112 lines
8.4 KiB
XML
112 lines
8.4 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZBK="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva" docDate="2018" docId="9A7A105F8F0252F640D666A6056635E8" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaminus layangensis Schilthuizen & Perreau, sp. n." docType="treatment" docUuid="A840208F-B55E-486C-887F-A26A62243AEB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="91" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="89" updateTime="1668166102793" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perreau, Michel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Njunjic, Iva</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>777</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>108</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.777.23212</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.777.23212</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-777-57</mods:identifier>
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<mods:identifier type="ZBK">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
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<mods:identifier type="ZooBank">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="146746183" LSID="urn:lsid:zoobank.org:act:A840208F-B55E-486C-887F-A26A62243AEB" httpUri="http://treatment.plazi.org/id/9A7A105F8F0252F640D666A6056635E8" lastPageId="34" lastPageNumber="91" pageId="32" pageNumber="89">
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<subSubSection pageId="32" pageNumber="89" type="nomenclature">
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<paragraph pageId="32" pageNumber="89">
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<taxonomicName LSID="http://zoobank.org/A840208F-B55E-486C-887F-A26A62243AEB" authority="Schilthuizen & Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaminus layangensis" order="Coleoptera" pageId="32" pageNumber="89" phylum="Arthropoda" rank="species" species="layangensis">Ptomaphaminus layangensis Schilthuizen & Perreau</taxonomicName>
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<taxonomicNameLabel pageId="32" pageNumber="89">sp. n.</taxonomicNameLabel>
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Figures 11c, 12s, t, 13o, p
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="90" type="reference_group">
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<paragraph pageId="33" pageNumber="90">
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<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus" order="Coleoptera" pageId="33" pageNumber="90" phylum="Arthropoda" rank="genus">
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<pageBreakToken pageId="33" pageNumber="90" start="start">Ptomaphaginus</pageBreakToken>
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</taxonomicName>
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sp. n.
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<taxonomicName class="Insecta" family="Leiodidae" genus="Staphylinoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Staphylinoidea bryanti" order="Coleoptera" pageId="33" pageNumber="90" phylum="Arthropoda" rank="species" species="bryanti">bryanti</taxonomicName>
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complex Merckx et al., 2015: extended data fig. 6.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="90" type="materials_examined">
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<paragraph pageId="33" pageNumber="90">Material.</paragraph>
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<paragraph pageId="33" pageNumber="90">
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Holotype: Malaysia, Sabah, Kinabalu Park, Layang-Layang, 2750 m elev.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="6.0458">6°02.748'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="116.56053">116°33.632'E</geoCoordinate>
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, 13-18.ix.2012 (leg. M. Schilthuizen, Crocker Range / Kinabalu Expedition, RMNH.INS.555619), 1 male. Paratypes: Sabah. Kinabalu Park, Layang-Layang,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="6.0458">6°02.748'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="116.56053">116°33.632'E</geoCoordinate>
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, 2750 m elev., 13-18.ix.2012 (leg. M. Schilthuizen, Crocker Range / Kinabalu Expedition, CMPR, RMNH, incl. RMNH.INS.555618-555622), 11 paratypes.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="34" lastPageNumber="91" pageId="33" pageNumber="90" type="description">
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<paragraph pageId="33" pageNumber="90">Description.</paragraph>
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<paragraph lastPageId="34" lastPageNumber="91" pageId="33" pageNumber="90">
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Length: 1.6-2.5 mm. General colour dark brown; the tarsi and two first antennomeres lighter. Winged. Eyes well developed. Pronotum 1.6 times as wide as long. Elytra 1.45 times as long as wide. Sutural angle of female elytra rounded, without noticeable sexual dimorphism. Male protarsi 0.75 times as wide as the protibia. Spiculum gastrale of the male genital segment short, moderately dilated at the apex (Figure 13o). Aedeagus 5.4 times shorter than the body length, the sides regularly arcuate in dorsal view (Figure 12s), ventrally straight and dorsally obtusely angular in lateral view. Apex of the median lobe with a ventrally deflexed, arcuate hook in lateral
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<pageBreakToken pageId="34" pageNumber="91" start="start">view</pageBreakToken>
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(Figure 12t). Six lateroventral preapical setae and one lateroapical seta on each side. Internal stylus of the endophallus significantly sinuate. Female genital segment with reduced gonocoxites. Spermiduct long and helical (Figure 13p). Apex of spermatheca rounded, without apical sclerotised plate.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="91" type="differential diagnosis">
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<paragraph pageId="34" pageNumber="91">Differential diagnosis.</paragraph>
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<paragraph pageId="34" pageNumber="91">
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Normal-sized species with unreduced eyes and non-spiniform female elytra. Aedeagus with a long, retroverted apical hook. Distinguishable from
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<taxonomicName lsidName="P. alabensis" pageId="34" pageNumber="91" rank="species" species="alabensis">P. alabensis</taxonomicName>
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by the anteriorly widened spiculum gastrale and the broader apex of the median lobe of the aedeagus. Females are distinguishable from
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<taxonomicName lsidName="P. nanus" pageId="34" pageNumber="91" rank="species" species="nanus">P. nanus</taxonomicName>
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by their larger size.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="91" type="dna barcodes">
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<paragraph pageId="34" pageNumber="91">DNA barcodes.</paragraph>
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<paragraph pageId="34" pageNumber="91">COI barcodes are available in BOLD for the holotype, RMNH.INS.555619, as well as for the paratypes RMNH.INS.555618 and 555620-555622, jointly forming the BIN ACK0070.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="91" type="habitat">
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<paragraph pageId="34" pageNumber="91">Habitat and distribution.</paragraph>
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<paragraph pageId="34" pageNumber="91">Known from high altitude (above 2000 m) on Gunung Kinabalu, Sabah, Malaysia.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="91" type="etymology">
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<paragraph pageId="34" pageNumber="91">Etymology.</paragraph>
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<paragraph pageId="34" pageNumber="91">Named for the type locality on Gunung Kinabalu.</paragraph>
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</subSubSection>
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</treatment>
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</document> |