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<document id="4DAA5A1C513311044E2BDEE2ADF3FA25" ID-DOI="10.11646/zootaxa.4012.2.1" ID-GBIF-Dataset="cfd66b91-8db3-4362-b35c-f7b8e95a2cef" ID-ISSN="1175-5326" ID-Zenodo-Dep="236912" ID-ZooBank="F8A186B9-EE1C-43CC-AB9C-F8CCB1A03794" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461240191041" checkinUser="plazi" docAuthor="Ozerov, A. L. &amp; Krivosheina, M. G." docDate="2015" docId="3A407A65FF9DA71280FAFC59291FFE2A" docLanguage="en" docName="zt04012p257.pdf" docOrigin="Zootaxa 4012 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonarcticus Becker 1894" docType="treatment" docVersion="9" lastPageNumber="212" masterDocId="C679021DFF94A719806DFFE82B63FFA1" masterDocTitle="A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia" masterLastPageNumber="258" masterPageNumber="201" pageNumber="210" updateTime="1698605136340" updateUser="plazi">
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<mods:title id="39A5F89367B3E97A5FEF7C62AEAE383D">A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia</mods:title>
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<mods:title id="BDAB1F5FA7B3C5B3428CD896F09FE5C7">Zootaxa</mods:title>
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<mods:date id="54707C8878B62EC75C7BC11EF3091638">2015</mods:date>
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<treatment id="3A407A65FF9DA71280FAFC59291FFE2A" ID-DOI="http://doi.org/10.5281/zenodo.6096206" ID-GBIF-Taxon="119632792" ID-Zenodo-Dep="6096206" LSID="urn:lsid:plazi:treatment:3A407A65FF9DA71280FAFC59291FFE2A" httpUri="http://treatment.plazi.org/id/3A407A65FF9DA71280FAFC59291FFE2A" lastPageId="11" lastPageNumber="212" pageId="9" pageNumber="210">
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<taxonomicName id="75E9B0F0FF9DA71080FAFC592AB7FC6A" authority="Becker, 1894" authorityName="Becker" authorityYear="1894" box="[151,468,945,971]" class="Insecta" family="Scathophagidae" genus="Gonarcticus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="809D1761FF9DA71080FAFC592AB7FC6A" bold="true" box="[151,468,945,971]" pageId="9" pageNumber="210">
<emphasis id="809D1761FF9DA71080FAFC592A4EFC6A" bold="true" box="[151,301,945,971]" italics="true" pageId="9" pageNumber="210">Gonarcticus</emphasis>
<bibRefCitation id="D678B682FF9DA7108159FC592AB7FC6A" author="Becker" box="[308,468,945,971]" pageId="9" pageNumber="210" refString="Becker, T. (1894) Dipterologische Studien. I. Scatomyzidae. Berliner Entomologische Zeitschrift, 39 (1), 77 - 196. http: // dx. doi. org / 10.1002 / mmnd. 18940390108" type="journal article" year="1894">Becker, 1894</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="B256CB73FF9DA71080FAFC112A2BFB8D" blockId="9.[151,1436,1016,1068]" pageId="9" pageNumber="210">
<treatmentCitationGroup id="92F9EC5DFF9DA71080FAFC112A2BFB8D" pageId="9" pageNumber="210">
<taxonomicName id="75E9B0F0FF9DA71080FAFC112A8BFBAF" authority="Becker, 1894: 103" authorityName="Becker" authorityPageNumber="103" authorityYear="1894" box="[151,488,1016,1038]" class="Insecta" family="Scathophagidae" genus="Gonarcticus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="809D1761FF9DA71080FAFC112A76FBAF" box="[151,277,1017,1038]" italics="true" pageId="9" pageNumber="210">Gonarcticus</emphasis>
<treatmentCitation id="3348ED62FF9DA7108173FC102A8BFBAF" author="Becker" box="[286,488,1016,1038]" page="103" pageId="9" pageNumber="210" year="1894">
<bibRefCitation id="D678B682FF9DA7108173FC102AD3FBAF" author="Becker" box="[286,432,1016,1038]" pageId="9" pageNumber="210" refString="Becker, T. (1894) Dipterologische Studien. I. Scatomyzidae. Berliner Entomologische Zeitschrift, 39 (1), 77 - 196. http: // dx. doi. org / 10.1002 / mmnd. 18940390108" type="journal article" year="1894">Becker, 1894</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
. Gender: masculine.
<typeStatus id="6D5275D1FF9DA71082BFFC112865FBAF" box="[722,774,1017,1038]" pageId="9" pageNumber="210">Type</typeStatus>
species:
<taxonomicName id="75E9B0F0FF9DA7108302FC112E75FBAF" authority="Zetterstedt, 1838" authorityName="Zetterstedt" authorityYear="1838" box="[879,1302,1016,1038]" class="Insecta" family="Scathophagidae" genus="Scatomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="antennata">
<emphasis id="809D1761FF9DA7108302FC112F30FBAF" box="[879,1107,1017,1038]" italics="true" pageId="9" pageNumber="210">Scatomyza antennata</emphasis>
<bibRefCitation id="D678B682FF9DA7108430FC102E75FBAF" author="Zetterstedt" box="[1117,1302,1016,1038]" pageId="9" pageNumber="210" refString="Zetterstedt, J. W. (1838) Sectio tertia. Diptera. In: Insecta Lapponica. Leopold Voss, Lipsiae [= Leipzig], pp. 477 - 868. [' 1840 ']" type="journal article" year="1838">Zetterstedt, 1838</bibRefCitation>
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B256CB73FF9DA71080FAFBB12A45FB37" blockId="9.[151,1437,1113,2038]" pageId="9" pageNumber="210">
The world fauna comprises four species, two of them recorded from the Palaearctic, including
<collectingCountry id="CAFE8BE3FF9DA71084A4FBB12E76FBD3" box="[1225,1301,1113,1138]" name="Russia" pageId="9" pageNumber="210">Russia</collectingCountry>
(
<bibRefCitation id="D678B682FF9DA710854EFBB12BB1FB37" author="Gorodkov" pageId="9" pageNumber="210" refString="Gorodkov, K. B. (1970) Family Scathophagidae (Cordyluridae, Scatomyzidae, Scopeumatidae). In: Bei-Bienko, G. Y. (Ed.), Key to the insects of the European part of the USSR, V. Diptera, Siphonaptera, 2, pp. 440 - 458. [in Russian]" type="journal article" year="1970">Gorodkov, 1970</bibRefCitation>
,
<bibRefCitation id="D678B682FF9DA71080B0FB952A7AFB37" author="Gorodkov" box="[221,281,1149,1174]" pageId="9" pageNumber="210" refString="Gorodkov, K. B. (1986) Family Scathophagidae. In Soos, A. &amp; Papp, L. (Eds), Catalogue of Palaearctic Diptera. Vol. 11. Scathophagidae-Hypodermatidae. Akademiai Kiado, Budapest, pp. 11 - 41." type="book" year="1986">1986</bibRefCitation>
).
</paragraph>
<paragraph id="B256CB73FF9DA71080AAFB4A293BFA17" blockId="9.[151,1437,1113,2038]" pageId="9" pageNumber="210">
<taxonomicName id="75E9B0F0FF9DA71080AAFB4A2A31FB18" box="[199,338,1186,1209]" class="Insecta" family="Scathophagidae" genus="Gonarcticus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="809D1761FF9DA71080AAFB4A2A31FB18" box="[199,338,1186,1209]" italics="true" pageId="9" pageNumber="210">Gonarcticus</emphasis>
</taxonomicName>
species are slender, medium-sized flies, about
<quantity id="75116696FF9DA7108309FB4928A1FB18" box="[868,962,1185,1210]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="210" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long (
<figureCitation id="2AD2D7F6FF9DA7108467FB492F3CFB1B" box="[1034,1119,1185,1210]" captionStart="FIGURES 39 – 44" captionStartId="13.[151,263,1874,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@13.[151,1436,193,1853]" captionTargetPageId="13" captionText="FIGURES 39 – 44. Gonarcticus arcticus (Becker) (39 – 43) and Gonarcticus abdominalis (Zetterstedt) (44): 39 — end of female abdomen, ventral view; 40 — same, dorsal view; 41 — same, lateral view; 42 — male, habitus; 43, 44 — distribution maps." httpUri="https://zenodo.org/record/236919/files/figure.png" pageId="9" pageNumber="210">Fig. 42</figureCitation>
).
<emphasis id="809D1761FF9DA710841EFB492FD3FB1B" box="[1139,1200,1185,1210]" italics="true" pageId="9" pageNumber="210">Head</emphasis>
. Frontal vitta yellow or blackish in upper half and yellow in lower half, frontal plate black, grey dusted. Face, parafacial and gena yellow. Postcranium black completely or black in upper half and yellow in lower half, greyish dusted, covered with yellow hairs in lower half or third, with black postocular setulae and black setae in upper part. Chaetotaxy: 3 orbital, 4–5 frontal, 1 ocellar, 1 inner vertical, 1 outer vertical, 1 postocellar; 1 pair of strong vibrissae and 2–4 pairs of subvibrissae present. Antenna yellow or black. Postpedicel with slightly acute upper apical corner, approximately 2.5–3.0 times as long as wide. Arista black, bare or pubescent throughout its length. Palpus long, yellow completely or darkened apically.
</paragraph>
<paragraph id="B256CB73FF9DA71080AAFA292E01F92C" blockId="9.[151,1437,1113,2038]" pageId="9" pageNumber="210">
<emphasis id="809D1761FF9DA71080AAFA292A7BFA7B" box="[199,280,1473,1498]" italics="true" pageId="9" pageNumber="210">Thorax</emphasis>
black, grey dusted. Chaetotaxy of scutum: acrostichals in two rows, prescutellar pair slightly longer than the other acrostichals, dorsocentrals (3–4)+3, intra-alars 1+2, supra-alars 1+2, postpronotals 2, notopleurals 2, postalars 2. Proepisternum with hairs in central part, with 1 seta near lower margin. Proepimeron with 1 seta. Anepisternum covered with hairs in posterior half and with 2–5 black setae along posterior margin. Katepisternum covered with hairs in ventral corner and with 2 or 3 strong setae; in
<taxonomicName id="75E9B0F0FF9DA71083CCF9BA2F79F9C8" box="[929,1050,1618,1641]" class="Insecta" family="Scathophagidae" genus="Gonarcticus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="arcticus">
<emphasis id="809D1761FF9DA71083CCF9BA2F79F9C8" box="[929,1050,1618,1641]" italics="true" pageId="9" pageNumber="210">G. arcticus</emphasis>
</taxonomicName>
anterior katepisternal seta absent. Anepimeron bare. Scutellum black, grey dusted, with pairs of strong basal scutellar and apical scutellar setae.
</paragraph>
<paragraph id="B256CB73FF9DA71080AAF97228F6F857" blockId="9.[151,1437,1113,2038]" pageId="9" pageNumber="210">
<emphasis id="809D1761FF9DA71080AAF9722B9FF910" box="[199,252,1690,1713]" italics="true" pageId="9" pageNumber="210">Legs</emphasis>
usually completely yellow, but in
<taxonomicName id="75E9B0F0FF9DA71082E4F9722860F910" box="[649,771,1690,1713]" class="Insecta" family="Scathophagidae" genus="Gonarcticus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="arcticus">
<emphasis id="809D1761FF9DA71082E4F9722860F910" box="[649,771,1690,1713]" italics="true" pageId="9" pageNumber="210">G. arcticus</emphasis>
</taxonomicName>
coxae of all legs blackish, tarsi of all legs and fore femur posterodorsally darkened. Fore femur with rows of posterior, posterodorsal and posteroventral setae. Fore tibia with 1–2 posterodorsal, 1–2 anterodorsal and 0–1 posterior setae at middle, and apical setae: 0–1 posterodorsal, 1 dorsal and 1 posterior. Mid femur with row of anterior and posteroventral setae, 2 preapical posterior and 3–4 anteroventral setae on apical half, in
<emphasis id="809D1761FF9DA7108259F8C229CBF8E0" box="[564,680,1834,1857]" italics="true" pageId="9" pageNumber="210">G. a rc t i cu s</emphasis>
additionally with a row of posteroventral setae. Mid tibia with 1–2 anterodorsal, 1–2 posterodorsal, 0–1 posterior (posteroventral), 0–1 anteroventral setae, and ring of apicals.
<collectingCountry id="CAFE8BE3FF9DA7108509F8A52EFFF8C7" box="[1380,1436,1869,1894]" name="India" pageId="9" pageNumber="210">Hind</collectingCountry>
femur with row of anterior (anterodorsal) setae, with 0–3 dorsal setae on apical third, 1 preapical posterior (posterodorsal) seta, 4–5 anteroventral setae on apical half, and usually with a row of thin posterovental setae.
<collectingCountry id="CAFE8BE3FF9DA7108509F87D2EFFF80F" box="[1380,1436,1941,1966]" name="India" pageId="9" pageNumber="210">Hind</collectingCountry>
tibia with 2–3 anterodorsal, 2–3 posterodorsal, 0–1 anteroventral, 1 preapical dorsal setae, and apical setae: 1 anteroventral, 0–1 posteroventral, 1 anterodorsal, 0–1 posterodorsal.
</paragraph>
<caption id="E6969BFBFF9EA71380FAF87F2FB5F84A" httpUri="https://zenodo.org/record/236917/files/figure.png" pageId="10" pageNumber="211" targetBox="[214,1411,226,1880]" targetPageId="10">
<paragraph id="B256CB73FF9EA71380FAF87F2FB5F84A" blockId="10.[151,1436,1943,2027]" pageId="10" pageNumber="211">
<emphasis id="809D1761FF9EA71380FAF87F2A35F80C" bold="true" box="[151,342,1943,1966]" pageId="10" pageNumber="211">FIGURES 22–29.</emphasis>
<taxonomicName id="75E9B0F0FF9EA7138132F87029EEF80F" ID-CoL="8H32B" authority="Ozerov" authorityName="Ozerov" box="[351,653,1944,1966]" class="Insecta" family="Scathophagidae" genus="Cleigastra" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="809D1761FF9EA7138132F870295AF80F" box="[351,569,1944,1966]" italics="true" pageId="10" pageNumber="211">Cleigastra sundukovi</emphasis>
Ozerov
</taxonomicName>
(22–24, 29) and
<taxonomicName id="75E9B0F0FF9EA713832BF8712FACF80F" authority="Zetterstedt" authorityName="Zetterstedt" box="[838,1231,1944,1966]" class="Insecta" family="Scathophagidae" genus="Gonarcticus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="809D1761FF9EA713832BF8712F2BF80F" box="[838,1096,1944,1966]" italics="true" pageId="10" pageNumber="211">Gonarcticus abdominalis</emphasis>
(Zetterstedt)
</taxonomicName>
(25–28): 22—male sternites 4 (lower) and 5 (upper); 23, 26—epandrium, cerci and surstyli, lateral view; 24, 27—same, dorsal view; 25—male sternites 5; 28—male sternites 4; 29—distribution map. Figures 22–24 after Ozerov (2013, Figures 21–23).
</paragraph>
</caption>
<paragraph id="B256CB73FF9FA71280AAFF7129E8FF76" blockId="11.[151,1436,151,395]" pageId="11" pageNumber="212">
<emphasis id="809D1761FF9FA71280AAFF712B9CFF11" box="[199,255,153,176]" italics="true" pageId="11" pageNumber="212">Wing</emphasis>
tinged with brownish; veins blackish; vein R1 bare or setulose on apical third of dorsal surface. Calypters, margins of calypters, and halteres yellowish.
</paragraph>
<paragraph id="B256CB73FF9FA71280AAFF0A2A56FEC6" blockId="11.[151,1436,151,395]" pageId="11" pageNumber="212">
<emphasis id="809D1761FF9FA71280AAFF0A2A52FF5A" box="[199,305,226,251]" italics="true" pageId="11" pageNumber="212">Abdomen</emphasis>
cylindrical, yellow or black, grey dusted, covered with black hairs and setae. Tergites 2–6 each with row of marginal setae. Sternite 4 simple, longer than wide (
<figureCitation id="2AD2D7F6FF9FA712833DFEEE28C9FEBE" box="[848,938,262,287]" captionStart="FIGURES 22 – 29" captionStartId="10.[151,263,1943,1966]" captionTargetBox="[214,1411,226,1880]" captionTargetId="figure@10.[168,1412,193,1902]" captionTargetPageId="10" captionText="FIGURES 22 – 29. Cleigastra sundukovi Ozerov (22 – 24, 29) and Gonarcticus abdominalis (Zetterstedt) (25 – 28): 22 — male sternites 4 (lower) and 5 (upper); 23, 26 — epandrium, cerci and surstyli, lateral view; 24, 27 — same, dorsal view; 25 — male sternites 5; 28 — male sternites 4; 29 — distribution map. Figures 22 – 24 after Ozerov (2013, Figures 21 – 23)." httpUri="https://zenodo.org/record/236917/files/figure.png" pageId="11" pageNumber="212">Figs 28</figureCitation>
,
<figureCitation id="2AD2D7F6FF9FA71283D4FEEF2F65FEBE" box="[953,1030,263,287]" captionStart="FIGURES 30 – 38" captionStartId="12.[151,263,1928,1951]" captionTargetBox="[206,1410,198,1877]" captionTargetId="figure@12.[159,1411,193,1894]" captionTargetPageId="12" captionText="FIGURES 30 – 38. Gonarcticus antennatus (Zetterstedt) (30 – 33, 38), Gonarcticus arcticus (Becker) (34 – 37): 30, 36 — male sternite 5; 31, 34 — epandrium, cerci and surstyli, lateral view; 32, 35 — same, dorsal view; 33, 37 — male sternite 4; 38 — distribution map." httpUri="https://zenodo.org/record/236918/files/figure.png" pageId="11" pageNumber="212">33, 37</figureCitation>
). Sternites 5 as in
<figureCitation id="2AD2D7F6FF9FA7128489FEEE2E5FFEBE" box="[1252,1340,262,287]" captionStart="FIGURES 22 – 29" captionStartId="10.[151,263,1943,1966]" captionTargetBox="[214,1411,226,1880]" captionTargetId="figure@10.[168,1412,193,1902]" captionTargetPageId="10" captionText="FIGURES 22 – 29. Cleigastra sundukovi Ozerov (22 – 24, 29) and Gonarcticus abdominalis (Zetterstedt) (25 – 28): 22 — male sternites 4 (lower) and 5 (upper); 23, 26 — epandrium, cerci and surstyli, lateral view; 24, 27 — same, dorsal view; 25 — male sternites 5; 28 — male sternites 4; 29 — distribution map. Figures 22 – 24 after Ozerov (2013, Figures 21 – 23)." httpUri="https://zenodo.org/record/236917/files/figure.png" pageId="11" pageNumber="212">Figs 25</figureCitation>
,
<figureCitation id="2AD2D7F6FF9FA7128521FEEF2EFBFEBE" box="[1356,1432,262,287]" captionStart="FIGURES 30 – 38" captionStartId="12.[151,263,1928,1951]" captionTargetBox="[206,1410,198,1877]" captionTargetId="figure@12.[159,1411,193,1894]" captionTargetPageId="12" captionText="FIGURES 30 – 38. Gonarcticus antennatus (Zetterstedt) (30 – 33, 38), Gonarcticus arcticus (Becker) (34 – 37): 30, 36 — male sternite 5; 31, 34 — epandrium, cerci and surstyli, lateral view; 32, 35 — same, dorsal view; 33, 37 — male sternite 4; 38 — distribution map." httpUri="https://zenodo.org/record/236918/files/figure.png" pageId="11" pageNumber="212">30, 36</figureCitation>
. Surstyli simple and wide, cerci fused (
<figureCitation id="2AD2D7F6FF9FA712823CFEC229B4FEE2" box="[593,727,298,323]" captionStart="FIGURES 22 – 29" captionStartId="10.[151,263,1943,1966]" captionTargetBox="[214,1411,226,1880]" captionTargetId="figure@10.[168,1412,193,1902]" captionTargetPageId="10" captionText="FIGURES 22 – 29. Cleigastra sundukovi Ozerov (22 – 24, 29) and Gonarcticus abdominalis (Zetterstedt) (25 – 28): 22 — male sternites 4 (lower) and 5 (upper); 23, 26 — epandrium, cerci and surstyli, lateral view; 24, 27 — same, dorsal view; 25 — male sternites 5; 28 — male sternites 4; 29 — distribution map. Figures 22 – 24 after Ozerov (2013, Figures 21 – 23)." httpUri="https://zenodo.org/record/236917/files/figure.png" pageId="11" pageNumber="212">Figs 26, 27</figureCitation>
,
<figureCitation id="2AD2D7F6FF9FA7128288FEC328EAFEE2" box="[741,905,298,323]" captionStart="FIGURES 30 – 38" captionStartId="12.[151,263,1928,1951]" captionTargetBox="[206,1410,198,1877]" captionTargetId="figure@12.[159,1411,193,1894]" captionTargetPageId="12" captionText="FIGURES 30 – 38. Gonarcticus antennatus (Zetterstedt) (30 – 33, 38), Gonarcticus arcticus (Becker) (34 – 37): 30, 36 — male sternite 5; 31, 34 — epandrium, cerci and surstyli, lateral view; 32, 35 — same, dorsal view; 33, 37 — male sternite 4; 38 — distribution map." httpUri="https://zenodo.org/record/236918/files/figure.png" pageId="11" pageNumber="212">31, 32, 34, 35</figureCitation>
). Ovipositor long, cylindrical, not compressed (
<figureCitation id="2AD2D7F6FF9FA71280F2FEA62A44FEC6" box="[159,295,334,359]" captionStart="FIGURES 39 – 44" captionStartId="13.[151,263,1874,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@13.[151,1436,193,1853]" captionTargetPageId="13" captionText="FIGURES 39 – 44. Gonarcticus arcticus (Becker) (39 – 43) and Gonarcticus abdominalis (Zetterstedt) (44): 39 — end of female abdomen, ventral view; 40 — same, dorsal view; 41 — same, lateral view; 42 — male, habitus; 43, 44 — distribution maps." httpUri="https://zenodo.org/record/236919/files/figure.png" pageId="11" pageNumber="212">Figs 39– 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FAF398F8FF9FA71280AAFE9A291FFE2A" box="[199,636,370,395]" pageId="11" pageNumber="212" type="biology_ecology">
<paragraph id="B256CB73FF9FA71280AAFE9A291FFE2A" blockId="11.[151,1436,151,395]" box="[199,636,370,395]" pageId="11" pageNumber="212">Biology of all species is still unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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