treatments-xml/data/03/D3/87/03D387E9FFB3776BFF6FEB91FBF6FA09.xml

<document id="5ECD6ED580E1CA03A67C4B45CDD4F8CB" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFB3776BFF6FEB91FBF6FA09" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trixagus parvulus Muona 2019, new species" docType="treatment" docVersion="5" lastPageNumber="542" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="541" updateTime="1698730058212" updateUser="plazi">
<mods:mods id="E36C84CC6B18C834B31BE21CF3AFF37E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="08D8D39382850A480ACDF487275CC857">
<mods:title id="FA457266A6BBFFE38C618D76F2816C66">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
</mods:titleInfo>
<mods:name id="112658276B3813CFAA40D5E74E5AC030" type="personal">
<mods:role id="53D8747CC063501747A6488ABC9E60B7">
<mods:roleTerm id="28DA51ED4CC22E97BCD394D0C1497FFC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD23EB620E174C378175DFE0D6AF6C64">Muona, Jyrki</mods:namePart>
</mods:name>
<mods:typeOfResource id="079F89A35763892FDBD7A03F065C4679">text</mods:typeOfResource>
<mods:relatedItem id="E6E5AFC426033D1C7A9A67204AAE04E8" type="host">
<mods:titleInfo id="919D5792156C89D00E1088ACBE3D4ECA">
<mods:title id="75360169EE318407CD72BAB56766E489">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A9E4651FEA5FDD8053876FA21F9D47F9">
<mods:date id="5602E3D14E54F826657EE991AF531467">2019</mods:date>
<mods:detail id="B64B34C6E2A4CE34060CBFEFB899FFC8" type="pubDate">
<mods:number id="038D29C230ABE799AE18D81173CDCE44">2019-04-03</mods:number>
</mods:detail>
<mods:detail id="19C6D16C20DCA8F6A3709F8292F499BE" type="volume">
<mods:number id="97D6B0ACBEF9A4FC2D6BAD11D990AF1F">4576</mods:number>
</mods:detail>
<mods:detail id="2A29461F75BA3D12D66DCBE61821EB80" type="issue">
<mods:number id="D6038D00088CCA74009ADA13438DF383">3</mods:number>
</mods:detail>
<mods:extent id="6666BF38EF79BD46D34ED2695228D765" unit="page">
<mods:start id="8EAC285D7E5C9B41EE15CF9A6119B6A6">521</mods:start>
<mods:end id="F23F75DF6DE78BF55D3BD4DD6655BE54">543</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="042C92A9F23C2CD2032DAE9F5C696D47">journal article</mods:classification>
<mods:identifier id="9C62C6F387FCBB5F09494C0466C861E7" type="DOI">10.11646/zootaxa.4576.3.6</mods:identifier>
<mods:identifier id="33DA870D06214061F1D3882EE667154E" type="GBIF-Dataset">7abdc62c-c350-49dd-8060-4ed64da07362</mods:identifier>
<mods:identifier id="F406165A500B4ADA942E789ED7CACFCC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="293CE5D1387DF60FA38EFC231CD7D335" type="Zenodo-Dep">3715772</mods:identifier>
<mods:identifier id="32F85FAEC38A6A165FBA741D86052F71" type="ZooBank">56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD</mods:identifier>
</mods:mods>
<treatment id="03D387E9FFB3776BFF6FEB91FBF6FA09" ID-DOI="http://doi.org/10.5281/zenodo.3716790" ID-GBIF-Taxon="162639247" ID-Zenodo-Dep="3716790" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09" httpUri="http://treatment.plazi.org/id/03D387E9FFB3776BFF6FEB91FBF6FA09" lastPageId="21" lastPageNumber="542" pageId="20" pageNumber="541">
<subSubSection id="C3606574FFB3776AFF6FEB91FD8AFA4C" box="[151,523,1489,1515]" pageId="20" pageNumber="541" type="nomenclature">
<paragraph id="8BC536FFFFB3776AFF6FEB91FD8AFA4C" blockId="20.[151,523,1489,1549]" box="[151,523,1489,1515]" pageId="20" pageNumber="541">
<heading id="D08D8193FFB3776AFF6FEB91FD8AFA4C" bold="true" box="[151,523,1489,1515]" fontSize="11" level="1" pageId="20" pageNumber="541" reason="1">
<emphasis id="B90EEAEDFFB3776AFF6FEB91FD8AFA4C" bold="true" box="[151,523,1489,1515]" pageId="20" pageNumber="541">
<taxonomicName id="4C7A4D7CFFB3776AFF6FEB91FEF5FA4C" ID-CoL="82NN" ID-ENA="195323" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,372,1489,1515]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB3776AFF6FEB91FEF5FA4C" bold="true" box="[151,372,1489,1515]" italics="true" pageId="20" pageNumber="541">Trixagus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFB3776AFE83EB91FD8AFA4C" box="[379,523,1489,1515]" pageId="20" pageNumber="541" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFB3776AFF6FEBB4FF67F9AA" box="[151,230,1524,1549]" pageId="20" pageNumber="541" type="description">
<paragraph id="8BC536FFFFB3776AFF6FEBB4FF67F9AA" blockId="20.[151,523,1489,1549]" box="[151,230,1524,1549]" pageId="20" pageNumber="541">
<figureCitation id="13412A7AFFB3776AFF6FEBB4FF67F9AA" box="[151,230,1524,1549]" captionStart="FIGURE 15" captionStartId="15.[151,250,971,993]" captionTargetBox="[151,1436,220,934]" captionTargetId="figure@15.[151,1436,220,934]" captionTargetPageId="15" captionText="FIGURE 15. Trixagus parvulus n. sp., head, frontal view. ExE = excised eyes; L = labrum." figureDoi="http://doi.org/10.5281/zenodo.3715804" httpUri="https://zenodo.org/record/3715804/files/figure.png" pageId="20" pageNumber="541">Fig. 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFB3776AFF6FE87CFC06F8AF" pageId="20" pageNumber="541" type="materials_examined">
<paragraph id="8BC536FFFFB3776AFF6FE87CFC06F8AF" blockId="20.[151,1436,1596,1981]" pageId="20" pageNumber="541">
<emphasis id="B90EEAEDFFB3776AFF6FE87CFEC2F9F2" bold="true" box="[151,323,1596,1621]" pageId="20" pageNumber="541">Type material</emphasis>
. 
<typeStatus id="54C1885DFFB3776AFEA0E87CFE41F9F2" box="[344,448,1596,1621]" pageId="20" pageNumber="541" type="holotype">Holotype</typeStatus>
labeled: 
<taxonomicName id="4C7A4D7CFFB3776AFDCBE87CFC82F9F2" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[563,771,1596,1621]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="genus" status="sp. nov.">Trixagus parvulus</taxonomicName>
<emphasis id="B90EEAEDFFB3776AFCF7E87CFCD5F9F2" bold="true" box="[783,852,1596,1621]" pageId="20" pageNumber="541">
<taxonomicNameLabel id="A23D5796FFB3776AFCF7E87CFCD5F9F2" box="[783,852,1596,1621]" pageId="20" pageNumber="541" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ 
<materialsCitation id="3B123CA2FFB3776AFCA5E87CFC02F8AF" ID-GBIF-Occurrence="2578498275" collectingDate="2016-10" collectorName="J. Muona &amp; Age &amp; My. Hirsute &amp; Dorsum &amp; Length" country="Russia" county="Baltic Amber" location="Found" municipality="Embedded" pageId="20" pageNumber="541" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C1885DFFB3776AFCA5E87CFC75F9F3" box="[861,1012,1596,1620]" pageId="20" pageNumber="541" type="holotype">HOLOTYPE</typeStatus>
/ 
<collectorName id="268F5329FFB3776AFC04E87CFBD8F9F3" box="[1020,1113,1596,1620]" pageId="20" pageNumber="541">J.Muona</collectorName>
des. 10-2015. 
<location id="8EA56024FFB3776AFAE9E87CFAD9F9F2" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFAE9E87CFAD9F9F2" box="[1297,1368,1596,1621]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Found" pageId="20" pageNumber="541">Found</location>
from Jantarny.Another label with following data included: 
<location id="8EA56024FFB3776AFD15E820FC52F9DE" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFD15E820FC52F9DE" box="[749,979,1632,1657]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Baltischer Bernstein" pageId="20" pageNumber="541">Baltischer Bernstein</location>
/ 
<location id="8EA56024FFB3776AFC23E820FBB7F9DE" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFC23E820FBB7F9DE" box="[987,1078,1632,1657]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Fundort" pageId="20" pageNumber="541">Fundort</location>
: Balticum. Jantarny (
<collectingCountry id="F36D766FFFB3776AFADBE820FA0DF9DE" box="[1315,1420,1632,1657]" name="Russia" pageId="20" pageNumber="541">Russland</collectingCountry>
)/ 
<location id="8EA56024FFB3776AFF6FE8C4FF54F93A" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFF6FE8C4FF54F93A" box="[151,213,1668,1693]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Alter" pageId="20" pageNumber="541">Alter</location>
: 
<location id="8EA56024FFB3776AFF1AE8C4FEAFF93A" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFF1AE8C4FEAFF93A" box="[226,302,1668,1693]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Tertiar" pageId="20" pageNumber="541">Tertiär</location>
, 
<location id="8EA56024FFB3776AFEC4E8C5FE03F93B" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFEC4E8C5FE03F93B" box="[316,386,1669,1692]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Eozan" pageId="20" pageNumber="541">Eozän</location>
28-54 Mio. Jahre/ 
<location id="8EA56024FFB3776AFDA3E8C4FD3CF93A" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFDA3E8C4FD3CF93A" box="[603,701,1668,1693]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Inklusen" pageId="20" pageNumber="541">Inklusen</location>
: 
<location id="8EA56024FFB3776AFD32E8C4FC8DF93A" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFD32E8C4FC8DF93A" box="[714,780,1668,1693]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Kafer" pageId="20" pageNumber="541">Käfer</location>
, 
<location id="8EA56024FFB3776AFCE2E8C4FC29F93A" LSID="urn:lsid:plazi:treatment:03D387E9FFB3776BFF6FEB91FBF6FA09:8EA56024FFB3776AFCE2E8C4FC29F93A" box="[794,936,1668,1693]" country="Russia" county="Baltic Amber" municipality="Embedded" name="Dungmucke" pageId="20" pageNumber="541">Dungmücke</location>
/www.fossilien.de. 
<collectingMunicipality id="6BA1AC85FFB3776AFB85E8C4FB74F93A" box="[1149,1269,1668,1693]" pageId="20" pageNumber="541">Embedded</collectingMunicipality>
in rectangular piece 
<collectingCounty id="62A44E73FFB3776AFF20E8E8FEF1F966" box="[216,368,1704,1729]" pageId="20" pageNumber="541">Baltic Amber</collectingCounty>
piece, 
<quantity id="4C829B1AFFB3776AFE47E8E8FD8CF967" box="[447,525,1704,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="541" unit="mm" value="19.0">19 mm</quantity>
<date id="FFC4103FFFB3776AFDEBE8E9FDC5F966" box="[531,580,1704,1729]" pageId="20" pageNumber="541" value="2016-10">
<collectingDate id="EF80E9D7FFB3776AFDEBE8E9FDC5F966" box="[531,580,1704,1729]" pageId="20" pageNumber="541" value="2016-10">
x 
<quantity id="4C829B1AFFB3776AFDD0E8E8FDC5F966" box="[552,580,1704,1729]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="541" unit="mm" value="16.0">16</quantity>
</collectingDate>
</date>
mm 
<date id="FFC4103FFFB3776AFD85E8E9FD1EF967" box="[637,671,1704,1728]" pageId="20" pageNumber="541">
<collectingDate id="EF80E9D7FFB3776AFD85E8E9FD1EF967" box="[637,671,1704,1728]" pageId="20" pageNumber="541">
x 
<quantity id="4C829B1AFFB3776AFD69E8E8FD1EF967" box="[657,671,1704,1728]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="541" unit="mm" value="3.0">3</quantity>
</collectingDate>
</date>
mm, sex unknown but most likely female. 
<collectorName id="268F5329FFB3776AFB79E8E9FB31F966" box="[1153,1200,1705,1729]" pageId="20" pageNumber="541">Age</collectorName>
50 
<collectorName id="268F5329FFB3776AFB21E8E9FADFF966" box="[1241,1374,1704,1729]" pageId="20" pageNumber="541">My. Hirsute</collectorName>
plant remains are present as well as a nematoceran dipteran (
<figureCitation id="13412A7AFFB3776AFCC5E88CFC19F942" box="[829,920,1740,1765]" captionStart="FIGURE 26" captionStartId="21.[151,250,1220,1242]" captionTargetBox="[163,1428,188,1209]" captionTargetId="figure@21.[163,1428,188,1209]" captionTargetPageId="21" captionText="FIGURE 26. Trixagus parvulus n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715824" httpUri="https://zenodo.org/record/3715824/files/figure.png" pageId="20" pageNumber="541">Fig. 26</figureCitation>
). 
<collectorName id="268F5329FFB3776AFC4AE88DFB8DF943" box="[946,1036,1741,1764]" pageId="20" pageNumber="541">Dorsum</collectorName>
poorly visible, specimen slightly dorsoventrally compressed, venter entirely visible. 
<collectorName id="268F5329FFB3776AFD2CE8B0FCA2F8AE" box="[724,803,1776,1801]" pageId="20" pageNumber="541">Length</collectorName>
2.0 mm
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFB3776AFF3FE954FBEFF81B" pageId="20" pageNumber="541" type="diagnosis">
<paragraph id="8BC536FFFFB3776AFF3FE954FBEFF81B" blockId="20.[151,1436,1596,1981]" pageId="20" pageNumber="541">
<emphasis id="B90EEAEDFFB3776AFF3FE954FEBBF88A" bold="true" box="[199,314,1812,1837]" pageId="20" pageNumber="541">Diagnosis</emphasis>
. The presence of flattened, grooved tibiae in combination with absence of tarsal grooves places this species in 
<taxonomicName id="4C7A4D7CFFB3776AFEF5E979FEEEF8F7" authorityName="Kugelann" authorityYear="1794" box="[269,367,1849,1872]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEAEDFFB3776AFEF5E979FEEEF8F7" box="[269,367,1849,1872]" italics="true" pageId="20" pageNumber="541">Trixagus</emphasis>
</taxonomicName>
. The distinctly but not deeply emarginated eyes (
<figureCitation id="13412A7AFFB3776AFC59E978FC73F8F6" box="[929,1010,1848,1873]" captionStart="FIGURE 15" captionStartId="15.[151,250,971,993]" captionTargetBox="[151,1436,220,934]" captionTargetId="figure@15.[151,1436,220,934]" captionTargetPageId="15" captionText="FIGURE 15. Trixagus parvulus n. sp., head, frontal view. ExE = excised eyes; L = labrum." figureDoi="http://doi.org/10.5281/zenodo.3715804" httpUri="https://zenodo.org/record/3715804/files/figure.png" pageId="20" pageNumber="541">Fig. 15</figureCitation>
) separate 
<taxonomicName id="4C7A4D7CFFB3776AFB9FE979FB64F8F6" authorityName="Muona" authorityYear="2019" box="[1127,1253,1848,1873]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B90EEAEDFFB3776AFB9FE979FB64F8F6" box="[1127,1253,1848,1873]" italics="true" pageId="20" pageNumber="541">T. parvulus</emphasis>
</taxonomicName>
from all known species except the extant 
<taxonomicName id="4C7A4D7CFFB3776AFE4EE91DFDE7F8D2" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[438,614,1884,1909]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="species" species="dermestoides">
<emphasis id="B90EEAEDFFB3776AFE4EE91DFDE7F8D2" box="[438,614,1884,1909]" italics="true" pageId="20" pageNumber="541">T. dermestoides</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7A4D7CFFB3776AFD8CE91DFD73F8D2" authorityName="Muona" authorityYear="2019" box="[628,754,1884,1909]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B90EEAEDFFB3776AFD8CE91DFD73F8D2" box="[628,754,1884,1909]" italics="true" pageId="20" pageNumber="541">T. parvulus</emphasis>
</taxonomicName>
is smaller, more dorsally convex, has a wider pronotum and less emarginated eyes than 
<taxonomicName id="4C7A4D7CFFB3776AFE32E9C1FDFBF83E" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[458,634,1920,1945]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="species" species="dermestoides">
<emphasis id="B90EEAEDFFB3776AFE32E9C1FDFBF83E" box="[458,634,1920,1945]" italics="true" pageId="20" pageNumber="541">T. dermestoides</emphasis>
</taxonomicName>
. The Eocene fossil species 
<taxonomicName id="4C7A4D7CFFB3776AFC4AE9C1FA03F83E" authority="Kovalev, Kirejtshuk &amp; Nel" authorityName="Kovalev, Kirejtshuk &amp; Nel" box="[946,1410,1920,1945]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="species" species="majusculus">
<emphasis id="B90EEAEDFFB3776AFC4AE9C1FBCAF83E" box="[946,1099,1920,1945]" italics="true" pageId="20" pageNumber="541">T. majusculus</emphasis>
Kovalev, Kirejtshuk &amp; Nel
</taxonomicName>
is quite similar in form, but it is larger and has deeply excised eyes (Kovalev 
<emphasis id="B90EEAEDFFB3776AFC27E9E5FB91F81A" box="[991,1040,1956,1981]" italics="true" pageId="20" pageNumber="541">et al</emphasis>
., 2012).
</paragraph>
</subSubSection>
<caption id="DF056677FFB2776BFF6FEA84FD08FB7D" ID-DOI="http://doi.org/10.5281/zenodo.3715824" ID-Zenodo-Dep="3715824" box="[151,649,1220,1242]" httpUri="https://zenodo.org/record/3715824/files/figure.png" pageId="21" pageNumber="542" startId="21.[151,250,1220,1242]" targetBox="[163,1428,188,1209]" targetPageId="21">
<paragraph id="8BC536FFFFB2776BFF6FEA84FD08FB7D" blockId="21.[151,649,1220,1242]" box="[151,649,1220,1242]" pageId="21" pageNumber="542">
<emphasis id="B90EEAEDFFB2776BFF6FEA84FE9EFB7D" bold="true" box="[151,287,1220,1242]" pageId="21" pageNumber="542">FIGURE 26.</emphasis>
<taxonomicName id="4C7A4D7CFFB2776BFEDDEA85FE5DFB7D" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,476,1220,1242]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="542" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB2776BFEDDEA85FE5DFB7D" box="[293,476,1220,1242]" italics="true" pageId="21" pageNumber="542">Trixagus parvulus</emphasis>
</taxonomicName>
<emphasis id="B90EEAEDFFB2776BFE1BEA84FD9DFB7D" bold="true" box="[483,540,1220,1242]" pageId="21" pageNumber="542">
<taxonomicNameLabel id="A23D5796FFB2776BFE1BEA84FD9DFB7D" box="[483,540,1220,1242]" pageId="21" pageNumber="542" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype.
</paragraph>
</caption>
<subSubSection id="C3606574FFB2776BFF3FEB46FAD1FA2C" pageId="21" pageNumber="542" type="description">
<paragraph id="8BC536FFFFB2776BFF3FEB46FAD1FA2C" blockId="21.[151,1436,1286,1455]" pageId="21" pageNumber="542">
<emphasis id="B90EEAEDFFB2776BFF3FEB46FED0FAB8" bold="true" box="[199,337,1286,1311]" pageId="21" pageNumber="542">Description</emphasis>
. Form fairly wide, pronotum convex, sides evenly rounded to hind angles (
<figureCitation id="13412A7AFFB2776BFB1CEB46FABFFAB8" box="[1252,1342,1286,1311]" captionStart="FIGURE 16" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[151,1436,1060,1917]" captionTargetId="figure@15.[151,1436,1060,1917]" captionTargetPageId="15" captionText="FIGURE 16. Trixagus parvulus n. sp., ventrolateral view." figureDoi="http://doi.org/10.5281/zenodo.3715806" httpUri="https://zenodo.org/record/3715806/files/figure.png" pageId="21" pageNumber="542">Fig. 16</figureCitation>
). Elytra bulbous, with sharp, minutely punctate striae, interstices with much larger, flat, moderately dense punctation (
<figureCitation id="13412A7AFFB2776BFA97EB6AFF37FAC0" captionStart="FIGURE 17" captionStartId="16.[151,250,1107,1129]" captionTargetBox="[171,1420,193,1097]" captionTargetId="figure@16.[171,1420,193,1097]" captionTargetPageId="16" captionText="FIGURE 17. Trixagus parvulus n. sp., dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.3715808" httpUri="https://zenodo.org/record/3715808/files/figure.png" pageId="21" pageNumber="542">Fig. 17</figureCitation>
). Abdomen and metathorax with coarse, moderately dense punctation, hypomera with dense and coarse punctation. Antennae with stout, well-developed club, a9 and a10 transverse, a11 longer than wide (
<figureCitation id="13412A7AFFB2776BFB0FEB32FAC2FA2C" box="[1271,1347,1394,1419]" captionStart="FIGURE 16" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[151,1436,1060,1917]" captionTargetId="figure@15.[151,1436,1060,1917]" captionTargetPageId="15" captionText="FIGURE 16. Trixagus parvulus n. sp., ventrolateral view." figureDoi="http://doi.org/10.5281/zenodo.3715806" httpUri="https://zenodo.org/record/3715806/files/figure.png" pageId="21" pageNumber="542">Fig.16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFB2776BFF3FEBD6FBF6FA09" box="[199,1143,1430,1455]" pageId="21" pageNumber="542" type="etymology">
<paragraph id="8BC536FFFFB2776BFF3FEBD6FBF6FA09" blockId="21.[151,1436,1286,1455]" box="[199,1143,1430,1455]" pageId="21" pageNumber="542">
<emphasis id="B90EEAEDFFB2776BFF3FEBD6FEC7FA08" bold="true" box="[199,326,1430,1455]" pageId="21" pageNumber="542">Etymology</emphasis>
. One of the smallest known 
<taxonomicName id="4C7A4D7CFFB2776BFD7FEBD7FD68FA09" authorityName="Kugelann" authorityYear="1794" box="[647,745,1431,1454]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEAEDFFB2776BFD7FEBD7FD68FA09" box="[647,745,1431,1454]" italics="true" pageId="21" pageNumber="542">Trixagus</emphasis>
</taxonomicName>
species and the smallest fossil one.
</paragraph>
</subSubSection>
</treatment>
</document>