AMNH_PBI 00343312, AMNH_PBI 00343314-AMNH_PBI 00343325, AMNH_PBI 00343333, AMNH_PBI 00343328) (ZISP); 25♂, 34♀ (SOFM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="90423A06FEEAC9137FD4C952E1066CB0" pageId="10" pageNumber="225" type="diagnosis">
|
||
<paragraph id="090AB27960BDCDB3AB002DBDCFBE9AB1" pageId="10" pageNumber="225">Diagnosis.</paragraph>
|
||
<paragraph id="3BE5C63AB3CAB5DDC4D8337B5F014E38" pageId="10" pageNumber="225">Size large, clypeus straight, not bulging apically in either sex (Figs 14, 26), head and abdomen not shiny (Figs 5, 8), antennal segment II in male shorter than width of pronotum; shape of the left paramere as in Figs 30-32, lobes of aedeagus membranous (Figure 37), and sclerotized rings of dorsal labiate plate broadly oval (Figs 51, 55).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C2361F25987BD49D4E2C3ADE084BF30E" pageId="10" pageNumber="225" type="remarks">
|
||
<paragraph id="C7D59108AA3256E5DD4C0902E29CD0CB" pageId="10" pageNumber="225">Remarks.</paragraph>
|
||
<paragraph id="7B64ADFDF1B207960A01659C689CBD72" pageId="10" pageNumber="225">
|
||
Males of
|
||
<taxonomicName id="883269ECF53F70F28F92086D5106F5F6" lsidName="M. armeniacus" pageId="10" pageNumber="225" rank="species" species="armeniacus">M. armeniacus</taxonomicName>
|
||
are most similar to
|
||
<taxonomicName id="D2B34F30DA527791D0B628A485011BE3" lsidName="M. heterocerus" pageId="10" pageNumber="225" rank="species" species="heterocerus">M. heterocerus</taxonomicName>
|
||
in structure of the head and male genitalia, but the latter species can be distinguished by the smaller size, shiny head and abdomen, and structure of the left paramere with the apical process narrowing before the T-shaped blade and equipped with a prominent outgrowth at base (compare Figs 30, 34). Females of
|
||
<taxonomicName id="B652F73953D5CD65F34BC1BA74DEDB92" lsidName="M. armeniacus" pageId="10" pageNumber="225" rank="species" species="armeniacus">M. armeniacus</taxonomicName>
|
||
and
|
||
<taxonomicName id="8542F00708BFABCA35D2126A1885311E" lsidName="M. nasutus" pageId="10" pageNumber="225" rank="species" species="nasutus">M. nasutus</taxonomicName>
|
||
are roughly similar in size and vestiture, and the surface of the dorsum is not shiny; these species can be separated by their differently shaped sclerotized rings (compare Figs 51, 54).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="47B4730F77C6F15212C19AF1193F0F54" lastPageId="11" lastPageNumber="226" pageId="10" pageNumber="225" type="redescription">
|
||
<paragraph id="DE04EA7C8D0837EABDB42EF3A6A36AA7" pageId="10" pageNumber="225">Redescription.</paragraph>
|
||
<paragraph id="9191D7C5F80422B850D8547B2490F9A8" pageId="10" pageNumber="225">
|
||
Male. Total length 3.4-3.7. Coloration: (Figure 5): entirely black, with whitish transverse stripe along apex of wing pad; fore and middle femora usually, hind femur rarely with pale brown apices. Surface and vestiture: Dorsum matt, head and abdomen at most moderately shiny; vestiture as in subgeneric description. Structure: Body 3.9-4.2
|
||
<normalizedToken id="355F907B3E692BD5B50962C577526B7C" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Head: Clypeus not bulging (Figure 26), flat at base and slightly convex in apical part, with apical margin straight; vertex 1.8-2.0
|
||
<normalizedToken id="B14703B74BDCD9E91D16E363A4F193FD" originalValue="×">x</normalizedToken>
|
||
as wide as eye; antennal segment I strongly swollen, short, 0.8-0.9
|
||
<normalizedToken id="E0C2C3477BF8F5B69E460847330C5CCB" originalValue="×">x</normalizedToken>
|
||
as long as length of pronotum, segment II slightly curved and widened apically, rather short, 1.4-1.5
|
||
<normalizedToken id="C5AC377AA371AEE174541FC4EDC71D0E" originalValue="×">x</normalizedToken>
|
||
as long as I, 0.8
|
||
<normalizedToken id="2B1CE9F42C80934A5FC4DAD8AECC71FF" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle, and 0.6
|
||
<normalizedToken id="EC16178497D76E313436DCF00CEA2FF1" originalValue="×">x</normalizedToken>
|
||
as long as width of head; segment III more than 2.8-2.9
|
||
<normalizedToken id="B6B83848911F8E7674CE7CA275B327C5" originalValue="×">x</normalizedToken>
|
||
as long as II and distinctly longer than I and II combined; segment IV about 1.5
|
||
<normalizedToken id="A81C2CCDB77EF616F52A3BC9766FEAB9" originalValue="×">x</normalizedToken>
|
||
as long as II. Thorax: Pronotum at middle 1.3
|
||
<normalizedToken id="CC66E416BC392E66B8A0937C4DA53A25" originalValue="×">x</normalizedToken>
|
||
as wide as long, 0.7-0.8
|
||
<normalizedToken id="26813B6D7EA39B84187D04E45E5E6516" originalValue="×">x</normalizedToken>
|
||
as wide as head; hind tibia 3.6-4.0
|
||
<normalizedToken id="40651EC0EAF7D5FC2F3BA30FCA64CD29" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Genitalia: genital capsule comparatively small, about 0.3 of abdomen; left paramere sickle-shaped, thin, apical process of uniform width along entire length, long, somewhat sinuate, flattened, ending with characteristic oblique T-shaped blade; body of paramere with small but distinct rounded outgrowth at base of apical process with long setae (Figs 30-32); right paramere typically flag-shaped, somewhat larger than in
|
||
<taxonomicName id="D620F9859074A5529A0F61089866185D" lsidName="M. heterocerus" pageId="10" pageNumber="225" rank="species" species="heterocerus">M. heterocerus</taxonomicName>
|
||
(Figure 33); endosoma of aedeagus voluminous, folded, entirely membranous, with several small, minutely dentate eversible lobes (Figs 37, 38).
|
||
</paragraph>
|
||
<paragraph id="186FDA285B61192B282869E545C8689A" lastPageId="11" lastPageNumber="226" pageId="10" pageNumber="225">
|
||
Female. Body large, total length 3.9-4.1. Coloration: As in male, but antennal segment I and femora ranging from uniformly rust brown to totally black (Figure 8). Surface and vestiture: As in subgeneric description; dorsum matt, head and abdomen at most moderately shiny; frons and vertex with few erect spinelike setae; antennal segment I with 6-8 similar setae on mesial surface. Structure: Body 4.4-4.6
|
||
<normalizedToken id="CEC824A225A8A9EB95252BC05DC0FA79" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Head: Vertex 1.9-2.1
|
||
<normalizedToken id="CF075E5E553F485A1AC1890FEC2033C9" originalValue="×">x</normalizedToken>
|
||
as wide as eye; antennal segment I 0.8
|
||
<normalizedToken id="C3E4013EF2BB43E870F76154083C3CDA" originalValue="×">x</normalizedToken>
|
||
as long as length of pronotum, segment II 1.9-2.1
|
||
<normalizedToken id="D42A5E4070DC4F2163A6C473BA4F0BA9" originalValue="×">x</normalizedToken>
|
||
as long as I, 1.1-1.2
|
||
<normalizedToken id="D13BD92FD9FEF853E561068D4E17C1FE" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle, and 0.8-0.9
|
||
<normalizedToken id="C8DE1753D08AC3193A877594AF8191FB" originalValue="×">x</normalizedToken>
|
||
as long as width of head; segment III 1.5-1.6
|
||
<normalizedToken id="6A6D0FF32C9888C8AB0D12572DF6C95F" originalValue="×">x</normalizedToken>
|
||
as long as II; segment IV equal to II in length. Thorax: Pronotum at middle 1.3-1.4
|
||
<normalizedToken id="DF3F471CA9815D57372AA17F8A2BB0A7" originalValue="×">x</normalizedToken>
|
||
as wide as long, 0.7-0.8
|
||
<normalizedToken id="C4209162C46E8D388D0E659554F3A331" originalValue="×">x</normalizedToken>
|
||
as wide as head; hind tibia 3.3-3.5
|
||
<normalizedToken id="C3BFE09F4BEF3444BF9102021F14E05F" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Genitalia: Sclerotized rings of
|
||
<pageBreakToken id="EF0E8ABC271956C6D8062756DEF71898" pageId="11" pageNumber="226" start="start">dorsal</pageBreakToken>
|
||
labiate plate broadly oval (Figs 51, 59), interramal sclerites of posterior wall almost flat, densely covered with minute denticles (Figure 55).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4A77EFBF2988AA28CE861B14D249659F" pageId="11" pageNumber="226" type="distribution">
|
||
<paragraph id="2D0BEB90A48752609D42C54318A77ED2" pageId="11" pageNumber="226">Distribution.</paragraph>
|
||
<paragraph id="9E54ADF4CD7AE169EF3E9AF2B53C6FDB" pageId="11" pageNumber="226">
|
||
The current distribution of
|
||
<taxonomicName id="33BD07CE7E0F55983939EF4B25F874EB" lsidName="M. armeniacus" pageId="11" pageNumber="226" rank="species" species="armeniacus">M. armeniacus</taxonomicName>
|
||
spans a distance of less than 200 km from northeastern Turkey in the west to northwestern Armenia in the east (Figure 64).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |