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<document id="C4ACB20EC3569287E91A3EF609DA6F8C" ID-CLB-Dataset="299680" ID-DOI="10.11646/zootaxa.5477.1.3" ID-GBIF-Dataset="418a2845-412e-4bbc-9b2b-9040c88d861c" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681470" ID-ZooBank="7FDBA6B8-8D37-44FD-8E12-11FFAF159C36" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1720376501025" checkinUser="plazi" docAuthor="Núñez, Jorge &amp; Martín, Guillermo San" docDate="2024" docId="087487942A7C070169E478C7D6D1FCC6" docLanguage="en" docName="zootaxa.5477.1.3.pdf" docOrigin="Zootaxa 5477 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5477.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exogone martinsosai Núñez &amp; Martín, 2024, n. sp." docType="treatment" docVersion="6" lastPageNumber="52" masterDocId="F44DFFEC2A7B070869737B01D701FFFF" masterDocTitle="Syllidae Grube, 1850 (Annelida) from Conception, Amanay and Banquete Banks off Canary Islands, with the description of two new species" masterLastPageNumber="65" masterPageNumber="43" pageNumber="50" updateTime="1720704280211" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="41FB2BCEAD507B670E0144CED14392F2">Syllidae Grube, 1850 (Annelida) from Conception, Amanay and Banquete Banks off Canary Islands, with the description of two new species</mods:title>
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<mods:role id="7F4ABE621C44B571F4EADCFA03FBFC23">
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<mods:namePart id="A6ACBE417FA5811C3A0073E5C3E7DB15">Núñez, Jorge</mods:namePart>
<mods:affiliation id="8CB796217B16A681793CFFF31EAF551A">Laboratorio de Bentos, Departamento de Biología Animal, Edafología y Geología, Facultad de Ciencias, Universidad de La Laguna, 38200 La Laguna, Tenerife, Canary Islands, Spain</mods:affiliation>
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<mods:namePart id="C70AE698952F560AA15CE76F2152219A">Martín, Guillermo San</mods:namePart>
<mods:nameIdentifier id="70C3ED8296078FC571E96C79623D6036" type="ORCID">0000-0002-8360-6221</mods:nameIdentifier>
<mods:nameIdentifier id="958FBC5CFBB60E8D5D25AB341F20D3CC" type="email">guillermo.sanmartin@uam.es</mods:nameIdentifier>
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<mods:title id="F52050FD1D028809B9C84FC0D4A9B623">Zootaxa</mods:title>
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<mods:part id="51143A4B636E7ED37F5F0DA40B95C89C">
<mods:date id="7B9902F1A2E0C64008C6C21F675A6545">2024</mods:date>
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<mods:number id="39093C1F0FE23E7781FC817D0F44E9B0">2024-07-05</mods:number>
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<mods:url id="D58A21414E50984E040FF4546D5A8CEA">http://dx.doi.org/10.11646/zootaxa.5477.1.3</mods:url>
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<mods:classification id="F6AB548F4B0B058761232AA18A92A181">journal article</mods:classification>
<mods:identifier id="675627BF539632359B44C846F33B6180" type="CLB-Dataset">299680</mods:identifier>
<mods:identifier id="7E9CEA368A46124DBD473B405B90F8B8" type="DOI">10.11646/zootaxa.5477.1.3</mods:identifier>
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<treatment id="087487942A7C070169E478C7D6D1FCC6" ID-DOI="http://doi.org/10.5281/zenodo.12681616" ID-GBIF-Taxon="235037301" ID-Zenodo-Dep="12681616" LSID="urn:lsid:plazi:treatment:087487942A7C070169E478C7D6D1FCC6" httpUri="http://treatment.plazi.org/id/087487942A7C070169E478C7D6D1FCC6" lastPageId="9" lastPageNumber="52" pageId="7" pageNumber="50">
<subSubSection id="C8C765092A7C070F69E478C7D6E3FC1F" box="[151,482,966,992]" pageId="7" pageNumber="50" type="nomenclature">
<paragraph id="806236822A7C070F69E478C7D6E3FC1F" blockId="7.[151,482,966,1028]" box="[151,482,966,992]" pageId="7" pageNumber="50">
<heading id="DB2A81EE2A7C070F69E478C7D6E3FC1F" bold="true" box="[151,482,966,992]" fontSize="11" level="1" pageId="7" pageNumber="50" reason="1">
<emphasis id="B2A9EA902A7C070F69E478C7D6E3FC1F" bold="true" box="[151,482,966,992]" pageId="7" pageNumber="50">
<taxonomicName id="47DD4D012A7C070F69E478C7D699FC1F" authorityName="Núñez &amp; Martín" authorityYear="2024" box="[151,408,966,992]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="50" phylum="Annelida" rank="species" species="martinsosai" status="sp. nov.">
<emphasis id="B2A9EA902A7C070F69E478C7D699FC1F" bold="true" box="[151,408,966,992]" italics="true" pageId="7" pageNumber="50">Exogone martinsosai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A99A57EB2A7C070F68EC78C7D6E3FC1F" box="[415,482,966,992]" pageId="7" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8C765092A7C070F69E478EBD7F4FBFB" box="[151,245,1002,1028]" pageId="7" pageNumber="50" type="description">
<paragraph id="806236822A7C070F69E478EBD7F4FBFB" blockId="7.[151,482,966,1028]" box="[151,245,1002,1028]" pageId="7" pageNumber="50">
<figureCitation id="18E62A072A7C070F69E478EBD7F4FBFB" box="[151,245,1002,1028]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8C765092A7C070F69E47F33D6FCFB6B" pageId="7" pageNumber="50" type="materials_examined">
<paragraph id="806236822A7C070F69E47F33D6FCFB6B" blockId="7.[151,1437,1074,1893]" pageId="7" pageNumber="50">
<emphasis id="B2A9EA902A7C070F69E47F33D685FBB3" bold="true" box="[151,388,1074,1100]" pageId="7" pageNumber="50">Material examined.</emphasis>
<materialsCitation id="30B53CDF2A7C070F68FC7F33D215FBB2" ID-GBIF-Occurrence="4905126308" box="[399,1300,1074,1101]" collectionCode="MNCN" elevation="645" pageId="7" pageNumber="50" specimenCode="MNCN 16.01" specimenCount="1" typeStatus="holotype">
INCOECO station: BC3 0611,
<elevation id="0BF0D1B12A7C070F6B897F33D491FBB3" box="[762,912,1074,1101]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="7" pageNumber="50" unit="m" value="645.0">
<quantity id="47259B672A7C070F6B897F33D444FBB2" box="[762,837,1074,1101]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="7" pageNumber="50" unit="m" value="645.0">645 m</quantity>
depth
</elevation>
,
<typeStatus id="5F6688202A7C070F6AEC7F33D306FBB3" box="[927,1031,1074,1100]" pageId="7" pageNumber="50" type="holotype">Holotype</typeStatus>
(
<specimenCode id="D07B9EF92A7C070F6D697F33D3BBFBB2" box="[1050,1210,1074,1101]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="7" pageNumber="50" type="Museum">MNCN 16.01</specimenCode>
/18955)
</materialsCitation>
;
<materialsCitation id="30B53CDF2A7C070F6C507F33D385FB8F" ID-GBIF-Occurrence="4905126301" collectionCode="MNCN" elevation="237" pageId="7" pageNumber="50" specimenCode="MNCN 16.01" specimenCount="2">
INFUECO station: BC5 0710,
<specimenCount id="96DBFD0B2A7C070F68037F57D6F0FB8E" box="[368,497,1110,1137]" count="2" pageId="7" pageNumber="50" type="generic" typeStatus="paratype">2 Paratypes</specimenCount>
(
<specimenCode id="D07B9EF92A7C070F688C7F57D59BFB8E" box="[511,666,1110,1137]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="7" pageNumber="50" type="Museum">MNCN 16.01</specimenCode>
/18956),
<elevation id="0BF0D1B12A7C070F6B8C7F57D48FFB8F" box="[767,910,1110,1137]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="7" pageNumber="50" unit="m" value="237.0">
<quantity id="47259B672A7C070F6B8C7F57D444FB8E" box="[767,837,1110,1137]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="7" pageNumber="50" unit="m" value="237.0">237 m</quantity>
depth
</elevation>
.
<typeStatus id="5F6688202A7C070F6AE47F57D385FB8F" box="[919,1156,1110,1136]" isOtherTypeStatus="true" pageId="7" pageNumber="50">
<emphasis id="B2A9EA902A7C070F6AE47F57D385FB8F" bold="true" box="[919,1156,1110,1136]" pageId="7" pageNumber="50">Additional material</emphasis>
</typeStatus>
</materialsCitation>
<emphasis id="B2A9EA902A7C070F6DF67F57D38BFB8F" bold="true" box="[1157,1162,1110,1136]" pageId="7" pageNumber="50">.</emphasis>
INFUECO station: BC6 0611,
<specimenCount id="96DBFD0B2A7C070F69AF7F7BD65FFB6B" box="[220,350,1146,1172]" count="1" pageId="7" pageNumber="50" type="generic">1 specimen</specimenCount>
,
<quantity id="47259B672A7C070F681A7F7BD6B1FB6A" box="[361,432,1146,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="50" unit="m" value="254.0">254 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C8C765092A7C070F69B47F9FD380F89B" pageId="7" pageNumber="50" type="description">
<paragraph id="806236822A7C070F69B47F9FD380F89B" blockId="7.[151,1437,1074,1893]" pageId="7" pageNumber="50">
<emphasis id="B2A9EA902A7C070F69B47F9FD650FB47" bold="true" box="[199,337,1182,1208]" pageId="7" pageNumber="50">Description</emphasis>
. Body long, slender, filiform;
<typeStatus id="5F6688202A7C070F6BC07F9FD41AFB47" box="[691,795,1182,1208]" pageId="7" pageNumber="50" type="holotype">Holotype</typeStatus>
incomplete,
<quantity id="47259B672A7C070F6AC27F9FD30BFB46" box="[945,1034,1182,1209]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="50" unit="mm" value="3.7">3.7 mm</quantity>
long,
<quantity id="47259B672A7C070F6D267F9FD3AFFB46" box="[1109,1198,1182,1209]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="50" unit="mm" value="0.1">0.1 mm</quantity>
wide, 38 chaetigers; longest
<typeStatus id="5F6688202A7C070F69867FC2D656FB22" box="[245,343,1219,1245]" pageId="7" pageNumber="50" type="paratype">Paratype</typeStatus>
complete,
<quantity id="47259B672A7C070F68AE7FC3D522FB22" box="[477,547,1218,1245]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="50" unit="mm" value="6.0">6 mm</quantity>
long, 55 chaetigers. Prostomium ovate to subrectangular, wider than long; apparently without eyes (probably lost after fixation). Antennae apparently lost;
<typeStatus id="5F6688202A7C070F6D637FE7D373FAFF" box="[1040,1138,1254,1280]" pageId="7" pageNumber="50" type="holotype">holotype</typeStatus>
with a short, ovate median antenna and
<specimenCount id="96DBFD0B2A7C070F68567E0AD6B5FADA" box="[293,436,1291,1317]" count="1" pageId="7" pageNumber="50" type="generic" typeStatus="paratype">one paratype</specimenCount>
with a similar lateral antenna, all similar in length. Palps broad, longer than prostomium, fused along their length, with a distinct dorsal notch. Peristomium shorter than subsequent segments; tentacular cirri ovoid, small. Dorsal cirri similar to tentacular cirri but longer, shorter than parapodial lobes, absent on chaetiger 2 (
<figureCitation id="18E62A072A7C070F69C67E77D610FA6F" box="[181,273,1398,1424]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2A</figureCitation>
). Anterior three parapodia each with about 6–7 compound falcigers, with spinose shafts and somewhat elongated, bidentate blades, subdistal tooth long and minute, indistinct distal tooth, with fine, moderately long marginal spines, 10 μm long, without compound spiniger-like chaetae (
<figureCitation id="18E62A072A7C070F6AED7EBFD4F5FA27" box="[926,1012,1470,1496]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2B</figureCitation>
). From about chaetiger four, parapodia each with solitary spiniger-like chaeta, shaft enlarged, distally expanded, with numerous spines, and proportionally short, thin, unidentate, filiform, triangular on bases blade, appearing like a long spine under low magnifications, apparently smooth, 35–40 μm long (
<figureCitation id="18E62A072A7C070F6B497D2BD5B7F9BB" box="[570,694,1578,1604]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2C, E</figureCitation>
); in addition, 5–6 (4-
<quantity id="47259B672A7C070F6ADE7D2BD4DBF9BB" box="[941,986,1578,1604]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="50" unit="in" value="3.0">3 in</quantity>
midbody) falcigers, similar to those of anteriormost parapodia but slightly shorter (
<figureCitation id="18E62A072A7C070F6BF07D4FD5FBF997" box="[643,762,1614,1640]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2D, F</figureCitation>
); posterior parapodia each with solitary compound spiniger-like chaetae similar to those of midbody segments (
<figureCitation id="18E62A072A7C070F6BA97D73D434F973" box="[730,821,1650,1676]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2G</figureCitation>
); in addition, three falcigers with short blades, 7–8 μm long (
<figureCitation id="18E62A072A7C070F69A47D97D631F94F" box="[215,304,1686,1712]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2H</figureCitation>
). Dorsal simple chaetae from chaetiger 1, with rounded tips and finely spinulose subterminally, thicker posteriorly (
<figureCitation id="18E62A072A7C070F686C7DBBD68BF92A" box="[287,394,1722,1749]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2I–L</figureCitation>
). Ventral simple chaetae on most posterior parapodia, thick, sigmoid, smooth, bidentate, with small distal tooth and large subdistal tooth (
<figureCitation id="18E62A072A7C070F6BFC7DDFD5F1F907" box="[655,752,1758,1784]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2M</figureCitation>
). A single aciculae per parapodium along the body, slender, distally rounded, apparently hollow in tip (
<figureCitation id="18E62A072A7C070F6BF57C03D5E3F8E3" box="[646,738,1794,1820]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2N</figureCitation>
). Pharynx long, everted in all specimens, through about five segments; pharyngeal tooth long, on anterior rim (
<figureCitation id="18E62A072A7C070F6BB87C27D424F8BF" box="[715,805,1830,1856]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2A</figureCitation>
). Proventricle similar in length to pharynx, through four segments, with 15 very thick muscle cell rows (
<figureCitation id="18E62A072A7C070F6BDF7C4BD406F89B" box="[684,775,1866,1892]" captionStart="FIGURE 2" captionStartId="8.[152,255,1826,1850]" captionTargetBox="[173,1414,190,1794]" captionTargetId="figure-31@8.[164,1423,181,1803]" captionTargetPageId="8" captionText="FIGURE 2. Exogone martinsosai n. sp. A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium.F, falcigers, midbody parapodium.G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="7" pageNumber="50">Fig. 2A</figureCitation>
). Pygidium with 2 long anal cirri.
</paragraph>
</subSubSection>
<caption id="D4A2660A2A73070069EB7C23D56DF810" ID-DOI="http://doi.org/10.5281/zenodo.12681472" ID-Zenodo-Dep="12681472" httpUri="https://zenodo.org/record/12681472/files/figure.png" pageId="8" pageNumber="51" startId="8.[152,255,1826,1850]" targetBox="[173,1414,190,1794]" targetPageId="8" targetType="figure">
<paragraph id="806236822A73070069EB7C23D56DF810" blockId="8.[151,1437,1826,2031]" pageId="8" pageNumber="51">
<emphasis id="B2A9EA902A73070069EB7C23D548F8C4" bold="true" box="[152,585,1826,1851]" pageId="8" pageNumber="51">
FIGURE 2.
<taxonomicName id="47DD4D012A73070068577C22D502F8C4" authorityName="Núñez &amp; Martín" authorityYear="2024" box="[292,515,1827,1851]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="51" phylum="Annelida" rank="species" species="martinsosai" status="sp. nov.">
<emphasis id="B2A9EA902A73070068577C22D502F8C4" bold="true" box="[292,515,1827,1851]" italics="true" pageId="8" pageNumber="51">Exogone martinsosai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A99A57EB2A7307006B7E7C22D548F8C4" box="[525,585,1827,1851]" pageId="8" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A, anterior end, dorsal view. B, falcigers, most anterior parapodium. C, spinigerlike compound chaeta, anterior parapodium. D, falcigers, anterior parapodium. E, spiniger-like compound chaetae, midbody parapodium. F, falcigers, midbody parapodium. G, spiniger-like compound chaetae, posterior parapodium. H, falcigers, posterior parapodium. I, dorsal simple chaeta, most anterior parapodium. J, dorsal simple chaeta, anterior parapodium. K, dorsal simple chaeta, midbody parapodium. L, dorsal simple chaeta, posterior parapodium. M, ventral simple chaeta. N, acicula, posterior parapodium. Scale. A: 0.05 mm. B–N: 20 µm.
</paragraph>
</caption>
<subSubSection id="C8C765092A72070169B47B96D46CFD7A" pageId="9" pageNumber="52" type="discussion">
<paragraph id="806236822A72070169B47B96D7F1FDE6" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="52">
<emphasis id="B2A9EA902A72070169B47B96D63AFF4E" bold="true" box="[199,315,151,177]" pageId="9" pageNumber="52">Remarks.</emphasis>
<taxonomicName id="47DD4D012A72070168317B96D52EFF4F" authorityName="Núñez &amp; Martín" authorityYear="2024" box="[322,559,151,176]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="52" phylum="Annelida" rank="species" species="martinsosai" status="sp. nov.">
<emphasis id="B2A9EA902A72070168317B96D52EFF4F" box="[322,559,151,176]" italics="true" pageId="9" pageNumber="52">Exogone martinsosai</emphasis>
</taxonomicName>
<emphasis id="B2A9EA902A7207016B447B96D577FF4E" bold="true" box="[567,630,151,177]" pageId="9" pageNumber="52">
<taxonomicNameLabel id="A99A57EB2A7207016B447B96D577FF4E" box="[567,630,151,177]" pageId="9" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to a group of this genus sharing some characters, having shafts of spiniger-like compound chaetae distally enlarged, spinose and blades relatively short, triangular, which are absent on most anterior segments, and absence of dorsal cirri on the second chaetiger. The related species are
<taxonomicName id="47DD4D012A7207016C497BDED6E0FEE2" authority="McIntosh, 1885" authorityName="McIntosh" authorityYear="1885" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="52" phylum="Annelida" rank="species" species="heterosetosa">
<emphasis id="B2A9EA902A7207016C497BDED625FEE2" italics="true" pageId="9" pageNumber="52">Exogone heterosetosa</emphasis>
<bibRefCitation id="E44C4B732A72070168587A02D6E0FEE2" author="McIntosh, W. C." box="[299,481,258,285]" pageId="9" pageNumber="52" pagination="1 - 554" refId="ref13936" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger, 12, 1 - 554." type="journal article" year="1885">McIntosh, 1885</bibRefCitation>
</taxonomicName>
, from sub-Antarctic seas,
<collectingCountry id="F8CA76122A7207016A7B7A02D487FEE2" box="[776,902,259,285]" name="Philippines" pageId="9" pageNumber="52">Philippines</collectingCountry>
and
<collectingCountry id="F8CA76122A7207016ACE7A02D325FEE2" box="[957,1060,259,285]" name="Australia" pageId="9" pageNumber="52">Australia</collectingCountry>
(
<bibRefCitation id="E44C4B732A7207016D407A02D3F7FEE2" author="San Martin, G." box="[1075,1270,258,285]" pageId="9" pageNumber="52" pagination="39 - 152" refId="ref15404" refString="San Martin, G. (2005) Exogoninae (Polychaeta: Syllidae) from Australia with the description of a new genus and twenty-two new species. Records of the Australian Museum, 57, 39 - 152. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1438" type="journal article" year="2005">San Martín 2005</bibRefCitation>
;
<bibRefCitation id="E44C4B732A7207016C717A02D7F7FEBE" author="San Martin, G. &amp; Lucas, Y. &amp; Westheide, W." pageId="9" pageNumber="52" pagination="149 - 195" refId="ref15899" refString="San Martin, G., Lucas, Y. &amp; Westheide, W. (2021) The hidden worms on the beach: Interstitial Syllidae (Annelida) from the Indo-Pacific. Proceedings of the Biological Society of Washington, 134, 149 - 195. https: // doi. org / 10.2988 / 0006 - 324 X- 134.1.149" type="journal article" year="2021">
San Martín
<emphasis id="B2A9EA902A7207016CFB7A02D7B5FEBE" italics="true" pageId="9" pageNumber="52">et al.</emphasis>
2021
</bibRefCitation>
),
<taxonomicName id="47DD4D012A720701687B7A26D5EFFEBE" authority="Hartmann-Schroder, 1979" authorityName="Hartmann-Schroder" authorityYear="1979" box="[264,750,295,321]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="52" phylum="Annelida" rank="species" species="heterosetoides">
<emphasis id="B2A9EA902A720701687B7A26D6C6FEBE" box="[264,455,295,321]" italics="true" pageId="9" pageNumber="52">E. heterosetoides</emphasis>
Hartmann-Schröder, 1979
</taxonomicName>
from
<collectingCountry id="F8CA76122A7207016A5C7A26D498FEBE" box="[815,921,295,321]" name="Australia" pageId="9" pageNumber="52">Australia</collectingCountry>
,
<collectingCountry id="F8CA76122A7207016AD17A26D33BFEBE" box="[930,1082,295,321]" name="New Zealand" pageId="9" pageNumber="52">New Zealand</collectingCountry>
,
<collectingCountry id="F8CA76122A7207016D377A26D3C4FEBE" box="[1092,1221,295,321]" name="Philippines" pageId="9" pageNumber="52">Philippines</collectingCountry>
, and southern areas of
<collectingCountry id="F8CA76122A72070169C57A4AD7F2FE9A" box="[182,243,331,357]" name="Chile" pageId="9" pageNumber="52">Chile</collectingCountry>
(
<bibRefCitation id="E44C4B732A72070168717A4AD6C5FE9A" author="San Martin, G." box="[258,452,330,357]" pageId="9" pageNumber="52" pagination="39 - 152" refId="ref15404" refString="San Martin, G. (2005) Exogoninae (Polychaeta: Syllidae) from Australia with the description of a new genus and twenty-two new species. Records of the Australian Museum, 57, 39 - 152. https: // doi. org / 10.3853 / j. 0067 - 1975.57.2005.1438" type="journal article" year="2005">San Martín 2005</bibRefCitation>
;
<bibRefCitation id="E44C4B732A72070168A37A4AD5D3FE9A" author="San Martin, G. &amp; Lucas, Y. &amp; Westheide, W." box="[464,722,331,357]" pageId="9" pageNumber="52" pagination="149 - 195" refId="ref15899" refString="San Martin, G., Lucas, Y. &amp; Westheide, W. (2021) The hidden worms on the beach: Interstitial Syllidae (Annelida) from the Indo-Pacific. Proceedings of the Biological Society of Washington, 134, 149 - 195. https: // doi. org / 10.2988 / 0006 - 324 X- 134.1.149" type="journal article" year="2021">
San Martín
<emphasis id="B2A9EA902A7207016B267A4AD58FFE9A" box="[597,654,331,357]" italics="true" pageId="9" pageNumber="52">et al.</emphasis>
2021
</bibRefCitation>
), and
<taxonomicName id="47DD4D012A7207016A667A4AD299FE9A" authority="Martinez, Adarraga &amp; San Martin, 2002" authorityName="Martinez, Adarraga &amp; San Martin" authorityYear="2002" box="[789,1432,331,357]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="52" phylum="Annelida" rank="species" species="mompasensis">
<emphasis id="B2A9EA902A7207016A667A4AD4CBFE9B" box="[789,970,331,356]" italics="true" pageId="9" pageNumber="52">E. mompasensis</emphasis>
<bibRefCitation id="E44C4B732A7207016AA17A4AD299FE9A" author="Martinez, J. &amp; Adarraga, I. &amp; San Martin, G." box="[978,1432,331,357]" pageId="9" pageNumber="52" pagination="676 - 680" refId="ref13880" refString="Martinez, J., Adarraga, I. &amp; San Martin, G. (2002) Exogone (Exogone) mompasensis (Exogoninae: Syllidae: Polychaeta), a new species of the Iberian Peninsula. Proceedings of the Biological Society of Washington, 115 (3), 676 - 680." type="journal article" year="2002">Martínez, Adarraga &amp; San Martín, 2002</bibRefCitation>
</taxonomicName>
, which is known only from the
<typeStatus id="5F6688202A72070168817A6ED523FE76" box="[498,546,367,393]" pageId="9" pageNumber="52">type</typeStatus>
locality Mompás (Guipúzcoa,
<collectingCountry id="F8CA76122A7207016AF37A6ED4C0FE76" box="[896,961,367,393]" name="Spain" pageId="9" pageNumber="52">Spain</collectingCountry>
) (
<bibRefCitation id="E44C4B732A7207016AAB7A6ED3C2FE76" author="Martinez, J. &amp; Adarraga, I. &amp; San Martin, G." box="[984,1219,367,393]" pageId="9" pageNumber="52" pagination="676 - 680" refId="ref13880" refString="Martinez, J., Adarraga, I. &amp; San Martin, G. (2002) Exogone (Exogone) mompasensis (Exogoninae: Syllidae: Polychaeta), a new species of the Iberian Peninsula. Proceedings of the Biological Society of Washington, 115 (3), 676 - 680." type="journal article" year="2002">
Martínez
<emphasis id="B2A9EA902A7207016D347A6ED381FE76" box="[1095,1152,367,393]" italics="true" pageId="9" pageNumber="52">et al.</emphasis>
2002
</bibRefCitation>
;
<bibRefCitation id="E44C4B732A7207016DBC7A6ED290FE76" author="San Martin, G." box="[1231,1425,367,393]" pageId="9" pageNumber="52" refId="ref15352" refString="San Martin, G. (2003) Familia Syllidae. In: Ramos, M. A. (Ed.), Annelida, Polychaeta II. Fauna Iberica. Vol. 21. Museo Nacional de Ciencias Naturales, CSIC, Madrid, pp. 1 - 554." type="book" year="2003">San Martín 2003</bibRefCitation>
). The new species,
<taxonomicName id="47DD4D012A72070168117A92D551FE53" authorityName="Núñez &amp; Martín" authorityYear="2024" box="[354,592,403,428]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="52" phylum="Annelida" rank="species" species="martinsosai" status="sp. nov.">
<emphasis id="B2A9EA902A72070168117A92D551FE53" box="[354,592,403,428]" italics="true" pageId="9" pageNumber="52">Exogone martinsosai</emphasis>
</taxonomicName>
<emphasis id="B2A9EA902A7207016B2A7A92D598FE52" bold="true" box="[601,665,403,429]" pageId="9" pageNumber="52">
<taxonomicNameLabel id="A99A57EB2A7207016B2A7A92D598FE52" box="[601,665,403,429]" pageId="9" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, has an appreciable smaller thickening of the distal end of the shaft of the spiniger-like chaetae, similar to
<emphasis id="B2A9EA902A7207016B367AB6D5F6FE2E" box="[581,759,439,465]" italics="true" pageId="9" pageNumber="52">
<taxonomicName id="47DD4D012A7207016B367AB6D5F2FE2E" authorityName="McIntosh" authorityYear="1885" box="[581,755,439,465]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="52" phylum="Annelida" rank="species" species="heterosetosa">E. heterosetosa</taxonomicName>
,
</emphasis>
but in the new species the blades are appreciably longer and unidentated. Also it is remarkable the wide muscle cell rows of the proventricle, being much slender in the other species.
</paragraph>
<paragraph id="806236822A72070169B47922D46CFD7A" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="52">
None of the examined specimens have all the antennae; the
<typeStatus id="5F6688202A7207016A017922D4D5FDC2" box="[882,980,547,573]" pageId="9" pageNumber="52" type="holotype">holotype</typeStatus>
has only the median one and one of the
<typeStatus id="5F6688202A72070169E47946D603FD9E" box="[151,258,583,609]" pageId="9" pageNumber="52" type="paratype">paratypes</typeStatus>
has only a lateral one, being similar to the median antenna of the
<typeStatus id="5F6688202A7207016A917946D345FD9E" box="[994,1092,583,609]" pageId="9" pageNumber="52" type="holotype">holotype</typeStatus>
in size; so, we assume that the species has three antennae, short, but longer than the dorsal cirri.
</paragraph>
</subSubSection>
<subSubSection id="C8C765092A72070169B4798ED387FD0E" pageId="9" pageNumber="52" type="etymology">
<paragraph id="806236822A72070169B4798ED4CCFD32" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="52">
<emphasis id="B2A9EA902A72070169B4798ED649FD56" bold="true" box="[199,328,655,681]" pageId="9" pageNumber="52">Etymology</emphasis>
. We dedicate the species to Pablo Martín-Sosa, a researcher at IEO-Canarias who directed the oceanographic campaigns that provided the material studied in this work.
</paragraph>
<paragraph id="806236822A72070169B479D7D387FD0E" blockId="9.[151,1437,151,825]" box="[199,1158,726,753]" pageId="9" pageNumber="52">
<emphasis id="B2A9EA902A72070169B479D7D62BFD0F" bold="true" box="[199,298,726,752]" pageId="9" pageNumber="52">Habitat.</emphasis>
In deep bottoms where coarse sand substrates dominate, from
<quantity id="47259B672A7207016A9979D6D380FD0E" box="[1002,1153,727,753]" metricMagnitude="2" metricUnit="m" metricValue="5.35" metricValueMax="8.33" metricValueMin="2.37" pageId="9" pageNumber="52" unit="m" value="535.0" valueMax="833.0" valueMin="237.0">237 to 833 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C8C765092A72070169B479FBD6D1FCC6" pageId="9" pageNumber="52" type="distribution">
<paragraph id="806236822A72070169B479FBD6D1FCC6" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="52">
<emphasis id="B2A9EA902A72070169B479FBD661FCEA" bold="true" box="[199,352,762,789]" pageId="9" pageNumber="52">Distribution.</emphasis>
Amanay and Banquete Banks, to the south of Fuerteventura and Conception Bank to the north of Lanzarote (
<collectingRegion id="4219F8602A7207016865781ED6C2FCC6" box="[278,451,799,825]" country="Spain" name="Canarias" pageId="9" pageNumber="52">Canary Islands</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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