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<document id="ECAB83599CCC6F0404203B432DCBF67C" ID-CLB-Dataset="8782" ID-DOI="10.11646/zootaxa.4822.3.2" ID-GBIF-Dataset="3c15254c-086c-448f-bfe2-e3cd208dcc87" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401617" ID-ZooBank="86DC1241-FE11-49CC-9674-ACAB1DF92F62" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609337343636" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="525F87CCFF90FF989BEDFF75FBCFFB57" docLanguage="en" docName="zootaxa.4822.3.2.pdf" docOrigin="Zootaxa 4822 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nogunius Jałoszyński 2020, gen. n." docType="treatment" docVersion="7" lastPageNumber="350" masterDocId="AE66FFB4FF9CFF889B7AFFE2FFD9FFF7" masterDocTitle="Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="346" updateTime="1698849732245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FFAD41E59BD45AEC8C17D1E389EFE69D">Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae)</mods:title>
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<mods:namePart id="4D38A6BEC800DFAA458EFEB17347AD8E">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="A24529E1B09379F6AE20A67A70203B7B">2020</mods:date>
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<mods:number id="8F9E28F989900D9E328D814D02B70E61">2020-08-06</mods:number>
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<treatment id="525F87CCFF90FF989BEDFF75FBCFFB57" ID-DOI="http://doi.org/10.5281/zenodo.4450827" ID-GBIF-Taxon="177009892" ID-Zenodo-Dep="4450827" LSID="urn:lsid:plazi:treatment:525F87CCFF90FF989BEDFF75FBCFFB57" httpUri="http://treatment.plazi.org/id/525F87CCFF90FF989BEDFF75FBCFFB57" lastPageId="16" lastPageNumber="350" pageId="12" pageNumber="346">
<subSubSection id="92EC6551FF90FF849BEDFF75FEB1FF46" box="[151,360,151,177]" pageId="12" pageNumber="346" type="nomenclature">
<paragraph id="DA4936DAFF90FF849BEDFF75FEB1FF46" blockId="12.[151,360,151,177]" box="[151,360,151,177]" pageId="12" pageNumber="346">
<heading id="810181B6FF90FF849BEDFF75FEB1FF46" bold="true" box="[151,360,151,177]" fontSize="11" level="1" pageId="12" pageNumber="346" reason="1">
<emphasis id="E882EAC8FF90FF849BEDFF75FEB1FF46" bold="true" box="[151,360,151,177]" pageId="12" pageNumber="346">
<taxonomicName id="1DF64D59FF90FF849BEDFF75FED7FF46" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,270,151,177]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E882EAC8FF90FF849BEDFF75FED7FF46" bold="true" box="[151,270,151,177]" italics="true" pageId="12" pageNumber="346">Nogunius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF90FF849A6CFF75FEB1FF46" box="[278,360,151,177]" pageId="12" pageNumber="346" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF90FF849BEDFF3DFD6FFF0F" box="[151,694,222,249]" pageId="12" pageNumber="346" type="materials_examined">
<paragraph id="DA4936DAFF90FF849BEDFF3DFD6FFF0F" blockId="12.[151,1437,222,2009]" box="[151,694,222,249]" pageId="12" pageNumber="346">
<typeStatus id="054D8878FF90FF849BEDFF3DFF16FF0E" box="[151,207,223,249]" pageId="12" pageNumber="346">Type</typeStatus>
species:
<taxonomicName id="1DF64D59FF90FF849A4FFF3DFE28FF0F" authorityName="Jałoszyński" authorityYear="2020" box="[309,497,222,248]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="species" species="sokani">
<emphasis id="E882EAC8FF90FF849A4FFF3DFE28FF0F" box="[309,497,222,248]" italics="true" pageId="12" pageNumber="346">Nogunius sokani</emphasis>
</taxonomicName>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF90FF849BBDFEE3FB1DFD35" pageId="12" pageNumber="346" type="diagnosis">
<paragraph id="DA4936DAFF90FF849BBDFEE3FB1DFD35" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
<emphasis id="E882EAC8FF90FF849BBDFEE3FE9BFEEC" bold="true" box="[199,322,257,283]" pageId="12" pageNumber="346">Diagnosis.</emphasis>
Body elongate, with distinct constriction between head and pronotum and between pronotum and elytra; head short and round, with eyes situated anteriorly, adjacent to antennal fossae, and with long tempora; thick bristles present on head and prothorax; antennal fossae narrowly separated at middle; antenna with large, sharply delimited dimerous club; submentum with lateral sutures continuous with gular sutures; hypostomal ridges incomplete, running only slightly obliquely to lateral margins of cardines and reaching neither submentum nor posterior tentorial pits; pronotum subconical, broadest shortly in front of base and narrowing anterad, with rounded sides; pronotum with two pairs of antebasal pits; prosternum lacking prosternal process, with basisternal portion not demarcated; notosternal sutures distinct in front of procoxal cavities, but postcoxal process of hypomeron firmly fused with postcoxal process of coxosternum without a trace of suture, so that procoxal cavities are closed; hypomeral ridges absent; mesoventral intercoxal process carinate, broad; pleural structures of meso- and metaventrite partly fused laterad mesocoxal projections; metaventral intercoxal process subtrapezoidal, short, with median notch, narrowly separating metacoxae; each elytron with one small but distinct and asetose basal fovea; aedeagus with symmetrical median lobe and endophallus, and with free, slender parameres; spermatheca elongate.
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF90FF869BBDFD29FB8DFE59" lastPageId="14" lastPageNumber="348" pageId="12" pageNumber="346" type="description">
<paragraph id="DA4936DAFF90FF849BBDFD29FC80FCFF" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
<emphasis id="E882EAC8FF90FF849BBDFD29FE80FD12" bold="true" box="[199,345,715,741]" pageId="12" pageNumber="346">Description.</emphasis>
Body form (
<figureCitation id="42CD2A5FFF90FF849A90FD29FDB3FD11" box="[490,618,715,742]" captionStart="FIGURES 4552" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[151,1436,862,1918]" captionTargetId="figure-321@13.[151,1436,849,1918]" captionTargetPageId="13" captionText="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." figureDoi="http://doi.org/10.5281/zenodo.4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="12" pageNumber="346">Figs 4551</figureCitation>
)
<taxonomicName id="1DF64D59FF90FF849903FD2EFD3EFD12" box="[633,743,716,741]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF90FF849903FD2EFD3EFD12" box="[633,743,716,741]" italics="true" pageId="12" pageNumber="346">Euconnus</emphasis>
</taxonomicName>
-like, elongate, moderately slender, with distinct constrictions between head and pronotum and between pronotum and elytra.
</paragraph>
<paragraph id="DA4936DAFF90FF849BBDFCF3FC8EFA52" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
Head (
<figureCitation id="42CD2A5FFF90FF849A6EFCF3FDC4FCDB" box="[276,541,785,812]" captionStart-0="FIGURES 4552" captionStart-1="FIGURES 5359" captionStartId-0="13.[151,264,1942,1966]" captionStartId-1="17.[151,264,1834,1858]" captionTargetBox-0="[151,1436,862,1918]" captionTargetBox-1="[151,1428,580,1800]" captionTargetId-0="figure-321@13.[151,1436,849,1918]" captionTargetId-1="figure-438@17.[151,1436,571,1810]" captionTargetPageId-0="13" captionTargetPageId-1="17" captionText-0="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." captionText-1="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi-0="http://doi.org/10.5281/zenodo.4401631" figureDoi-1="http://doi.org/10.5281/zenodo.4401633" httpUri-0="https://zenodo.org/record/4401631/files/figure.png" httpUri-1="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 46, 48, 50, 5257</figureCitation>
) round and strongly convex; eyes situated closer to mandibular bases than to occipital constriction; vertex not bulging posterodorsad, confluent with frons; frontoclypeal groove present, short; tempora and posterior margin of frons with thick, long bristles. Occipital constriction (
<figureCitation id="42CD2A5FFF90FF849F18FCBAFB6CFC84" box="[1122,1205,856,883]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849FBBFCBBFB31FC85" box="[1217,1256,857,882]" italics="true" pageId="12" pageNumber="346">occ</emphasis>
) about as broad as frons; neck region slightly broadening posterad. Antennal fossae large and situated anterolaterally, narrowly separated at middle. Gular plate transverse, with distinct gular sutures marked on neck region (
<figureCitation id="42CD2A5FFF90FF849FBFFC7CFAC1FC4E" box="[1221,1304,926,953]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849E59FC7DFAE5FC4F" box="[1315,1340,927,952]" italics="true" pageId="12" pageNumber="346">gs</emphasis>
), anteriorly confluent with complete lateral sutures of submentum (
<figureCitation id="42CD2A5FFF90FF84984AFC23FC5AFC2B" box="[816,899,961,988]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF8498F4FC23FC75FC2C" box="[910,940,961,987]" italics="true" pageId="12" pageNumber="346">lss</emphasis>
); posterior tentorial pits (
<figureCitation id="42CD2A5FFF90FF849FBDFC23FAC3FC2B" box="[1223,1306,961,988]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849E5FFC20FA90FC2C" box="[1317,1353,962,987]" italics="true" pageId="12" pageNumber="346">ptp</emphasis>
) small, elongate, adjacent to gular sutures. Submentum (
<figureCitation id="42CD2A5FFF90FF8499CEFC07FCDEFC08" box="[692,775,996,1023]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849868FC07FCE6FC09" box="[786,831,997,1022]" italics="true" pageId="12" pageNumber="346">smn</emphasis>
) subtrapezoidal; mentum (
<figureCitation id="42CD2A5FFF90FF849F10FC07FB65FC08" box="[1130,1212,996,1023]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849FBDFC07FB30FC09" box="[1223,1257,997,1022]" italics="true" pageId="12" pageNumber="346">mn</emphasis>
) subtrapezoidal with concave anterior margin; prementum (
<figureCitation id="42CD2A5FFF90FF8499F8FBEAFD0FFBD4" box="[642,726,1032,1059]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849999FBEBFCCAFBD5" box="[739,787,1033,1058]" italics="true" pageId="12" pageNumber="346">pmn</emphasis>
) largely membranous, flexible, lacking ligula, with moderately broadly separated bases of labial palps and small, inconspicuous lateral lobes of hypopharynx covered with microtrichia; labial palpomere I (
<figureCitation id="42CD2A5FFF90FF849971FBACFD86FB9E" box="[523,607,1102,1129]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849911FBACFD56FB9F" box="[619,655,1102,1128]" italics="true" pageId="12" pageNumber="346">lp1</emphasis>
) short, ring-like, palpomere II (
<figureCitation id="42CD2A5FFF90FF84988FFBACFB90FB9E" box="[1013,1097,1102,1129]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849F2FFBACFBA0FB9F" box="[1109,1145,1102,1128]" italics="true" pageId="12" pageNumber="346">lp2</emphasis>
) elongate, subcylindrical, palpomere III (
<figureCitation id="42CD2A5FFF90FF849A44FB93FE48FB7B" box="[318,401,1137,1164]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849AE6FB93FE66FB7C" box="[412,447,1137,1163]" italics="true" pageId="12" pageNumber="346">lp3</emphasis>
) narrow, elongate, tapering distad and slightly curved. Maxilla (
<figureCitation id="42CD2A5FFF90FF849FF3FB93FB03FB7B" box="[1161,1242,1137,1164]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
) with large cardo (
<figureCitation id="42CD2A5FFF90FF849BDAFB77FF2BFB58" box="[160,242,1172,1199]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849B87FB77FECEFB58" box="[253,279,1173,1199]" italics="true" pageId="12" pageNumber="346">cd</emphasis>
), subtriangular basistipes (
<figureCitation id="42CD2A5FFF90FF849938FB77FD4DFB58" box="[578,660,1172,1199]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF8499E5FB77FD19FB58" box="[671,704,1173,1199]" italics="true" pageId="12" pageNumber="346">bst</emphasis>
), broad mediostipes with an oblique groove demarcating elongate lacinia with a row of robust setae along mesal and distal margin, galea (
<figureCitation id="42CD2A5FFF90FF8498A9FB5AFBF1FB24" box="[979,1064,1208,1235]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849F4CFB5AFB83FB25" box="[1078,1114,1208,1234]" italics="true" pageId="12" pageNumber="346">gal</emphasis>
) elongate, with robust setae along distal and distomesal margin, palpifer (
<figureCitation id="42CD2A5FFF90FF8499F7FB39FD06FB01" box="[653,735,1243,1270]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849990FB39FCD7FB02" box="[746,782,1243,1269]" italics="true" pageId="12" pageNumber="346">ppf</emphasis>
) elongate, maxillary palp with weakly elongate palpomere I (
<figureCitation id="42CD2A5FFF90FF849BCBFB1CFEDFFAEE" box="[177,262,1278,1305]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849A69FB1CFE96FAEF" box="[275,335,1278,1304]" italics="true" pageId="12" pageNumber="346">mxp1</emphasis>
), long, pipe-like palpomere II (
<figureCitation id="42CD2A5FFF90FF8499CCFB1CFCD2FAEE" box="[694,779,1278,1305]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849862FB1CFC8CFAEF" box="[792,853,1278,1304]" italics="true" pageId="12" pageNumber="346">mxp2</emphasis>
), broad and relatively short palpomere III (
<figureCitation id="42CD2A5FFF90FF849E39FB1CFA4EFAEE" box="[1347,1431,1278,1305]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF849BEDFAC0FF0DFACB" box="[151,212,1314,1340]" italics="true" pageId="12" pageNumber="346">mxp3</emphasis>
), and subconical palpomere IV (
<figureCitation id="42CD2A5FFF90FF849938FAC3FD4FFACB" box="[578,662,1313,1340]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 55</figureCitation>
;
<emphasis id="E882EAC8FF90FF8499DBFAC0FD07FACB" box="[673,734,1314,1340]" italics="true" pageId="12" pageNumber="346">mxp4</emphasis>
) with distal half nearly rod-like. Mandibles (
<figureCitation id="42CD2A5FFF90FF849FAFFAC3FA8CFACB" box="[1237,1365,1313,1340]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 54, 59</figureCitation>
) symmetrical, each subtriangular, with broad basal portion and narrow apical tooth, which is indistinctly bifurcate; setose prostheca present but short and indistinct. Labrum (
<figureCitation id="42CD2A5FFF90FF849997FA8AFC98FA74" box="[749,833,1384,1411]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 58</figureCitation>
) transverse, with rounded anterior margin, and with sparse, symmetrically distributed dorsal setae directed anterad.
</paragraph>
<paragraph id="DA4936DAFF90FF849BBDFA4CFA9AF9C5" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
Tentorium (
<figureCitation id="42CD2A5FFF90FF849A33FA4CFE1FFA3E" box="[329,454,1454,1481]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 54, 57</figureCitation>
) with small corporatentorium (
<figureCitation id="42CD2A5FFF90FF849867FA4CFC45FA3E" box="[797,924,1454,1481]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 54, 57</figureCitation>
;
<emphasis id="E882EAC8FF90FF8498DDFA4DFC62FA3F" box="[935,955,1455,1480]" italics="true" pageId="12" pageNumber="346">ct</emphasis>
), bisinuate tentorial bridge (
<figureCitation id="42CD2A5FFF90FF849F8EFA4CFAAAFA3E" box="[1268,1395,1454,1481]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 54, 57</figureCitation>
;
<emphasis id="E882EAC8FF90FF849E07FA4CFA4AFA3F" box="[1405,1427,1454,1480]" italics="true" pageId="12" pageNumber="346">tb</emphasis>
) with anteromedian projection, long gular ridges (
<figureCitation id="42CD2A5FFF90FF8499C3FA33FCE3FA1B" box="[697,826,1489,1516]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 54, 57</figureCitation>
;
<emphasis id="E882EAC8FF90FF84983CFA30FC86FA1C" box="[838,863,1490,1515]" italics="true" pageId="12" pageNumber="346">gr</emphasis>
; with membranous ventral portions and sclerotized dorsal portion, the latter visible in lateral and dorsal view as long arms), lacking dorsal tentorial arms, with short and stout posterior tentorial arms (
<figureCitation id="42CD2A5FFF90FF849963F9FAFDB5F9C4" box="[537,620,1560,1587]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Fig. 57</figureCitation>
;
<emphasis id="E882EAC8FF90FF849902F9FBFD45F9C5" box="[632,668,1561,1586]" italics="true" pageId="12" pageNumber="346">pta</emphasis>
), and long, slender anterior tentorial arms (
<figureCitation id="42CD2A5FFF90FF849FF8F9FAFADAF9C4" box="[1154,1283,1560,1587]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 54, 57</figureCitation>
;
<emphasis id="E882EAC8FF90FF849E75F9FBFAEAF9C5" box="[1295,1331,1561,1586]" italics="true" pageId="12" pageNumber="346">ata</emphasis>
).
</paragraph>
<paragraph id="DA4936DAFF90FF849BBDF9D9FDE9F96B" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
Antenna (
<figureCitation id="42CD2A5FFF90FF849A4CF9D9FE6EF9A1" box="[310,439,1595,1622]" captionStart="FIGURES 5359" captionStartId="17.[151,264,1834,1858]" captionTargetBox="[151,1428,580,1800]" captionTargetId="figure-438@17.[151,1436,571,1810]" captionTargetPageId="17" captionText="FIGURES 5359. Morphological structures of Nogunius sokani sp. n., female. Head in dorsal (5354), ventral (55) and lateral (5657) views; labrum in dorsal view (58); and left mandible in dorsal view (59). Abbreviations: ata, anterior tentorial arm; bst, basistipes; cd, cardo; ct, corporatentorium; fo, foramen occipitale; gal, galea; gr, gular ridge; gs, gular suture (post-tentorial); hr, hypostomal ridge; lac, lacinia; lbr, labrum; lp13, labial palpomere IIII; lss, lateral suture of submentum (= pre-tentorial gular suture); mn, mentum; mxp14, maxillary palpomere IIV; occ, occipital constriction; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; tb, tentorial bridge." figureDoi="http://doi.org/10.5281/zenodo.4401633" httpUri="https://zenodo.org/record/4401633/files/figure.png" pageId="12" pageNumber="346">Figs 53, 56</figureCitation>
) slender, with scape and pedicel each strongly elongate, short and compactly assembled antennomeres IIIVIII, slightly enlarged antennomere IX, and large, sharply delimited dimerous club, with antennomere XI slightly narrower than X.
</paragraph>
<paragraph id="DA4936DAFF90FF849BBDF947FB96F898" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
Prothorax in dorsal view (
<figureCitation id="42CD2A5FFF90FF849A94F947FDA9F948" box="[494,624,1700,1727]" captionStart="FIGURES 4552" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[151,1436,862,1918]" captionTargetId="figure-321@13.[151,1436,849,1918]" captionTargetPageId="13" captionText="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." figureDoi="http://doi.org/10.5281/zenodo.4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="12" pageNumber="346">Figs 4551</figureCitation>
) subconical, broadest shortly in front of base but relatively weakly narrowing anterad. Anterior pronotal corners distinct, posterior corners broadly rounded. Pronotal base with two pairs of distinct pits. Sides of pronotum with thick bristles. Prosternum (
<figureCitation id="42CD2A5FFF90FF8498F2F909FBD0F8F1" box="[904,1033,1771,1798]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="12" pageNumber="346">Figs 6061</figureCitation>
) short in relation to prothorax, with basisternal portion not demarcated and lacking prosternal process, laterally in front of procoxal cavities separated from hypomera by notosternal sutures (
<figureCitation id="42CD2A5FFF90FF84992CF8D3FD72F8BC" box="[598,683,1841,1867]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="12" pageNumber="346">Fig. 61</figureCitation>
;
<emphasis id="E882EAC8FF90FF8499C2F8D0FD05F8BC" box="[696,732,1842,1867]" italics="true" pageId="12" pageNumber="346">nss</emphasis>
), posteriorly lateral process of coxosternum firmly fused with postcoxal process of hypomeron, closing procoxal cavities; hypomeral ridges absent.
</paragraph>
<paragraph id="DA4936DAFF90FF849BBDF89AFD00F82E" blockId="12.[151,1437,222,2009]" pageId="12" pageNumber="346">
Mesonotum (
<figureCitation id="42CD2A5FFF90FF849A24F89AFE68F864" box="[350,433,1912,1939]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="12" pageNumber="346">Fig. 62</figureCitation>
) with mesoscutum (Fig, 62;
<emphasis id="E882EAC8FF90FF849986F89AFCFBF865" box="[764,802,1912,1938]" italics="true" pageId="12" pageNumber="346">sc2</emphasis>
) and mesoscutellum (
<figureCitation id="42CD2A5FFF90FF849F64F89AFBAAF865" box="[1054,1139,1912,1938]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="12" pageNumber="346">Fig. 62</figureCitation>
;
<emphasis id="E882EAC8FF90FF849FFAF89AFB77F865" box="[1152,1198,1912,1938]" italics="true" pageId="12" pageNumber="346">scl2</emphasis>
) demarcated by scutoscutellar suture (
<figureCitation id="42CD2A5FFF90FF849A10F879FE19F842" box="[362,448,1947,1973]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="12" pageNumber="346">Fig. 62</figureCitation>
;
<emphasis id="E882EAC8FF90FF849AB7F87EFE37F842" box="[461,494,1948,1973]" italics="true" pageId="12" pageNumber="346">sss</emphasis>
); only tiny posterior area of scutellar shield visible between elytral bases in intact specimens. Hind wings fully developed, functional.
</paragraph>
<paragraph id="DA4936DAFF91FF859BBDFF75FAC4FE5A" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="347">
Mesoventrite (
<figureCitation id="42CD2A5FFF91FF859A16FF75FE36FF45" box="[364,495,151,178]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Figs 6466</figureCitation>
) transverse, with short prepectus, transverse and setose anteromedian impressions functioning as procoxal rests (
<figureCitation id="42CD2A5FFF91FF859A88FF59FD9EFF22" box="[498,583,186,213]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF85992FFF59FDA3FF23" box="[597,634,187,212]" italics="true" pageId="13" pageNumber="347">pcr</emphasis>
) separated at middle by long, carinate mesoventral intercoxal process (
<figureCitation id="42CD2A5FFF91FF859BDAFF3DFF2DFF0E" box="[160,244,222,249]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF859A7AFF3DFEE3FF0F" box="[256,314,223,248]" italics="true" pageId="13" pageNumber="347">msvp</emphasis>
), which anteriorly nearly reaches anterior margin of mesoventrite. Two shallow foveae can be seen on sides of mesoventrite, each opening on dorsal surface of ventrite (i.e., not opening outside, but inside thorax): fovea 2 (
<figureCitation id="42CD2A5FFF91FF859B86FEC5FE88FEB6" box="[252,337,294,321]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF859A24FEC5FE57FEB6" box="[350,398,295,321]" italics="true" pageId="13" pageNumber="347">fov2</emphasis>
) with opening at base of mesocoxal projection near tip of mesofurca, and fovea 3 (
<figureCitation id="42CD2A5FFF91FF859E39FEC5FA4EFEB6" box="[1347,1431,294,321]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF859BEDFEA9FF11FE92" box="[151,200,331,357]" italics="true" pageId="13" pageNumber="347">fov3</emphasis>
) opening on anteropleural region at the level of lateral corner of procoxal rest. Mesoventral pleural structures partly fused with metaventrite laterad mesocoxal projection (
<figureCitation id="42CD2A5FFF91FF85983EFE8DFC40FE7D" box="[836,921,367,394]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 66</figureCitation>
); the latter (
<figureCitation id="42CD2A5FFF91FF859F5AFE8DFBADFE7E" box="[1056,1140,366,393]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF859FFAFE8DFB76FE7F" box="[1152,1199,367,392]" italics="true" pageId="13" pageNumber="347">mcp</emphasis>
) prominent, strongly projecting laterad, with mesocoxal cavities situated on ventromesal region and exposed in ventral view.
</paragraph>
<paragraph id="DA4936DAFF91FF859BBDFE55FBF7FDCA" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="347">
Metanotum (
<figureCitation id="42CD2A5FFF91FF859A2DFE55FE7EFE25" box="[343,423,439,466]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="13" pageNumber="347">Fig. 63</figureCitation>
) with narrow alacristae (
<figureCitation id="42CD2A5FFF91FF8599C0FE55FCD4FE26" box="[698,781,439,465]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="13" pageNumber="347">Fig. 63</figureCitation>
;
<emphasis id="E882EAC8FF91FF859862FE55FCE3FE26" box="[792,826,439,465]" italics="true" pageId="13" pageNumber="347">alc</emphasis>
), large metascutum (
<figureCitation id="42CD2A5FFF91FF859F59FE55FBACFE26" box="[1059,1141,439,465]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="13" pageNumber="347">Fig. 63</figureCitation>
;
<emphasis id="E882EAC8FF91FF859FFAFE55FB7FFE26" box="[1152,1190,439,465]" italics="true" pageId="13" pageNumber="347">sc3</emphasis>
) with oblique internal apodemes (
<figureCitation id="42CD2A5FFF91FF859A63FE39FEB7FE02" box="[281,366,475,501]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="13" pageNumber="347">Fig. 63</figureCitation>
;
<emphasis id="E882EAC8FF91FF859A06FE39FE41FE03" box="[380,408,475,500]" italics="true" pageId="13" pageNumber="347">ap</emphasis>
), and a broad median membranous area (
<figureCitation id="42CD2A5FFF91FF859802FE39FC14FE02" box="[888,973,475,501]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="13" pageNumber="347">Fig. 63</figureCitation>
;
<emphasis id="E882EAC8FF91FF8598A1FE39FBC8FE03" box="[987,1041,475,500]" italics="true" pageId="13" pageNumber="347">mma</emphasis>
); metascutellum not recognizable; metapostnotum (
<figureCitation id="42CD2A5FFF91FF859A2EFE1DFE71FDEE" box="[340,424,511,537]" captionStart="FIGURES 6063" captionStartId="18.[151,264,1688,1712]" captionTargetBox="[155,1436,535,1660]" captionTargetId="figure-166@18.[151,1436,525,1665]" captionTargetPageId="18" captionText="FIGURES 6063. Morphological structures of Nogunius sokani sp. n., female. Prothorax in ventral view (6063); mesonotum (62) and metanotum (63) in dorsal view. Abbreviations: alc, alacrista; ap, apodeme; mma, median membranous area; nss, noto- sternal suture; pf, profurca; pff, profurcal fovea; ph1, phragma 1; pn3, metapostnotum; pst, prosternum; sc2, mesoscutum; sc3, metascutum; scl2, mesoscutellum; sss, scutoscutellar suture." figureDoi="http://doi.org/10.5281/zenodo.4401635" httpUri="https://zenodo.org/record/4401635/files/figure.png" pageId="13" pageNumber="347">Fig. 63</figureCitation>
;
<emphasis id="E882EAC8FF91FF859ACFFE1DFE06FDEE" box="[437,479,511,537]" italics="true" pageId="13" pageNumber="347">pn3</emphasis>
) short. Elytra (
<figureCitation id="42CD2A5FFF91FF8599F0FE1DFCBBFDEE" box="[650,866,510,537]" captionStart="FIGURES 4552" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[151,1436,862,1918]" captionTargetId="figure-321@13.[151,1436,849,1918]" captionTargetPageId="13" captionText="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." figureDoi="http://doi.org/10.5281/zenodo.4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="13" pageNumber="347">Figs 45, 47, 49, 51</figureCitation>
,
<figureCitation id="42CD2A5FFF91FF859815FE1DFC55FDED" box="[879,908,511,538]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">67</figureCitation>
) together oval, each with small humeral callus and round basal impression with a single asetose basal elytral fovea (
<figureCitation id="42CD2A5FFF91FF8598E6FDC1FC36FDCA" box="[924,1007,547,573]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 67</figureCitation>
;
<emphasis id="E882EAC8FF91FF859881FDC1FBC4FDCA" box="[1019,1053,547,573]" italics="true" pageId="13" pageNumber="347">bef</emphasis>
).
</paragraph>
<paragraph id="DA4936DAFF91FF859BBDFDA5FE39FCE2" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="347">
Metaventrite (
<figureCitation id="42CD2A5FFF91FF859A1EFDA5FE3DFD96" box="[356,484,582,609]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Figs 6465</figureCitation>
) subrectangular, anteriorly fused with mesoventrite, lacking anterior metaventral process, with metaventral intermetacoxal process (
<figureCitation id="42CD2A5FFF91FF8599CAFD89FCDCFD72" box="[688,773,618,645]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF859868FD89FC91FD73" box="[786,840,619,644]" italics="true" pageId="13" pageNumber="347">mtvp</emphasis>
) short, subtrapezoidal, with median notch, narrowly separating metacoxae. Mesofurcal foveae (
<figureCitation id="42CD2A5FFF91FF859900FD6DFD16FD5E" box="[634,719,654,681]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 65</figureCitation>
;
<emphasis id="E882EAC8FF91FF8599A1FD6DFCD3FD5E" box="[731,778,655,681]" italics="true" pageId="13" pageNumber="347">msff</emphasis>
) open in median area of each mesocoxal rest, base of mesofurcal arm bears a deep projection with fovea 1 (
<figureCitation id="42CD2A5FFF91FF8599A3FD51FC85FD3A" box="[729,860,690,717]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Figs 6566</figureCitation>
;
<emphasis id="E882EAC8FF91FF859813FD51FC40FD3A" box="[873,921,691,717]" italics="true" pageId="13" pageNumber="347">fov1</emphasis>
) opening internally and its walls forming additional projection (apophysa) inside pterothorax. Metanepisterna (
<figureCitation id="42CD2A5FFF91FF8598FAFD35FC0AFD06" box="[896,979,727,753]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 66</figureCitation>
;
<emphasis id="E882EAC8FF91FF8598A4FD35FBFEFD06" box="[990,1063,727,753]" italics="true" pageId="13" pageNumber="347">aest23</emphasis>
) and metepimera (
<figureCitation id="42CD2A5FFF91FF859F82FD35FA92FD06" box="[1272,1355,727,753]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="13" pageNumber="347">Fig. 66</figureCitation>
;
<emphasis id="E882EAC8FF91FF859E2CFD35FA4AFD06" box="[1366,1427,727,753]" italics="true" pageId="13" pageNumber="347">epm3</emphasis>
) fully demarcated and narrow.
</paragraph>
<paragraph id="DA4936DAFF91FF859BBDFCFDFDA4FCCE" blockId="13.[151,1437,151,825]" box="[199,637,799,825]" pageId="13" pageNumber="347">Legs slender, lacking peculiar features.</paragraph>
<caption id="8E896652FF91FF859BEDF874FB1FF825" ID-DOI="http://doi.org/10.5281/zenodo.4401631" ID-Zenodo-Dep="4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="13" pageNumber="347" startId="13.[151,264,1942,1966]" targetBox="[151,1436,862,1918]" targetPageId="13">
<paragraph id="DA4936DAFF91FF859BEDF874FB1FF825" blockId="13.[151,1436,1942,2002]" pageId="13" pageNumber="347">
<emphasis id="E882EAC8FF91FF859BEDF874FE8CF859" bold="true" box="[151,341,1942,1966]" pageId="13" pageNumber="347">FIGURES 4552.</emphasis>
Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52).
<taxonomicName id="1DF64D59FF91FF859F88F875FA45F859" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[1266,1436,1942,1966]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="347" phylum="Arthropoda" rank="species" species="sokani" status="sp. nov.">
<emphasis id="E882EAC8FF91FF859F88F875FA45F859" box="[1266,1436,1942,1966]" italics="true" pageId="13" pageNumber="347">Nogunius sokani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF91FF859BEDF858FF17F826" box="[151,206,1978,2001]" pageId="13" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
(4546);
<taxonomicName id="1DF64D59FF91FF859A4BF859FE38F825" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[305,481,1978,2002]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="347" phylum="Arthropoda" rank="species" species="aogashimanus" status="sp. nov.">
<emphasis id="E882EAC8FF91FF859A4BF859FE38F825" box="[305,481,1978,2002]" italics="true" pageId="13" pageNumber="347">N. aogashimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF91FF859A92F858FDC7F826" box="[488,542,1978,2001]" pageId="13" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
(4748);
<taxonomicName id="1DF64D59FF91FF8599F8F859FD09F825" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[642,720,1978,2002]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="347" phylum="Arthropoda" rank="species" species="kerri" status="sp. nov.">
<emphasis id="E882EAC8FF91FF8599F8F859FD09F825" box="[642,720,1978,2002]" italics="true" pageId="13" pageNumber="347">N. kerri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF91FF8599ADF858FCD4F826" box="[727,781,1978,2001]" pageId="13" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
(4950); and
<taxonomicName id="1DF64D59FF91FF8598E6F859FBF2F825" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[924,1067,1978,2002]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="347" phylum="Arthropoda" rank="species" species="fukuuzanus" status="sp. nov.">
<emphasis id="E882EAC8FF91FF8598E6F859FBF2F825" box="[924,1067,1978,2002]" italics="true" pageId="13" pageNumber="347">N. fukuuzanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF91FF859F48F858FBB1F826" box="[1074,1128,1978,2001]" pageId="13" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
(5152).
</paragraph>
</caption>
<paragraph id="DA4936DAFF92FF869BBDFF75FEDEFF22" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
Abdomen (
<figureCitation id="42CD2A5FFF92FF869A3CFF75FE40FF45" box="[326,409,151,178]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="14" pageNumber="348">Fig. 68</figureCitation>
) subtriangular, with six exposed sternites (sternites IIIVIII), suture between VII and VIII indistinct.
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDFF3DFD00FE92" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
Aedeagus (
<figureCitation id="42CD2A5FFF92FF869A3EFF3DFE1AFF0D" box="[324,451,223,250]" captionStart-0="FIGURES 6976" captionStart-1="FIGURES 7784" captionStartId-0="20.[151,264,1925,1949]" captionStartId-1="21.[151,264,1915,1939]" captionTargetBox-0="[152,1424,569,1894]" captionTargetBox-1="[164,1432,189,1882]" captionTargetId-0="figure-170@20.[151,1436,561,1902]" captionTargetId-1="figure-19@21.[151,1436,181,1892]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURES 6976. Aedeagus in ventral (69, 71, 73, 75) and lateral (70, 72, 74, 76) views. Nogunius sokani sp. n. (6972); N. aogashimanus sp. n. (7376)." captionText-1="FIGURES 7784. Aedeagus in ventral (77, 79, 81, 83) and lateral (78, 80, 82, 84) views. Nogunius kerri sp. n. (7780); and N. fukuuzanus sp. n. (8184)." figureDoi-0="http://doi.org/10.5281/zenodo.4401641" figureDoi-1="http://doi.org/10.5281/zenodo.4401643" httpUri-0="https://zenodo.org/record/4401641/files/figure.png" httpUri-1="https://zenodo.org/record/4401643/files/figure.png" pageId="14" pageNumber="348">Figs 6984</figureCitation>
) with symmetrical and nearly tubular median lobe with strongly curved basal region and basal foramen surrounded by large collar formed of parameral bases; ostium situated ventrally in subapical region; endophallus symmetrical and composed of paired sclerites; parameres free (i.e., not fused with walls of median lobe), slender, each with apical and subapical setae.
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDFE8DFC8EFE7E" blockId="14.[151,1437,151,2013]" box="[199,855,367,393]" pageId="14" pageNumber="348">
Spermatheca (
<figureCitation id="42CD2A5FFF92FF869A1DFE8DFE63FE7E" box="[359,442,367,393]" captionStart="FIGURES 6468" captionStartId="19.[151,264,1864,1888]" captionTargetBox="[164,1419,497,1831]" captionTargetId="figure-157@19.[164,1423,489,1841]" captionTargetPageId="19" captionText="FIGURES 6468. Morphological structures of Nogunius sokani sp. n., female. Pterothorax in ventral (6465) and lateral (66) views; elytral base (67); and abdomen in ventral view (68). Abbreviations: aest3, metanepisternum; bef, basal elytral fovea; epm3, metepimeron; fov13, fovea 13; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; msff, mesofurcal fovea; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; sp, spermatheca; v2, mesoven- trite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.4401637" httpUri="https://zenodo.org/record/4401637/files/figure.png" pageId="14" pageNumber="348">Fig. 68</figureCitation>
;
<emphasis id="E882EAC8FF92FF869ABCFE8DFE06FE7F" box="[454,479,367,392]" italics="true" pageId="14" pageNumber="348">sp</emphasis>
) small, thin-walled and elongate.
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDFE71FB8DFE59" blockId="14.[151,1437,151,2013]" box="[199,1108,403,430]" pageId="14" pageNumber="348">Sexual dimorphism distinctly expressed only in size of eyes (smaller in females).</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF92FF869BBDFE55FC3FFD5E" pageId="14" pageNumber="348" type="etymology">
<paragraph id="DA4936DAFF92FF869BBDFE55FB5EFD96" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
<emphasis id="E882EAC8FF92FF869BBDFE55FE97FE26" bold="true" box="[199,334,439,465]" pageId="14" pageNumber="348">Etymology.</emphasis>
This genus is named after Noguni Sôkan, an official who introduced the sweet potato to Okinawa from
<collectingCountry id="A2E1764AFF92FF869BAFFE39FEC1FE02" box="[213,280,475,501]" name="China" pageId="14" pageNumber="348">China</collectingCountry>
in the beginning of the seventeenth century. The sweet potato then spread from
<collectingRegion id="1832F838FF92FF869FE6FE39FAD8FE02" box="[1180,1281,475,501]" country="Japan" name="Okinawa" pageId="14" pageNumber="348">Okinawa</collectingRegion>
to
<collectingCountry id="A2E1764AFF92FF869E5CFE39FAB0FE02" box="[1318,1385,475,501]" name="Japan" pageId="14" pageNumber="348">Japan</collectingCountry>
, and in the Ryukyus it became a staple food that saved many lives from famine. It is not a personal name; sôkan was a manager-level position in the Ryukyu Kingdome, and Noguni is a district of the Kadena town on Okinawa-jima, where he was born; the true name of Noguni Sôkan remains unknown. Gender masculine.
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDFD88FC3FFD5E" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
<emphasis id="E882EAC8FF92FF869BBDFD88FDF6FD73" bold="true" box="[199,559,618,645]" pageId="14" pageNumber="348">Composition and distribution.</emphasis>
<taxonomicName id="1DF64D59FF92FF86994FFD89FD79FD73" authorityName="Jałoszyński" authorityYear="2020" box="[565,672,619,644]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF86994FFD89FD79FD73" box="[565,672,619,644]" italics="true" pageId="14" pageNumber="348">Nogunius</emphasis>
</taxonomicName>
is known to occur in the Ryukyus and Izu Islands; one female representing most likely an undescribed species was found on Shikoku (
<figureCitation id="42CD2A5FFF92FF8698FFFD6DFC00FD5E" box="[901,985,654,681]" captionStart="FIGURE 85" captionStartId="24.[151,250,1259,1283]" captionTargetBox="[151,1429,181,1230]" captionTargetId="figure-15@24.[151,1436,181,1236]" captionTargetPageId="24" captionText="FIGURE 85. Distribution of Nogunius." figureDoi="http://doi.org/10.5281/zenodo.4401647" httpUri="https://zenodo.org/record/4401647/files/figure.png" pageId="14" pageNumber="348">Fig. 85</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF92FF989BBDFD51FBCFFB57" lastPageId="16" lastPageNumber="350" pageId="14" pageNumber="348" type="discussion">
<paragraph id="DA4936DAFF92FF869BBDFD51FCE8FB8A" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
<emphasis id="E882EAC8FF92FF869BBDFD51FEE2FD3A" bold="true" box="[199,315,691,717]" pageId="14" pageNumber="348">Remarks.</emphasis>
Although adults of this genus resemble
<taxonomicName id="1DF64D59FF92FF869870FD51FCA1FD3B" box="[778,888,691,716]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869870FD51FCA1FD3B" box="[778,888,691,716]" italics="true" pageId="14" pageNumber="348">Euconnus</emphasis>
</taxonomicName>
and similar genera of
<taxonomicName id="1DF64D59FF92FF869F06FD51FB27FD3A" authorityName="Ganglbauer" authorityYear="1899" box="[1148,1278,691,717]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
, they have an unusual feature, so far within this tribe known only in
<taxonomicName id="1DF64D59FF92FF86987FFD35FCA7FD06" box="[773,894,727,753]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF86987FFD35FCA7FD06" box="[773,894,727,753]" italics="true" pageId="14" pageNumber="348">Stenichnus</emphasis>
</taxonomicName>
-like genera, i.e., complete lateral sutures of the submentum. In
<taxonomicName id="1DF64D59FF92FF869A31FD19FE6FFCE3" authorityName="Jałoszyński" authorityYear="2020" box="[331,438,763,788]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869A31FD19FE6FFCE3" box="[331,438,763,788]" italics="true" pageId="14" pageNumber="348">Nogunius</emphasis>
</taxonomicName>
, the sutures are clearly identifiable as anterior (pre-tentorial) portions of gular sutures, with posterior tentorial pits situated on their lateral margins far in front of the transverse gular plate on the neck region. The gular ridges extend along gular sutures inside the head capsule. Unambiguous homologization of the lateral sutures of the submentum with gular sutures was so far problematic, because the submental lateral sutures in
<taxonomicName id="1DF64D59FF92FF869BCEFC69FEF4FC52" box="[180,301,907,933]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869BCEFC69FEF4FC52" box="[180,301,907,933]" italics="true" pageId="14" pageNumber="348">Stenichnus</emphasis>
</taxonomicName>
-like genera meet the sutures of the posterior (post-tentorial) portion of the gular plate at some angle, and the posterior tentorial pits are hidden inside a deep transverse groove between the neck region and the anterior, exposed part of the head capsule (e.g.,
<bibRefCitation id="BE674B2BFF92FF86993CFC31FCD2FC1A" author="Jaloszynski, P." box="[582,779,979,1005]" pageId="14" pageNumber="348" pagination="39 - 79" refId="ref16969" refString="Jaloszynski, P. (2013) Revision of subgenera of Stenichnus Thomson, with review of Australo-Pacific species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3630 (1), 39 - 79. https: // doi. org / 10.11646 / zootaxa. 3630.1.2" type="journal article" year="2013">Jałoszyński 2013</bibRefCitation>
). Consequently, each lateral suture could not be unambiguously interpreted as a single, continuous structure extending from the anteroventral margin of the foramen occipitale up to the posterolateral margin of the mentum. The continuity of the suture on the entire ventral surface of the head capsule in
<taxonomicName id="1DF64D59FF92FF869A69FBDDFEA7FBAF" authorityName="Jałoszyński" authorityYear="2020" box="[275,382,1087,1112]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869A69FBDDFEA7FBAF" box="[275,382,1087,1112]" italics="true" pageId="14" pageNumber="348">Nogunius</emphasis>
</taxonomicName>
seems undisputed, and this arrangement is similar to the gular sutures found in many other
<taxonomicName id="1DF64D59FF92FF869BEDFB81FEE8FB8A" box="[151,305,1123,1149]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
(e.g.,
<bibRefCitation id="BE674B2BFF92FF869A02FB81FD0CFB8A" author="Solodovnikov, A. Y. &amp; Newton, A. F." box="[376,725,1122,1149]" pageId="14" pageNumber="348" pagination="398 - 441" refId="ref17355" refString="Solodovnikov, A. Y. &amp; Newton, A. F. (2005) Phylogenetic placement of Arrowinini trib. n. within the subfamily Staphylininae (Coleoptera: Staphylinidae), with revision of the relict South African genus Arrowinus and description of its larva. Systematic Entomology, 30 (3), 398 - 441. https: // doi. org / 10.1111 / j. 1365 - 3113.2004.00283. x" type="journal article" year="2005">Solodovnikov &amp; Newton 2005</bibRefCitation>
: fig. 2.).
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDFB65FC0CF96E" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
Among all genera of
<taxonomicName id="1DF64D59FF92FF869ACAFB65FDEBFB56" authorityName="Ganglbauer" authorityYear="1899" box="[432,562,1159,1185]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
, morphological structures of four are unknown to me:
<taxonomicName id="1DF64D59FF92FF869FF3FB65FA41FB56" authority="Franz" authorityName="Franz" authorityYear="1980" box="[1161,1432,1159,1185]" class="Insecta" family="Staphylinidae" genus="Napochomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869FF3FB65FA89FB56" box="[1161,1360,1159,1185]" italics="true" pageId="14" pageNumber="348">Napochomorphus</emphasis>
Franz
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869BEDFB49FE67FB32" authority="Reitter" authorityName="Reitter" authorityYear="1882" box="[151,446,1195,1221]" class="Insecta" family="Staphylinidae" genus="Neuraphomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869BEDFB49FEB0FB32" box="[151,361,1195,1221]" italics="true" pageId="14" pageNumber="348">Neuraphomorphus</emphasis>
Reitter
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869AB3FB49FD7DFB32" authority="King" authorityName="King" authorityYear="1864" box="[457,676,1195,1221]" class="Insecta" family="Staphylinidae" genus="Psepharobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869AB3FB49FDBBFB32" box="[457,610,1195,1221]" italics="true" pageId="14" pageNumber="348">Psepharobius</emphasis>
King
</taxonomicName>
, and
<taxonomicName id="1DF64D59FF92FF8699A4FB49FBC5FB32" authority="Casey. Each" authorityName="Casey. Each" authorityYear="1897" box="[734,1052,1195,1221]" class="Insecta" family="Staphylinidae" genus="Taphroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF8699A4FB49FC55FB32" box="[734,908,1195,1221]" italics="true" pageId="14" pageNumber="348">Taphroscydmus</emphasis>
Casey. Each
</taxonomicName>
of them includes one species. Despite long efforts, I was not able to borrow the
<typeStatus id="054D8878FF92FF8699DCFB2DFD0FFB1E" box="[678,726,1231,1257]" pageId="14" pageNumber="348">type</typeStatus>
material of the
<typeStatus id="054D8878FF92FF8698F2FB2DFC61FB1E" box="[904,952,1231,1257]" pageId="14" pageNumber="348">type</typeStatus>
species of the Brazilian
<taxonomicName id="1DF64D59FF92FF869FB7FB2DFA4DFB1E" authorityName="Franz" authorityYear="1980" box="[1229,1428,1231,1257]" class="Insecta" family="Staphylinidae" genus="Napochomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869FB7FB2DFA4DFB1E" box="[1229,1428,1231,1257]" italics="true" pageId="14" pageNumber="348">Napochomorphus</emphasis>
</taxonomicName>
; depositories of the
<typeStatus id="054D8878FF92FF869A11FB11FE42FAFA" box="[363,411,1267,1293]" pageId="14" pageNumber="348">type</typeStatus>
material of the
<typeStatus id="054D8878FF92FF869930FB11FDA3FAFA" box="[586,634,1267,1293]" pageId="14" pageNumber="348">type</typeStatus>
species of the Ethiopian
<taxonomicName id="1DF64D59FF92FF8698EBFB11FBBAFAFA" authorityName="Reitter" authorityYear="1882" box="[913,1123,1267,1293]" class="Insecta" family="Staphylinidae" genus="Neuraphomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF8698EBFB11FBBAFAFA" box="[913,1123,1267,1293]" italics="true" pageId="14" pageNumber="348">Neuraphomorphus</emphasis>
</taxonomicName>
, the Australian
<taxonomicName id="1DF64D59FF92FF869E69FB11FF69FAC7" authorityName="King" authorityYear="1864" class="Insecta" family="Staphylinidae" genus="Psepharobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869E69FB11FF69FAC7" italics="true" pageId="14" pageNumber="348">Psepharobius</emphasis>
</taxonomicName>
and the North American
<taxonomicName id="1DF64D59FF92FF869AB2FAF5FDAFFAC6" authorityName="Casey. Each" authorityYear="1897" box="[456,630,1303,1329]" class="Insecta" family="Staphylinidae" genus="Taphroscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869AB2FAF5FDAFFAC6" box="[456,630,1303,1329]" italics="true" pageId="14" pageNumber="348">Taphroscydmus</emphasis>
</taxonomicName>
remain unknown (none is present in Hungarian, Australian, and Russian museums that hold major parts of what remained from collections of Reitter, King and Motschulsky, respectively). The remaining genera can be divided into two groups based on the presence or absence of the lateral sutures of the submentum (the term “pre-tentorial gular sutures” will be used consistently in this and future studies). These groups may not be monophyletic, but such a preliminary division greatly facilitates identifications by excluding a large number of genera using a relatively easily observable character. The pre-tentorial gular sutures are known to occur in 23 genera, including
<taxonomicName id="1DF64D59FF92FF869ADFFA0DFDC9F9FF" authorityName="Jałoszyński" authorityYear="2020" box="[421,528,1519,1544]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869ADFFA0DFDC9F9FF" box="[421,528,1519,1544]" italics="true" pageId="14" pageNumber="348">Nogunius</emphasis>
</taxonomicName>
. Additionally, a re-assessment of cephalic structures in all specimens studied so far by scanning electron microscopy or as transparent mounts revealed that in
<taxonomicName id="1DF64D59FF92FF869F63F9F1FB3BF9DA" authorityName="Franz" authorityYear="1967" box="[1049,1250,1555,1581]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="E882EAC8FF92FF869F63F9F1FB3BF9DA" box="[1049,1250,1555,1581]" italics="true" pageId="14" pageNumber="348">Magellanoconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF92FF869F90F9F1FAFEF9DA" box="[1258,1319,1555,1581]" pageId="14" pageNumber="348" sensu="stricto">s. str.</taxonomicNameLabel>
structures interpreted previously as hypostomal ridges may in fact be the pre-tentorial gular sutures (discussion concerning the diagnosis and taxonomy of this genus will be included in a separate, phylogenetic paper; Jałoszyński in prep.). All genera with the pre-tentorial gular sutures can be divided into two groups:
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDF940FDDDF916" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
<emphasis id="E882EAC8FF92FF869BBDF940FF04F94B" bold="true" box="[199,221,1698,1724]" pageId="14" pageNumber="348">1.</emphasis>
The mesoventral intercoxal process absent:
<taxonomicName id="1DF64D59FF92FF86999CF941FC2DF94A" authority="Franz" authorityName="Franz" authorityYear="1971" box="[742,1012,1699,1725]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF86999CF941FC7FF94A" box="[742,934,1699,1725]" italics="true" pageId="14" pageNumber="348">Austrostenichnus</emphasis>
Franz
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869F78F941FB6FF94A" authority="Casey" authorityName="Casey" authorityYear="1897" box="[1026,1206,1699,1725]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869F78F941FBBFF94A" box="[1026,1126,1699,1725]" italics="true" pageId="14" pageNumber="348">Neladius</emphasis>
Casey
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869FBCF941FF03F916" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Palaeoscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869FBCF941FA44F94A" box="[1222,1437,1699,1725]" italics="true" pageId="14" pageNumber="348">Palaeoscydmaenus</emphasis>
Franz
</taxonomicName>
, and
<taxonomicName id="1DF64D59FF92FF869A6EF925FDDDF916" authority="Franz." authorityName="Franz." box="[276,516,1735,1761]" class="Insecta" family="Staphylinidae" genus="Stenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869A6EF925FE6FF916" box="[276,438,1735,1761]" italics="true" pageId="14" pageNumber="348">Stenichnaphes</emphasis>
Franz.
</taxonomicName>
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDF908FD96F862" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="348">
<emphasis id="E882EAC8FF92FF869BBDF908FF04F8F3" bold="true" box="[199,221,1770,1796]" pageId="14" pageNumber="348">2.</emphasis>
The mesoventral intercoxal process or carina present:
<taxonomicName id="1DF64D59FF92FF869829F909FBC2F8F2" authority="Franz" authorityName="Franz" authorityYear="1980" box="[851,1051,1771,1797]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869829F909FC17F8F2" box="[851,974,1771,1797]" italics="true" pageId="14" pageNumber="348">Alloraphes</emphasis>
Franz
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869F52F909FAFBF8F2" authority="Brendel" authorityName="Brendel" authorityYear="1889" box="[1064,1314,1771,1797]" class="Insecta" family="Staphylinidae" genus="Brachycepsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869F52F909FB64F8F2" box="[1064,1213,1771,1797]" italics="true" pageId="14" pageNumber="348">Brachycepsis</emphasis>
Brendel
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869E55F909FF05F8DE" authority="Casey" authorityName="Casey" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869E55F909FA45F8F2" box="[1327,1436,1771,1797]" italics="true" pageId="14" pageNumber="348">Catalinus</emphasis>
Casey
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869B91F8EDFE5AF8DE" authority="Casey" authorityName="Casey" authorityYear="1897" box="[235,387,1807,1833]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869B91F8EDFEEDF8DE" box="[235,308,1807,1833]" italics="true" pageId="14" pageNumber="348">Delius</emphasis>
Casey
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869AEBF8EDFD0CF8DE" authority="Croissandeau" authorityName="Croissandeau" box="[401,725,1807,1833]" class="Insecta" family="Staphylinidae" genus="Leptoderoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869AEBF8EDFDE9F8DE" box="[401,560,1807,1833]" italics="true" pageId="14" pageNumber="348">Leptoderoides</emphasis>
Croissandeau
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869998F8EDFBD8F8DE" authority="Jaloszynski" authorityName="Jaloszynski" box="[738,1025,1807,1833]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869998F8EDFCAAF8DE" box="[738,883,1807,1833]" italics="true" pageId="14" pageNumber="348">Madagaphes</emphasis>
Jałoszyński
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869F75F8EDFAFCF8DE" authority="Franz" authorityName="Franz" authorityYear="1967" box="[1039,1317,1807,1833]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869F75F8EDFB01F8DE" box="[1039,1240,1807,1833]" italics="true" pageId="14" pageNumber="348">Magellanoconnus</emphasis>
Franz
</taxonomicName>
,
<emphasis id="E882EAC8FF92FF869E49F8EDFF67F8BB" italics="true" pageId="14" pageNumber="348">Mexiconnus</emphasis>
Jałoszyński,
<taxonomicName id="1DF64D59FF92FF869A2CF8D1FD9DF8BA" authority="Thomson" authorityName="Thomson" box="[342,580,1843,1869]" class="Insecta" family="Staphylinidae" genus="Neuraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869A2CF8D1FE16F8BA" box="[342,463,1843,1869]" italics="true" pageId="14" pageNumber="348">Neuraphes</emphasis>
Thomson
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF86992AF8D1FC91F8BA" authority="Jaloszynski" authorityName="Jaloszynski" box="[592,840,1843,1869]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF86992AF8D1FD62F8BB" box="[592,699,1843,1868]" italics="true" pageId="14" pageNumber="348">Nogunius</emphasis>
Jałoszyński
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF86982EF8D1FBAEF8BA" authority="Jaloszynski" authorityName="Jaloszynski" authorityYear="2014" box="[852,1143,1843,1869]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF86982EF8D1FC32F8BA" box="[852,1003,1843,1869]" italics="true" pageId="14" pageNumber="348">Obesoconnus</emphasis>
Jałoszyński
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869FF9F8D1FABEF8BA" authority="Casey" authorityName="Casey" authorityYear="1897" box="[1155,1383,1843,1869]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869FF9F8D1FAC3F8BA" box="[1155,1306,1843,1869]" italics="true" pageId="14" pageNumber="348">Parascydmus</emphasis>
Casey
</taxonomicName>
,
<emphasis id="E882EAC8FF92FF869E0EF8D1FF20F886" italics="true" pageId="14" pageNumber="348">Rutaraphes</emphasis>
Jałoszyński,
<taxonomicName id="1DF64D59FF92FF869AE9F8B5FDB4F886" authority="King" authorityName="King" authorityYear="1864" box="[403,621,1879,1905]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869AE9F8B5FDF3F886" box="[403,554,1879,1905]" italics="true" pageId="14" pageNumber="348">Scydmaenilla</emphasis>
King
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869900F8B5FCADF886" authority="Reitter" authorityName="Reitter" authorityYear="1891" box="[634,884,1879,1905]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869900F8B5FCC4F886" box="[634,797,1879,1905]" italics="true" pageId="14" pageNumber="348">Scydmoraphes</emphasis>
Reitter
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF8698FAF8B5FBF2F886" authority="Franz" authorityName="Franz" authorityYear="1989" box="[896,1067,1879,1905]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF8698FAF8B5FC06F886" box="[896,991,1879,1905]" italics="true" pageId="14" pageNumber="348">Siamites</emphasis>
Franz
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869F4DF8B5FACEF886" authority="Franz" authorityName="Franz" authorityYear="1966" box="[1079,1303,1879,1905]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869F4DF8B5FB12F886" box="[1079,1227,1879,1905]" italics="true" pageId="14" pageNumber="348">Stenichnodes</emphasis>
Franz
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869E59F8B5FEDDF862" authority="Thomson" authorityName="Thomson" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869E59F8B5FA45F886" box="[1315,1436,1879,1905]" italics="true" pageId="14" pageNumber="348">Stenichnus</emphasis>
Thomson
</taxonomicName>
, and
<taxonomicName id="1DF64D59FF92FF869A44F899FD96F862" authority="Jaloszynski." box="[318,591,1915,1941]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869A44F899FE66F862" box="[318,447,1915,1941]" italics="true" pageId="14" pageNumber="348">Zeanichnus</emphasis>
Jałoszyński.
</taxonomicName>
</paragraph>
<paragraph id="DA4936DAFF92FF869BBDF87DFD39F84E" blockId="14.[151,1437,151,2013]" box="[199,736,1951,1977]" pageId="14" pageNumber="348">The group 2 can be divided into two subgroups:</paragraph>
<paragraph id="DA4936DAFF92FF879BBDF820FEC6FF22" blockId="14.[151,1437,151,2013]" lastBlockId="15.[151,1437,151,2014]" lastPageId="15" lastPageNumber="349" pageId="14" pageNumber="348">
<emphasis id="E882EAC8FF92FF869BBDF820FF29F82B" bold="true" box="[199,240,1986,2012]" pageId="14" pageNumber="348">2A.</emphasis>
Eyes clearly closer to occipital constriction than to mandibular bases:
<taxonomicName id="1DF64D59FF92FF869F4EF821FB76F82A" authorityName="Franz" authorityYear="1980" box="[1076,1199,1987,2013]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869F4EF821FB76F82A" box="[1076,1199,1987,2013]" italics="true" pageId="14" pageNumber="348">Alloraphes</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF92FF869FBBF821FA8FF82A" authorityName="Brendel" authorityYear="1889" box="[1217,1366,1987,2013]" class="Insecta" family="Staphylinidae" genus="Brachycepsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF92FF869FBBF821FA8FF82A" box="[1217,1366,1987,2013]" italics="true" pageId="14" pageNumber="348">Brachycepsis</emphasis>
</taxonomicName>
,
<emphasis id="E882EAC8FF92FF879E13F821FED5FF46" italics="true" lastPageId="15" lastPageNumber="349" pageId="14" pageNumber="348">Leptoderoides</emphasis>
,
<taxonomicName id="1DF64D59FF93FF879A65FF75FE69FF46" box="[287,432,151,177]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A65FF75FE69FF46" box="[287,432,151,177]" italics="true" pageId="15" pageNumber="349">Madagaphes</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF93FF879AB8FF75FDE2FF46" box="[450,571,151,177]" class="Insecta" family="Staphylinidae" genus="Neuraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AB8FF75FDE2FF46" box="[450,571,151,177]" italics="true" pageId="15" pageNumber="349">Neuraphes</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF93FF879934FF75FD3CFF46" authorityName="Casey" authorityYear="1897" box="[590,741,151,177]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879934FF75FD3CFF46" box="[590,741,151,177]" italics="true" pageId="15" pageNumber="349">Parascydmus</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF93FF879982FF75FCA0FF46" authorityName="Jaloszynski" authorityYear="2015" box="[760,889,151,177]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879982FF75FCA0FF46" box="[760,889,151,177]" italics="true" pageId="15" pageNumber="349">Rutaraphes</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF93FF8798F6FF75FBF6FF46" authorityName="Reitter" authorityYear="1891" box="[908,1071,151,177]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8798F6FF75FBF6FF46" box="[908,1071,151,177]" italics="true" pageId="15" pageNumber="349">Scydmoraphes</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF93FF879F3BFF75FB0CFF46" authorityName="Franz" authorityYear="1966" box="[1089,1237,151,177]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F3BFF75FB0CFF46" box="[1089,1237,151,177]" italics="true" pageId="15" pageNumber="349">Stenichnodes</emphasis>
</taxonomicName>
,
<taxonomicName id="1DF64D59FF93FF879F92FF75FAB8FF46" box="[1256,1377,151,177]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F92FF75FAB8FF46" box="[1256,1377,151,177]" italics="true" pageId="15" pageNumber="349">Stenichnus</emphasis>
</taxonomicName>
, and
<taxonomicName id="1DF64D59FF93FF879BEDFF59FEC1FF22" box="[151,280,187,213]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879BEDFF59FEC1FF22" box="[151,280,187,213]" italics="true" pageId="15" pageNumber="349">Zeanichnus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DA4936DAFF93FF879BBDFF3CFEB2FDED" blockId="15.[151,1437,151,2014]" pageId="15" pageNumber="349">
<emphasis id="E882EAC8FF93FF879BBDFF3CFF37FF0F" bold="true" box="[199,238,222,248]" pageId="15" pageNumber="349">2B.</emphasis>
Eyes either closer to mandibular bases than to the occipital constriction, or at middle, or so large that touching or nearly touching both mandibular bases and occipital constriction; or the placement of eye is ambiguous or variable within genus (e.g., because of the occurrence of macrophthalmous species with eyes close to middle and microphthalmous with eyes close to mandibular bases):
<taxonomicName id="1DF64D59FF93FF87986DFEA9FC5DFE92" authorityName="Casey" authorityYear="1897" box="[791,900,331,357]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF87986DFEA9FC5DFE92" box="[791,900,331,357]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
(eyes near middle),
<taxonomicName id="1DF64D59FF93FF879F0AFEA9FB60FE92" authorityName="Casey" authorityYear="1897" box="[1136,1209,331,357]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F0AFEA9FB60FE92" box="[1136,1209,331,357]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
(eyes near middle),
<taxonomicName id="1DF64D59FF93FF879BEDFE8DFEB9FE7E" authorityName="Franz" authorityYear="1967" box="[151,352,367,393]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="E882EAC8FF93FF879BEDFE8DFEB9FE7E" box="[151,352,367,393]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF93FF879A12FE8DFE7AFE7E" box="[360,419,367,393]" pageId="15" pageNumber="349" sensu="stricto">s. str.</taxonomicNameLabel>
(eyes clearly closer to mandibular bases),
<taxonomicName id="1DF64D59FF93FF8798FCFE8DFBD4FE7F" authorityName="Jaloszynski" authorityYear="2013" box="[902,1037,367,392]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8798FCFE8DFBD4FE7F" box="[902,1037,367,392]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
(eyes slightly closer to mandibular bases),
<taxonomicName id="1DF64D59FF93FF879B96FE71FE8EFE5B" box="[236,343,403,428]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879B96FE71FE8EFE5B" box="[236,343,403,428]" italics="true" pageId="15" pageNumber="349">Nogunius</emphasis>
</taxonomicName>
(eyes clearly closer to mandibular bases),
<taxonomicName id="1DF64D59FF93FF879847FE71FC0DFE5A" authorityName="Jaloszynski" authorityYear="2014" box="[829,980,403,429]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879847FE71FC0DFE5A" box="[829,980,403,429]" italics="true" pageId="15" pageNumber="349">Obesoconnus</emphasis>
</taxonomicName>
(huge eyes occupying entire lateral surface of head),
<taxonomicName id="1DF64D59FF93FF879A47FE55FE0DFE26" authorityName="King" authorityYear="1864" box="[317,468,439,465]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A47FE55FE0DFE26" box="[317,468,439,465]" italics="true" pageId="15" pageNumber="349">Scydmaenilla</emphasis>
</taxonomicName>
(eyes closer to occipital constriction or near middle),
<taxonomicName id="1DF64D59FF93FF879F38FE55FB78FE26" authorityName="Franz" authorityYear="1989" box="[1090,1185,439,465]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F38FE55FB78FE26" box="[1090,1185,439,465]" italics="true" pageId="15" pageNumber="349">Siamites</emphasis>
</taxonomicName>
(eyes near middle, or slightly closer to mandibular bases, in microphthalmous specimens adjacent to antennal fossae and clearly closer to mandibular bases).
</paragraph>
<paragraph id="DA4936DAFF93FF879BBDFDC1FC55FD5E" blockId="15.[151,1437,151,2014]" pageId="15" pageNumber="349">
Within group 2B,
<taxonomicName id="1DF64D59FF93FF879AF5FDC1FE23FDCB" box="[399,506,547,572]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AF5FDC1FE23FDCB" box="[399,506,547,572]" italics="true" pageId="15" pageNumber="349">Nogunius</emphasis>
</taxonomicName>
can be distinguished from all remaining genera by (1) the sharply delimited, dimerous antennal club; (2) antennal fossae narrowly separated at middle by distance narrower than width of scape; (3) anterior margin of the mentum distinctly concave; and (4) the pronotum broadest shortly in front of base. Additionally,
<taxonomicName id="1DF64D59FF93FF879BB7FD6DFEE1FD5F" box="[205,312,655,680]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879BB7FD6DFEE1FD5F" box="[205,312,655,680]" italics="true" pageId="15" pageNumber="349">Nogunius</emphasis>
</taxonomicName>
differs in several features from each of these genera:
</paragraph>
<paragraph id="DA4936DAFF93FF879BBDFD51FD0EFB55" blockId="15.[151,1437,151,2014]" pageId="15" pageNumber="349">
<emphasis id="E882EAC8FF93FF879BBDFD51FE53FD3A" bold="true" box="[199,394,690,717]" pageId="15" pageNumber="349">
- from
<taxonomicName id="1DF64D59FF93FF879A60FD50FE53FD3A" authorityName="Casey" authorityYear="1897" box="[282,394,690,717]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A60FD50FE53FD3A" bold="true" box="[282,394,690,717]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
</emphasis>
in thick bristles present on head and pronotum (lacking in
<taxonomicName id="1DF64D59FF93FF879F5EFD51FB48FD3A" authorityName="Casey" authorityYear="1897" box="[1060,1169,691,717]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F5EFD51FB48FD3A" box="[1060,1169,691,717]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
); tempora strongly narrowing posterad and forming an even arc with posteriorly convex posterior margin of vertex (in
<taxonomicName id="1DF64D59FF93FF879FBCFD35FAEAFD06" authorityName="Casey" authorityYear="1897" box="[1222,1331,727,753]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879FBCFD35FAEAFD06" box="[1222,1331,727,753]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
, tempora weakly narrowing posterad and delimited by distinct obtuse-angled corners from anteriorly convex posterior margin of vertex); posterior tentorial pits exposed and situated far in front of broad, diffuse transverse impression demarcating ventrally neck region (in
<taxonomicName id="1DF64D59FF93FF879A8AFCA1FD84FCAA" authorityName="Casey" authorityYear="1897" box="[496,605,835,861]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A8AFCA1FD84FCAA" box="[496,605,835,861]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
posterior tentorial pits hidden in deep, narrow, sharply marked transverse groove demarcating ventrally neck region); hypostomal ridges incomplete, short, developed only shortly behind cardines (in
<taxonomicName id="1DF64D59FF93FF879A5AFC69FE54FC52" authorityName="Casey" authorityYear="1897" box="[288,397,907,933]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A5AFC69FE54FC52" box="[288,397,907,933]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
hypostomal ridges complete, long, posteromesally confluent with transverse groove in which posterior tentorial pits are situated); pronotum with four antebasal pits (pits lacking in
<taxonomicName id="1DF64D59FF93FF879F2FFC4DFB1BFC3E" authorityName="Casey" authorityYear="1897" box="[1109,1218,943,969]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F2FFC4DFB1BFC3E" box="[1109,1218,943,969]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
); hypomeral ridges lacking (in
<taxonomicName id="1DF64D59FF93FF879A64FC31FE52FC1A" authorityName="Casey" authorityYear="1897" box="[286,395,979,1005]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A64FC31FE52FC1A" box="[286,395,979,1005]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
incomplete but present); posteromedian margin of prothoracic coxosternum with shallow emargination (in
<taxonomicName id="1DF64D59FF93FF879A2DFC15FE1DFBE6" authorityName="Casey" authorityYear="1897" box="[343,452,1015,1041]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A2DFC15FE1DFBE6" box="[343,452,1015,1041]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
with subtriangular projection); basal elytral fovea situated near middle of elytral articulating lobe, distant from sutural margin (in
<taxonomicName id="1DF64D59FF93FF87990FFBF9FD3BFBC2" authorityName="Casey" authorityYear="1897" box="[629,738,1051,1077]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF87990FFBF9FD3BFBC2" box="[629,738,1051,1077]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
close to sutural margin); mesoventral intercoxal process broad and behind mesocoxae indistinguishably fused with metaventrite (in
<taxonomicName id="1DF64D59FF93FF8798DDFBDDFBCDFBAE" authorityName="Casey" authorityYear="1897" box="[935,1044,1087,1113]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8798DDFBDDFBCDFBAE" box="[935,1044,1087,1113]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
process narrow and reaching only to middle of mesocoxae, where it forms a distinct posterior tip separated from metaventrite); and in symmetrical endophallic structures (in
<taxonomicName id="1DF64D59FF93FF879AC1FB65FDF1FB56" authorityName="Casey" authorityYear="1897" box="[443,552,1159,1185]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AC1FB65FDF1FB56" box="[443,552,1159,1185]" italics="true" pageId="15" pageNumber="349">Catalinus</emphasis>
</taxonomicName>
asymmetrical);
</paragraph>
<paragraph id="DA4936DAFF93FF879BBDFB49FCEDF9FD" blockId="15.[151,1437,151,2014]" pageId="15" pageNumber="349">
<emphasis id="E882EAC8FF93FF879BBDFB49FEBEFB32" bold="true" box="[199,359,1194,1221]" pageId="15" pageNumber="349">
- from
<taxonomicName id="1DF64D59FF93FF879A66FB48FEBEFB32" authorityName="Casey" authorityYear="1897" box="[284,359,1194,1221]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A66FB48FEBEFB32" bold="true" box="[284,359,1194,1221]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
</emphasis>
in tempora strongly narrowing posterad and forming an even arc with posteriorly convex posterior margin of vertex (in
<taxonomicName id="1DF64D59FF93FF879AA9FB2DFDC5FB1E" authorityName="Casey" authorityYear="1897" box="[467,540,1231,1257]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AA9FB2DFDC5FB1E" box="[467,540,1231,1257]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
, tempora weakly narrowing posterad and delimited by distinct obtuse-angled corners from anteriorly convex posterior margin of vertex); hypostomal ridges incomplete, short, developed only shortly behind cardines (in
<taxonomicName id="1DF64D59FF93FF879AA8FAF5FDC2FAC6" authorityName="Casey" authorityYear="1897" box="[466,539,1303,1329]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AA8FAF5FDC2FAC6" box="[466,539,1303,1329]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
hypostomal ridges complete, long, posteromesally reaching posterior tentorial pits); pronotum with inner antebasal pits not connected by transverse groove (groove present in
<taxonomicName id="1DF64D59FF93FF879FBEFAD9FAD4FAA2" authorityName="Casey" authorityYear="1897" box="[1220,1293,1339,1365]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879FBEFAD9FAD4FAA2" box="[1220,1293,1339,1365]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
); hypomeral ridges lacking (in
<taxonomicName id="1DF64D59FF93FF879A1EFABDFE74FA8E" authorityName="Casey" authorityYear="1897" box="[356,429,1375,1401]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A1EFABDFE74FA8E" box="[356,429,1375,1401]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
present); basal elytral fovea situated near middle of elytral articulating lobe, distant from sutural margin (in
<taxonomicName id="1DF64D59FF93FF879A13FA61FE6BFA6A" authorityName="Casey" authorityYear="1897" box="[361,434,1411,1437]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A13FA61FE6BFA6A" box="[361,434,1411,1437]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
basal elytral fovea close to sutural margin); mesoventral intercoxal process with straight lateral margins gradually diverging from anterior to posterior region (in
<taxonomicName id="1DF64D59FF93FF8798AEFA45FBC4FA36" authorityName="Casey" authorityYear="1897" box="[980,1053,1447,1473]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8798AEFA45FBC4FA36" box="[980,1053,1447,1473]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
process distinctly constricted between mesocoxae); and metacoxae laterally reaching sides of metaventrite (in
<taxonomicName id="1DF64D59FF93FF879F76FA29FB8CFA12" authorityName="Casey" authorityYear="1897" box="[1036,1109,1483,1509]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F76FA29FB8CFA12" box="[1036,1109,1483,1509]" italics="true" pageId="15" pageNumber="349">Delius</emphasis>
</taxonomicName>
lateral corners of metacoxae are broadly separated from lateral margins of metaventrite);
</paragraph>
<paragraph id="DA4936DAFF93FF879BBDF9F1FE96F8F2" blockId="15.[151,1437,151,2014]" pageId="15" pageNumber="349">
<emphasis id="E882EAC8FF93FF879BBDF9F1FDE4F9DA" bold="true" box="[199,573,1554,1581]" pageId="15" pageNumber="349">
- from
<taxonomicName id="1DF64D59FF93FF879A5AF9F0FE29F9DA" authorityName="Franz" authorityYear="1967" box="[288,496,1554,1581]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="E882EAC8FF93FF879A5AF9F0FE29F9DA" bold="true" box="[288,496,1554,1581]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF93FF879A81F9F1FDE4F9DA" box="[507,573,1555,1581]" pageId="15" pageNumber="349" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
in pronotum lacking sublateral carinae (present in
<taxonomicName id="1DF64D59FF93FF879FE9F9F1FA85F9DA" authorityName="Franz" authorityYear="1967" box="[1171,1372,1555,1581]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879FE9F9F1FA85F9DA" box="[1171,1372,1555,1581]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
); hypomeral ridges lacking (in
<taxonomicName id="1DF64D59FF93FF879AB6F9D5FD4CF9A6" authorityName="Franz" authorityYear="1967" box="[460,661,1591,1617]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AB6F9D5FD4CF9A6" box="[460,661,1591,1617]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
present); procoxal cavities closed (open in
<taxonomicName id="1DF64D59FF93FF879FF3F9D5FA8BF9A6" authorityName="Franz" authorityYear="1967" box="[1161,1362,1591,1617]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879FF3F9D5FA8BF9A6" box="[1161,1362,1591,1617]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
); posteromedian margin of prothoracic coxosternum with shallow emargination (in
<taxonomicName id="1DF64D59FF93FF879F7BF9B9FB13F982" authorityName="Franz" authorityYear="1967" box="[1025,1226,1627,1653]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879F7BF9B9FB13F982" box="[1025,1226,1627,1653]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
with subtriangular projection); one basal fovea on each elytron (two in
<taxonomicName id="1DF64D59FF93FF8799AEF99DFC44F96E" authorityName="Franz" authorityYear="1967" box="[724,925,1663,1689]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8799AEF99DFC44F96E" box="[724,925,1663,1689]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
); mesoventral intercoxal process long, behind mesocoxae fused with metaventrite (in
<taxonomicName id="1DF64D59FF93FF87992EF941FCC4F94A" authorityName="Franz" authorityYear="1967" box="[596,797,1699,1725]" class="Insecta" family="Staphylinidae" genus="Magellanoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF87992EF941FCC4F94A" box="[596,797,1699,1725]" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
</taxonomicName>
process reaching only to middle of mesocoxae, where it forms a distinct posterior tip separated from metaventrite); and anterior metaventral process lacking (in
<emphasis id="E882EAC8FF93FF879E60F925FF3FF8F3" italics="true" pageId="15" pageNumber="349">Magellanoconnus</emphasis>
present);
</paragraph>
<paragraph id="DA4936DAFF93FF879BBDF8EDFC09F829" blockId="15.[151,1437,151,2014]" pageId="15" pageNumber="349">
<emphasis id="E882EAC8FF93FF879BBDF8EDFE77F8DE" bold="true" box="[199,430,1806,1833]" pageId="15" pageNumber="349">
- from
<taxonomicName id="1DF64D59FF93FF879A65F8EDFE77F8DE" authorityName="Jaloszynski" authorityYear="2013" box="[287,430,1807,1833]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A65F8EDFE77F8DE" bold="true" box="[287,430,1807,1833]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
</emphasis>
in head with thick bristles (lacking in
<taxonomicName id="1DF64D59FF93FF87980EF8EDFC22F8DF" authorityName="Jaloszynski" authorityYear="2013" box="[884,1019,1807,1832]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF87980EF8EDFC22F8DF" box="[884,1019,1807,1832]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
); pronotum lacking transverse antebasal groove (groove present in
<taxonomicName id="1DF64D59FF93FF879A87F8D1FD5DF8BB" authorityName="Jaloszynski" authorityYear="2013" box="[509,644,1843,1868]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879A87F8D1FD5DF8BB" box="[509,644,1843,1868]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
); hypomeral ridges lacking (in
<taxonomicName id="1DF64D59FF93FF87989EF8D1FBB2F8BB" authorityName="Jaloszynski" authorityYear="2013" box="[996,1131,1843,1868]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF87989EF8D1FBB2F8BB" box="[996,1131,1843,1868]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
present); each elytron with one distinct basal fovea (in
<taxonomicName id="1DF64D59FF93FF879AACF8B5FD84F887" authorityName="Jaloszynski" authorityYear="2013" box="[470,605,1879,1904]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF879AACF8B5FD84F887" box="[470,605,1879,1904]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
two barely discernible vestiges of foveae); protarsi in males unmodified (strongly broadened and with long and dense ventral setal cushions in
<taxonomicName id="1DF64D59FF93FF8798CCF899FBE4F863" authorityName="Jaloszynski" authorityYear="2013" box="[950,1085,1915,1940]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8798CCF899FBE4F863" box="[950,1085,1915,1940]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
); and aedeagus with symmetrical endophallus and slender, free parameres bearing apical setae (in
<taxonomicName id="1DF64D59FF93FF8798CAF87DFBEEF84F" authorityName="Jaloszynski" authorityYear="2013" box="[944,1079,1951,1976]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF93FF8798CAF87DFBEEF84F" box="[944,1079,1951,1976]" italics="true" pageId="15" pageNumber="349">Mexiconnus</emphasis>
</taxonomicName>
endophallus asymmetrical and parameres in basal halves broader than median lobe, with asetose apices);
</paragraph>
<paragraph id="DA4936DAFF8CFF989BBDFF74FACFFDEE" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="350">
<emphasis id="E882EAC8FF8CFF989BBDFF74FE6AFF46" bold="true" box="[199,435,150,177]" pageId="16" pageNumber="350">
- from
<taxonomicName id="1DF64D59FF8CFF989A6DFF74FE6AFF46" authorityName="Jaloszynski" authorityYear="2014" box="[279,435,150,177]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989A6DFF74FE6AFF46" bold="true" box="[279,435,150,177]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
</emphasis>
in distinct tempora and distinct occipital constriction (both lacking in
<taxonomicName id="1DF64D59FF8CFF989FC0FF74FA88FF47" authorityName="Jaloszynski" authorityYear="2014" box="[1210,1361,150,176]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989FC0FF74FA88FF47" box="[1210,1361,150,176]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
); head and pronotum with thick bristles (absent in
<taxonomicName id="1DF64D59FF8CFF9899F8FF58FCC0FF23" authorityName="Jaloszynski" authorityYear="2014" box="[642,793,186,212]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF9899F8FF58FCC0FF23" box="[642,793,186,212]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
); posterior tentorial pits exposed and situated far in front of broad, diffuse transverse impression demarcating ventrally neck region (in
<taxonomicName id="1DF64D59FF8CFF989F73FF3CFB79FF0F" authorityName="Jaloszynski" authorityYear="2014" box="[1033,1184,222,248]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989F73FF3CFB79FF0F" box="[1033,1184,222,248]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
posterior tentorial pits hidden in a deep, narrow, sharply marked transverse groove demarcating ventrally neck region); pronotum lacking transverse antebasal groove (groove present in
<taxonomicName id="1DF64D59FF8CFF9899D1FEC4FC9BFEB7" authorityName="Jaloszynski" authorityYear="2014" box="[683,834,294,320]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF9899D1FEC4FC9BFEB7" box="[683,834,294,320]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
); maxillary palpomere III strongly elongate and with distal half nearly rod-like (stout, evenly subtriangular in
<taxonomicName id="1DF64D59FF8CFF98987FFEA8FC45FE93" authorityName="Jaloszynski" authorityYear="2014" box="[773,924,330,356]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF98987FFEA8FC45FE93" box="[773,924,330,356]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
); procoxal cavities closed (open in
<emphasis id="E882EAC8FF8CFF989E59FEA8FF67FE7F" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
); hypomeral ridges absent (present in
<taxonomicName id="1DF64D59FF8CFF98990AFE8CFCDEFE7F" authorityName="Jaloszynski" authorityYear="2014" box="[624,775,366,392]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF98990AFE8CFCDEFE7F" box="[624,775,366,392]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
); posteromedian emargination of prothoracic coxosternum shallow, barely marked (in
<taxonomicName id="1DF64D59FF8CFF989AB4FE70FDBCFE5B" authorityName="Jaloszynski" authorityYear="2014" box="[462,613,402,428]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989AB4FE70FDBCFE5B" box="[462,613,402,428]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
conspicuously deep, reaching nearly half length of prosternum); mesoscutellar shield tiny and barely discernible between elytral bases (in
<taxonomicName id="1DF64D59FF8CFF9898FDFE54FBC7FE27" authorityName="Jaloszynski" authorityYear="2014" box="[903,1054,438,464]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF9898FDFE54FBC7FE27" box="[903,1054,438,464]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
conspicuously large and well-visible); and aedeagus lacking basal pumping apparatus (in
<taxonomicName id="1DF64D59FF8CFF989867FE38FC6DFE03" authorityName="Jaloszynski" authorityYear="2014" box="[797,948,474,500]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989867FE38FC6DFE03" box="[797,948,474,500]" italics="true" pageId="16" pageNumber="350">Obesoconnus</emphasis>
</taxonomicName>
present, composed of basal diaphragm with sclerotized median area to which internal longitudinal apodeme with longitudinal muscles is attached);
</paragraph>
<paragraph id="DA4936DAFF8CFF989BBDFDC0FDCFFCAA" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="350">
<emphasis id="E882EAC8FF8CFF989BBDFDC0FE6EFDCA" bold="true" box="[199,439,546,573]" pageId="16" pageNumber="350">
- from
<taxonomicName id="1DF64D59FF8CFF989A61FDC0FE6EFDCA" authorityName="King" authorityYear="1864" box="[283,439,546,573]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989A61FDC0FE6EFDCA" bold="true" box="[283,439,546,573]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
</emphasis>
in head with thick bristles (absent in
<taxonomicName id="1DF64D59FF8CFF98981AFDC0FC2EFDCB" authorityName="King" authorityYear="1864" box="[864,1015,546,572]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF98981AFDC0FC2EFDCB" box="[864,1015,546,572]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
); posterior tentorial pits exposed and situated far in front of broad, diffuse transverse impression demarcating ventrally neck region (in
<taxonomicName id="1DF64D59FF8CFF989E7FFDA4FA45FD97" authorityName="King" authorityYear="1864" box="[1285,1436,582,608]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989E7FFDA4FA45FD97" box="[1285,1436,582,608]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
posterior tentorial pits are hidden in a deep, narrow, sharply marked transverse groove demarcating ventrally neck region); pronotum with two pairs of antebasal pits, lacking transverse groove (in
<taxonomicName id="1DF64D59FF8CFF989F5FFD6CFB65FD5F" authorityName="King" authorityYear="1864" box="[1061,1212,654,680]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989F5FFD6CFB65FD5F" box="[1061,1212,654,680]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
one pair of pits and transverse groove are present); postcoxal coxosternal and hypomeral processes firmly fused (in
<taxonomicName id="1DF64D59FF8CFF989FB4FD50FABCFD3B" authorityName="King" authorityYear="1864" box="[1230,1381,690,716]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989FB4FD50FABCFD3B" box="[1230,1381,690,716]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
posterior hypomeral process overlaps coxosternal process, but margins of each are well discernible); mesoventral intercoxal process long, behind mesocoxae fused with metaventrite (in
<taxonomicName id="1DF64D59FF8CFF9898E1FD18FBEBFCE3" authorityName="King" authorityYear="1864" box="[923,1074,762,788]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF9898E1FD18FBEBFCE3" box="[923,1074,762,788]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
process reaching only to middle of mesocoxae, where it forms a distinct posterior tip separated from metaventrite); and anterior metaventral process lacking (in
<taxonomicName id="1DF64D59FF8CFF989A6DFCA0FE77FCAB" authorityName="King" authorityYear="1864" box="[279,430,834,860]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989A6DFCA0FE77FCAB" box="[279,430,834,860]" italics="true" pageId="16" pageNumber="350">Scydmaenilla</emphasis>
</taxonomicName>
present);
</paragraph>
<paragraph id="DA4936DAFF8CFF989BBDFC84FBD7FBAE" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="350">
<emphasis id="E882EAC8FF8CFF989BBDFC84FEA1FC77" bold="true" box="[199,376,870,896]" pageId="16" pageNumber="350">
- from
<taxonomicName id="1DF64D59FF8CFF989A6CFC84FEA1FC77" authorityName="Franz" authorityYear="1989" box="[278,376,870,896]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989A6CFC84FEA1FC77" bold="true" box="[278,376,870,896]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
</emphasis>
in lacking paired, dense tufts of bristles on vertex, prosternum, meso- and metaventrite and posterior margins of abdominal sternites (in
<taxonomicName id="1DF64D59FF8CFF98991CFC69FD1CFC52" authorityName="Franz" authorityYear="1989" box="[614,709,907,933]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF98991CFC69FD1CFC52" box="[614,709,907,933]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
conspicuous tufts or rosettes of bristles present on each of these regions); pre-tentorial gular sutures separated at middle (in
<taxonomicName id="1DF64D59FF8CFF98983FFC4DFC7DFC3E" authorityName="Franz" authorityYear="1989" box="[837,932,943,969]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF98983FFC4DFC7DFC3E" box="[837,932,943,969]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
connected between posterior tentorial pits); incomplete, short hypostomal ridges (in
<taxonomicName id="1DF64D59FF8CFF989911FC31FD13FC1A" authorityName="Franz" authorityYear="1989" box="[619,714,979,1005]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989911FC31FD13FC1A" box="[619,714,979,1005]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
complete, reaching level of posterior tentorial pits); pronotum lacking transverse antebasal groove (in
<taxonomicName id="1DF64D59FF8CFF989921FC15FD63FBE6" authorityName="Franz" authorityYear="1989" box="[603,698,1015,1041]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989921FC15FD63FBE6" box="[603,698,1015,1041]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
present); procoxal cavities closed (open in
<taxonomicName id="1DF64D59FF8CFF989FD6FC15FAD2FBE6" authorityName="Franz" authorityYear="1989" box="[1196,1291,1015,1041]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989FD6FC15FAD2FBE6" box="[1196,1291,1015,1041]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
); hypomeral ridges lacking (present in
<taxonomicName id="1DF64D59FF8CFF989AC2FBF9FDCEFBC2" authorityName="Franz" authorityYear="1989" box="[440,535,1051,1077]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989AC2FBF9FDCEFBC2" box="[440,535,1051,1077]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
); and mesoventral intercoxal process behind mesocoxae fused with metaventrite (in
<taxonomicName id="1DF64D59FF8CFF989BC7FBDDFEC5FBAE" authorityName="Franz" authorityYear="1989" box="[189,284,1087,1113]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8CFF989BC7FBDDFEC5FBAE" box="[189,284,1087,1113]" italics="true" pageId="16" pageNumber="350">Siamites</emphasis>
</taxonomicName>
process forms a distinct posterior tip separated from metaventrite).
</paragraph>
<paragraph id="DA4936DAFF8CFF989BBDFB80FBCFFB57" blockId="16.[151,1437,150,1185]" pageId="16" pageNumber="350">
As two genera are here reported to occur in
<collectingCountry id="A2E1764AFF8CFF9899CAFB81FD29FB8A" box="[688,752,1123,1149]" name="Japan" pageId="16" pageNumber="350">Japan</collectingCountry>
for the first time, the identification key to Japanese genera of
<taxonomicName id="1DF64D59FF8CFF989BEDFB64FECEFB57" authorityName="Ganglbauer" authorityYear="1899" box="[151,279,1158,1184]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="tribe" tribe="Stenichnini">Stenichnini</taxonomicName>
(
<taxonomicName id="1DF64D59FF8CFF989A5FFB64FE14FB57" authorityName="L.W.Schaufuss" authorityYear="1889" box="[293,461,1158,1184]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="350" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
then) given by
<bibRefCitation id="BE674B2BFF8CFF989907FB64FC89FB56" author="Jaloszynski, P." box="[637,848,1158,1185]" pageId="16" pageNumber="350" pagination="330 - 339" refId="ref17019" refString="Jaloszynski, P. (2015) A new genus of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae) from Japan. Entomological Science, 18 (3), 330 - 339. https: // doi. org / 10.1111 / ens. 12136" type="journal article" year="2015">Jałoszyński (2015)</bibRefCitation>
must be updated.
</paragraph>
</subSubSection>
</treatment>
</document>