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<document id="0A941BFF1C313B732335624E780DC823" ID-CLB-Dataset="35132" ID-DOI="http://dx.doi.org/10.3897/dez.63.8386" ID-GBIF-Dataset="577b9d1a-d8ef-4c21-8658-1401eb2a0d8d" ID-Pensoft-Pub="1860-1324-1-155" ID-ZooBank="A137A3BD00F540409D7F11F68FC06A2F" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1860-1324-1-155" ModsDocOrigin="Deutsche Entomologische Zeitschrift 63 (1)" ModsDocTitle="A review of the genus Xotidium Löbl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with descriptions of five new species" checkinTime="1464726190790" checkinUser="pensoft" docAuthor="Ogawa, Ryo &amp; Loebl, Ivan" docDate="2016" docId="8A56A55AAB6AECE648197EC6F0DA4070" docLanguage="en" docName="DeutEntomolZeit 63(1): 155-169" docOrigin="Deutsche Entomologische Zeitschrift 63 (1)" docSource="http://dx.doi.org/10.3897/dez.63.8386" docTitle="Xotidium tarantulatum Ogawa &amp; Loebl, sp. n." docType="treatment" docUuid="5704E1D0-9D28-47ED-957B-6D5EB820D440" docUuidSource="ZooBank" docVersion="7" lastPageNumber="165" masterDocId="FFD5FFEC7540FFDBFFFA874BB035FF97" masterDocTitle="A review of the genus Xotidium Loebl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with descriptions of five new species" masterLastPageNumber="169" masterPageNumber="155" pageNumber="163" updateTime="1701378799930" updateUser="ExternalLinkService">
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<mods:title id="D34F38136EB41BE1BB74049D2607149D">A review of the genus Xotidium Loebl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with descriptions of five new species</mods:title>
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<mods:namePart id="63002F70ED97B79A68A27D383BC72044">Ogawa, Ryo</mods:namePart>
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<mods:namePart id="9A914B7EBB37828A0C0FF4CCD5FC6EEF">Loebl, Ivan</mods:namePart>
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<mods:date id="FA560DA09B558F24364E645A8CA572DD">2016</mods:date>
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<mods:identifier id="3C69A803E6265BC9FA6FFB893C8BC4CE" type="DOI">http://dx.doi.org/10.3897/dez.63.8386</mods:identifier>
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<treatment id="8A56A55AAB6AECE648197EC6F0DA4070" ID-GBIF-Taxon="127874123" LSID="urn:lsid:zoobank.org:act:5704E1D0-9D28-47ED-957B-6D5EB820D440" httpUri="http://treatment.plazi.org/id/8A56A55AAB6AECE648197EC6F0DA4070" lastPageId="10" lastPageNumber="165" pageId="8" pageNumber="163">
<subSubSection id="EB54995C9A20FDAB0885FF979D0FA321" pageId="8" pageNumber="163" type="multiple">
<paragraph id="7437E194C4E33E28CAEC3448245F3924" pageId="8" pageNumber="163">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
</subSubSection>
<subSubSection id="F5943F87E9FCC48FAC9AA848674F9C43" pageId="8" pageNumber="163" type="nomenclature">
<paragraph id="6BDF0EC5511FD8C4823AEA58B9DAF298" pageId="8" pageNumber="163">
<taxonomicName id="AFD6837169B4C783A6A67F9C35EFB9CC" ID-CoL="5CGQ4" LSID="http://zoobank.org/5704E1D0-9D28-47ED-957B-6D5EB820D440" authority="Ogawa &amp; Loebl" class="Insecta" family="Staphylinidae" genus="Xotidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xotidium tarantulatum" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="tarantulatum">
Xotidium tarantulatum Ogawa &amp;
<normalizedToken id="5BC1779EA0F4173637017E0C68F57CBD" originalValue="Löbl">Loebl</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="545E21A7E1E189AEF4318B66FCB841E0" pageId="8" pageNumber="163">sp. n.</taxonomicNameLabel>
Figs 2
<normalizedToken id="6EAA6B614BCCF634427BE6E298609048" originalValue="fi">f-i</normalizedToken>
, 5e, f, 6g, 8d, 9e
</paragraph>
</subSubSection>
<subSubSection id="B1577BA5EC9BB4388EC93BEFC6630ADA" lastPageId="9" lastPageNumber="164" pageId="8" pageNumber="163" type="type material">
<paragraph id="626619352F0D6011C73899B1E57620D0" pageId="8" pageNumber="163">Type material.</paragraph>
<paragraph id="C7DBB2D71DDE895DAF730D1334F2061A" pageId="8" pageNumber="163">
Holotype, 1♂, Mt Tilongkabila (Gunung Tilongkabila), N. Sulawesi, alt. ca. 1300m,
<geoCoordinate id="E22686B0CFA70BE847BE6B24AE7CE627" direction="north" orientation="latitude" precision="1" value="0.58837223">
0°35
<normalizedToken id="4B42EC5136A054303FBB0428CC563C37" originalValue="18.14”">'18.14&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="D62C9843D5338730998AC002359AE3F0" direction="east" orientation="longitude" precision="1" value="123.22298">
123°13
<normalizedToken id="030ECFCECE56C7860D7E69C0C5BB8916" originalValue="22.71”">'22.71&quot;</normalizedToken>
E
</geoCoordinate>
, 10. VI. 2012, R. Ogawa leg. (MZBI).
</paragraph>
<paragraph id="CD72A7CC8D15D18EDE5C26F00850204A" lastPageId="9" lastPageNumber="164" pageId="8" pageNumber="163">
Paratypes, 1♂1♀, same data as holotype (EUMJ); 2♂, Mt Tilongkabila (Gunung Tilongkabila), N. Sulawesi, alt. ca. 1300-1500m,
<geoCoordinate id="C4ABCB4176599D8A7897700D037A8B7E" direction="north" orientation="latitude" precision="1" value="0.58837223">
0°35
<normalizedToken id="A9B2695B7BE9E06AE905E95A41438BFB" originalValue="18.14”">'18.14&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="28CD73FD70659F24F0CEFEA9F712A082" direction="east" orientation="longitude" precision="1" value="123.22298">
123°13
<normalizedToken id="1A21A70F13FBCC96FB372B37912F7A8E" originalValue="22.71”">'22.71&quot;</normalizedToken>
E
</geoCoordinate>
-
<geoCoordinate id="6FD13A4FE45A82B1C79F63010DA1D468" direction="north" orientation="latitude" precision="1" value="0.58843607">
0°35
<normalizedToken id="F5B3055B65665EB730F7D00BB2AB8EE8" originalValue="18.37”">'18.37&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="2BFF4F12003B1F5D08306D92775ADF83" direction="east" orientation="longitude" precision="1" value="123.222946">
123°13
<normalizedToken id="94A8A1F07AC7732D9451E954667448AC" originalValue="22.61”">'22.61&quot;</normalizedToken>
E
</geoCoordinate>
, 10. VI. 2012, R. Ogawa leg. (EUMJ); 1♂, same data as holotype, but 800-1300m,
<geoCoordinate id="205531FFC95419703CD4E69A69937D5C" direction="north" orientation="latitude" precision="1" value="0.5745889">
0°34
<normalizedToken id="4C95C42152F8BBDCC98F9B536436BE7E" originalValue="28.52”">'28.52&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="DE05E8EC508B728E2EC2666E17745D28" direction="east" orientation="longitude" precision="1" value="123.19183">
123°11
<normalizedToken id="EC24F7FEB6AEE7B1324AA07A994B0D34" originalValue="30.61”">'30.61&quot;</normalizedToken>
E
</geoCoordinate>
-
<geoCoordinate id="ECA7C972D9B4EE9F451228AB7A2F575E" direction="north" orientation="latitude" precision="1" value="0.58837223">
0°35
<normalizedToken id="8252D5A35EFB5B402F4154C8B6221F9A" originalValue="18.14”">'18.14&quot;</normalizedToken>
N
</geoCoordinate>
, 123°13'
<geoCoordinate id="AFF8EF2018E1205F2D66AAE006FEB8C3" direction="east" orientation="longitude" precision="555" value="22.71">22.71E</geoCoordinate>
, 9. VI. 2012 (EUMJ); 1♂, same data
<pageBreakToken id="4C5B53B4AEDE065EDA98B819EABF7EF4" pageId="9" pageNumber="164" start="start">above</pageBreakToken>
, but preserved at MHNG; 1♂1♀, Mt. Pontolo, N. Sulawesi, alt. ca. 1400-1800m,
<geoCoordinate id="78B6DA927559EA36A3980BF658BB5D7F" direction="north" orientation="latitude" precision="1" value="0.91660273">
0°54
<normalizedToken id="1BF64CF25A6DD19D2781931301E65020" originalValue="59.77”">'59.77&quot;</normalizedToken>
N
</geoCoordinate>
, 122°04'
<geoCoordinate id="3AB6BEDDE6034FA07A4FF3989754E9AE" direction="east" orientation="longitude" precision="555" value="13.1">13.10E</geoCoordinate>
-
<geoCoordinate id="F9A5F914003E96810E5E1C6DC868E91E" direction="north" orientation="latitude" precision="1" value="0.9069639">
0°54
<normalizedToken id="102884024D6FD4B425F3E1CCA3BEE629" originalValue="25.07”">'25.07&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="18593BAE07E492EDCBBC7061117EECFE" direction="east" orientation="longitude" precision="1" value="122.072426">
122°04
<normalizedToken id="ADDFC74CDE7AE4902348F376B1E02061" originalValue="20.73”">'20.73&quot;</normalizedToken>
E
</geoCoordinate>
, 25. VII. 2012, R. Ogawa leg. (EUMJ); 1♂, under bark of rotten log, Plot C, ca 400m, Lowland forest, INDONESIA SULAWESI UTARA, Dumoga-Bone N.P., February, 1985, R. Ent. Soc. Lond., PROJECT WALLACE, B.M. 1985-10 (NHM).
</paragraph>
</subSubSection>
<subSubSection id="9BEFF8C99D8EAF0A851CF1EBE300D8CF" pageId="10" pageNumber="165" type="etymology">
<paragraph id="5309E72E78A704EFE742331A993C49A9" pageId="10" pageNumber="165">
<pageBreakToken id="6F423CC6904BF0D2A170173D27CD6094" pageId="10" pageNumber="165" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph id="FE3B1D1260EC29CD72706883541B8080" pageId="10" pageNumber="165">The species epithet is derived from Mapala Tarantula, the climbing club of Gorontalo State University.</paragraph>
</subSubSection>
<subSubSection id="880FF2D0A8F0B12CEB3C17A8F27AFBFA" pageId="10" pageNumber="165" type="description">
<paragraph id="691033D161009D0FB6711861A65AAF81" pageId="10" pageNumber="165">Description.</paragraph>
<paragraph id="3D32163D9E5906ABE60A77AF48F20FF4" pageId="10" pageNumber="165">
Dorsal and ventral surface almost black to dark reddish-brown (Fig. 5e, f), except for abdomen dark reddish-brown. Antennae almost yellowish-brown, antennomeres
<normalizedToken id="6A771A4A4FACA1B3A8E0E97262F3B91A" originalValue="VIXI">VI-XI</normalizedToken>
blackish. Propygidium and pygidium paler than other ventrite
<normalizedToken id="80D9172FDF2DFB052B42758EF41E1BAD" originalValue="IIII">I-III</normalizedToken>
. Legs reddish-brown; tarsi paler than tibiae and femora. Head, pronotum, and elytra sparsely and finely pubescent.
</paragraph>
<paragraph id="564E5839E8194D2E605AED169F7EFE30" pageId="10" pageNumber="165">Head with eye width almost the same as interocular distance. Punctuation sparse and fine.</paragraph>
<paragraph id="761E7AEDC8A01E4CC54D67BA49F1F0C6" pageId="10" pageNumber="165">Pronotum slightly wider than long, with an anterior bead. Punctuation sparse and fine, as on head. Scutellum almost as long as wide, with exposed apex.</paragraph>
<paragraph id="450011F2A3E4F56964ED585C133844E0" pageId="10" pageNumber="165">Elytra almost as long as wide, widest at basal sixth, lateral margins sharply narrowed apically, minutely serrate at inner part of posterior margin. Punctuation fine and sparse as on pronotum. Sutural striae extending outwards along basal margin to form basal striae, reaching humeral area and joined with lateral striae.</paragraph>
<paragraph id="3E7808271BD30271202DB93D0BF8C742" pageId="10" pageNumber="165">Hypomeron and lateral portion of mesoventrite smooth. Lateral portion of metaventrite finely and sparsely punctate. Metanepisternum about four times as long as wide, without longitudinal line. Mesepimeron almost as long as wide. Metacoxa about six times as wide as metacoxal process. Ventrite I sparsely and finely punctate, with microsculpture.</paragraph>
<paragraph id="0C84FADE8BB8A9484CF114E918D7C676" pageId="10" pageNumber="165">
Protarsomere I twice as long as each
<normalizedToken id="EA0C9DF73C911FBE16996B7E4A1BDC59" originalValue="IIIV">II-IV</normalizedToken>
; V 2.5 times as long as each
<normalizedToken id="9F552498044DCF172017E714EE15ED1E" originalValue="IIIV">II-IV</normalizedToken>
. Mesotarsomere I 2.5 times as long as II; II 1.2 times as long as each III and IV; V 2.5 times as long as each III and IV. Metatarsomere I about three times as long as each II and III; each II and III 1.1 times as long as IV; V about twice as long as IV.
</paragraph>
<paragraph id="8AA00C7F639F0AE8F6610CDBDD413F11" pageId="10" pageNumber="165">
Male. Protarsomeres
<normalizedToken id="E117A38448332690F097B761340101BA" originalValue="IIII">I-III</normalizedToken>
possibly with tenent setae, but invisible at
<normalizedToken id="3B7EFFA66DF1DFE9018B0FB6F38333CD" originalValue="×">x</normalizedToken>
80 magnification, not enlarged. Aedeagus about 0.44 mm long; parameres symmetrical, with weakly widened apical section as long as fourth of total parameral length; internal sac with a sclerite as flagellum, straight, evenly wide, sometimes bent at base.
</paragraph>
<paragraph id="E9A1CF7D6CC7491E127E21122BBBF32A" pageId="10" pageNumber="165">
Female. Protarsomeres
<normalizedToken id="41E512AFEC962FB5C520BF8E7C533826" originalValue="IIII">I-III</normalizedToken>
not enlarged. Ovipositor simple: gonostylus with a long apical seta, distinctly longer than wide; distal gonocoxites apparently elongated (Fig. 9e). Spermathecae undetected.
</paragraph>
</subSubSection>
<subSubSection id="AD472AB9400875EC85FAB98264AF7DEC" pageId="10" pageNumber="165" type="description">
<paragraph id="7C98584C77525F8D0F54B26A6CC6B44D" pageId="10" pageNumber="165">Measurements</paragraph>
<paragraph id="FE015FA76319AA24723AEC9223BEE3E8" pageId="10" pageNumber="165">(n = 9). Length (PL+EL): 1.32-1.43 mm, PW: 0.75-0.79 mm, EW: 0.79-0.89 mm, HW: 0.32-0.36 mm, ID: 0.09-0.13 mm, PL/PW: 0.69-0.76, EL/EW: 0.89-1.02. Approximate ratio of each antennal segment in length (width) (n = 1): II 1.3 (0.7): III 1.0 (0.3): IV 1.1 (0.2): V 1.4 (0.3): VI 1.6 (0.3): VII 1.7 (0.4): VIII 1.6 (0.2): IX 1.9 (0.4): X 1.7 (0.4): XI 2.2 (0.5).</paragraph>
</subSubSection>
<subSubSection id="3834AC2B4CD9E8057C2BA26684CDC103" pageId="10" pageNumber="165" type="distribution">
<paragraph id="30D6A10F8F93A1AABB6382FED97B14AF" pageId="10" pageNumber="165">Distribution.</paragraph>
<paragraph id="C1DF4BBE0F4F7E470105D400A80D7C0E" pageId="10" pageNumber="165">Indonesia: northern Sulawesi.</paragraph>
</subSubSection>
<subSubSection id="DED59589D8A5FE08078B1B6E466EC9C6" pageId="10" pageNumber="165" type="remarks">
<paragraph id="C2B295CCF14876EAD67D12563A9ADA8B" pageId="10" pageNumber="165">Remarks.</paragraph>
<paragraph id="BFA5E565A2994540BC6BE161D193EDD3" pageId="10" pageNumber="165">
This species is similar to
<taxonomicName id="A98714A4C73260D8F822E84E95D4D3B2" class="Insecta" family="Staphylinidae" genus="Xotidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xotidium bolmarum" order="Coleoptera" pageId="10" pageNumber="165" phylum="Arthropoda" rank="species" species="bolmarum">Xotidium bolmarum</taxonomicName>
<normalizedToken id="D09F9DF2350FE29EE3926528ABF317B1" originalValue="Löbl">Loebl</normalizedToken>
from Lombok in its color and body size, but may be easily distinguished by the straight and more elongate sclerite in the internal sac.
</paragraph>
</subSubSection>
</treatment>
</document>