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<document id="D9909159FEA3B2417BF51D8908C3E766" ID-CLB-Dataset="8782" ID-DOI="10.11646/zootaxa.4822.3.2" ID-GBIF-Dataset="3c15254c-086c-448f-bfe2-e3cd208dcc87" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401617" ID-ZooBank="86DC1241-FE11-49CC-9674-ACAB1DF92F62" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609337343636" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="525F87CCFF8AFF9F9BEDF8CAFC16FAFA" docLanguage="en" docName="zootaxa.4822.3.2.pdf" docOrigin="Zootaxa 4822 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nogunius kerri Jałoszyński 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="357" masterDocId="AE66FFB4FF9CFF889B7AFFE2FFD9FFF7" masterDocTitle="Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="356" updateTime="1698849732245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B601E650203BE6B06E7E460FF8352C62">Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae)</mods:title>
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<mods:namePart id="D3109F7ABA27394A4DE2DB1987065AB9">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="1CCC811FFC674FD55BB1DA9AA8167202">2020</mods:date>
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<treatment id="525F87CCFF8AFF9F9BEDF8CAFC16FAFA" ID-DOI="http://doi.org/10.5281/zenodo.4401603" ID-GBIF-Taxon="177009896" ID-Zenodo-Dep="4401603" LSID="urn:lsid:plazi:treatment:525F87CCFF8AFF9F9BEDF8CAFC16FAFA" httpUri="http://treatment.plazi.org/id/525F87CCFF8AFF9F9BEDF8CAFC16FAFA" lastPageId="23" lastPageNumber="357" pageId="22" pageNumber="356">
<subSubSection id="92EC6551FF8AFF9E9BEDF8CAFE43F8B5" box="[151,410,1831,1858]" pageId="22" pageNumber="356" type="nomenclature">
<paragraph id="DA4936DAFF8AFF9E9BEDF8CAFE43F8B5" blockId="22.[151,410,1831,1895]" box="[151,410,1831,1858]" pageId="22" pageNumber="356">
<heading id="810181B6FF8AFF9E9BEDF8CAFE43F8B5" bold="true" box="[151,410,1831,1858]" fontSize="11" level="1" pageId="22" pageNumber="356" reason="1">
<emphasis id="E882EAC8FF8AFF9E9BEDF8CAFE43F8B5" bold="true" box="[151,410,1831,1858]" pageId="22" pageNumber="356">
<taxonomicName id="1DF64D59FF8AFF9E9BEDF8CAFE88F8B5" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,337,1831,1858]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="356" phylum="Arthropoda" rank="species" species="kerri" status="sp. nov.">
<emphasis id="E882EAC8FF8AFF9E9BEDF8CAFE88F8B5" bold="true" box="[151,337,1831,1858]" italics="true" pageId="22" pageNumber="356">Nogunius kerri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3B157B3FF8AFF9E9A22F8CAFE43F8B5" box="[344,410,1832,1858]" pageId="22" pageNumber="356" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8AFF9E9BEDF8AEFEA2F891" box="[151,379,1868,1895]" pageId="22" pageNumber="356" type="description">
<paragraph id="DA4936DAFF8AFF9E9BEDF8AEFEA2F891" blockId="22.[151,410,1831,1895]" box="[151,379,1868,1895]" pageId="22" pageNumber="356">
(
<figureCitation id="42CD2A5FFF8AFF9E9BE5F8AEFEF8F890" box="[159,289,1868,1895]" captionStart="FIGURES 4552" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[151,1436,862,1918]" captionTargetId="figure-321@13.[151,1436,849,1918]" captionTargetPageId="13" captionText="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." figureDoi="http://doi.org/10.5281/zenodo.4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="22" pageNumber="356">Figs 4950</figureCitation>
,
<figureCitation id="42CD2A5FFF8AFF9E9A56F8AEFEAAF891" box="[300,371,1868,1894]" captionStart="FIGURES 7784" captionStartId="21.[151,264,1915,1939]" captionTargetBox="[164,1432,189,1882]" captionTargetId="figure-19@21.[151,1436,181,1892]" captionTargetPageId="21" captionText="FIGURES 7784. Aedeagus in ventral (77, 79, 81, 83) and lateral (78, 80, 82, 84) views. Nogunius kerri sp. n. (7780); and N. fukuuzanus sp. n. (8184)." figureDoi="http://doi.org/10.5281/zenodo.4401643" httpUri="https://zenodo.org/record/4401643/files/figure.png" pageId="22" pageNumber="356">7780</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8AFF9E9BEDF876FD70F801" pageId="22" pageNumber="356" type="materials_examined">
<paragraph id="DA4936DAFF8AFF9E9BEDF876FD70F801" blockId="22.[151,1437,1940,2040]" pageId="22" pageNumber="356">
<emphasis id="E882EAC8FF8AFF9E9BEDF876FE91F859" bold="true" box="[151,328,1940,1966]" pageId="22" pageNumber="356">Type material.</emphasis>
<materialsCitation id="6A9E3C87FF8AFF9E9A2BF876FD70F801" ID-GBIF-Occurrence="3019650329" collectingDate="1999-06-03" collectionCode="EUMJ" collectorName="H. Mizushima" country="Japan" location="Okinawa Pref." pageId="22" pageNumber="356" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="E882EAC8FF8AFF9E9A2BF876FD36F859" bold="true" box="[337,751,1940,1966]" pageId="22" pageNumber="356">
<typeStatus id="054D8878FF8AFF9E9A2BF876FE1BF859" box="[337,450,1940,1966]" pageId="22" pageNumber="356" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="A2E1764AFF8AFF9E9AABF876FDF2F859" box="[465,555,1940,1966]" name="Japan" pageId="22" pageNumber="356">JAPAN</collectingCountry>
(
<collectingRegion id="1832F838FF8AFF9E9946F876FD36F859" box="[572,751,1940,1966]" country="Japan" name="Okinawa" pageId="22" pageNumber="356">Okinawa Pref.</collectingRegion>
</emphasis>
):
<specimenCount id="CCF0FD53FF8AFF9E9871F877FCFBF858" box="[779,802,1941,1967]" pageId="22" pageNumber="356" type="male"></specimenCount>
, two labels: “Mt. OMOTO-DAKE / Ishigaki Is. / Okinawa Pref. /
<date id="AE48101AFF8AFF9E9A55F85AFE7CF825" box="[303,421,1976,2002]" pageId="22" pageNumber="356" value="1999-06-03">
<collectingDate id="BE0CE9F2FF8AFF9E9A55F85AFE7CF825" box="[303,421,1976,2002]" pageId="22" pageNumber="356" value="1999-06-03">3-VI-1999</collectingDate>
</date>
/
<collectorName id="7703530CFF8AFF9E9ABBF85BFDBCF825" box="[449,613,1976,2003]" pageId="22" pageNumber="356">H. Mizushima</collectorName>
leg” [white, handwritten], “
<taxonomicName id="1DF64D59FF8AFF9E98D7F85AFBE2F825" box="[941,1083,1976,2002]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="E882EAC8FF8AFF9E98D7F85AFBE2F825" box="[941,1083,1976,2002]" italics="true" pageId="22" pageNumber="356">NOGUNIUS</emphasis>
</taxonomicName>
/
<taxonomicName id="1DF64D59FF8AFF9E9F2DF85AFB54F825" authorityName="Jałoszyński" authorityYear="2020" box="[1111,1165,1976,2002]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="356" phylum="Arthropoda" rank="species" species="kerri">
<emphasis id="E882EAC8FF8AFF9E9F2DF85AFB54F825" box="[1111,1165,1976,2002]" italics="true" pageId="22" pageNumber="356">kerri</emphasis>
</taxonomicName>
m. /
<typeStatus id="054D8878FF8AFF9E9FABF85AFAA3F825" box="[1233,1402,1976,2002]" pageId="22" pageNumber="356" type="holotype">HOLOTYPUS</typeStatus>
, P. JAŁOSZYŃSKI, 2020” [red, printed] (
<collectionCode id="BCE7AE1FFF8AFF9E9934F83EFD44F801" box="[590,669,2012,2038]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="22" pageNumber="356">EUMJ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8BFF9F9BBDFF74FB0FFF23" pageId="23" pageNumber="357" type="diagnosis">
<paragraph id="DA4936DAFF8BFF9F9BBDFF74FB0FFF23" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
<emphasis id="E882EAC8FF8BFF9F9BBDFF74FE9BFF47" bold="true" box="[199,322,150,176]" pageId="23" pageNumber="357">Diagnosis.</emphasis>
Eyes in male small, in dorsal view only half as long as tempora; aedeagus in ventral view about three times as long as broad, with apical margin bearing an elongate median subtrapezoidal projection.
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8BFF9F9BBDFF3CFE48FC1B" pageId="23" pageNumber="357" type="description">
<paragraph id="DA4936DAFF8BFF9F9BBDFF3CFE64FEEA" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
<emphasis id="E882EAC8FF8BFF9F9BBDFF3CFE80FF0F" bold="true" box="[199,345,222,248]" pageId="23" pageNumber="357">Description.</emphasis>
Body of male (
<figureCitation id="42CD2A5FFF8BFF9F9979FF3CFD8AFF0F" box="[515,595,222,248]" captionStart="FIGURES 4552" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[151,1436,862,1918]" captionTargetId="figure-321@13.[151,1436,849,1918]" captionTargetPageId="13" captionText="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." figureDoi="http://doi.org/10.5281/zenodo.4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="23" pageNumber="357">Fig. 49</figureCitation>
) strongly convex, uniformly light brown, covered with setae slightly lighter than cuticle; BL
<quantity id="1D0E9B3FFF8BFF9F9A28FEE0FE60FEEA" box="[338,441,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="23" pageNumber="357" unit="mm" value="0.83">0.83 mm</quantity>
.
</paragraph>
<paragraph id="DA4936DAFF8BFF9F9BBDFEC4FDC4FDCB" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
Head (
<figureCitation id="42CD2A5FFF8BFF9F9A6AFEC4FEB9FEB6" box="[272,352,294,321]" captionStart="FIGURES 4552" captionStartId="13.[151,264,1942,1966]" captionTargetBox="[151,1436,862,1918]" captionTargetId="figure-321@13.[151,1436,849,1918]" captionTargetPageId="13" captionText="FIGURES 4552. Dorsal habitus of males (45, 47, 49, 51) and head of male in dorsal view (46, 48, 50, 52). Nogunius sokani sp. n. (4546); N. aogashimanus sp. n. (4748); N. kerri sp. n. (4950); and N. fukuuzanus sp. n. (5152)." figureDoi="http://doi.org/10.5281/zenodo.4401631" httpUri="https://zenodo.org/record/4401631/files/figure.png" pageId="23" pageNumber="357">Fig. 50</figureCitation>
) broadest at eyes, HL
<quantity id="1D0E9B3FFF8BFF9F992FFEC4FD63FEB6" box="[597,698,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="357" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="1D0E9B3FFF8BFF9F9982FEC4FC87FEB6" box="[760,862,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="23" pageNumber="357" unit="mm" value="0.16">0.16 mm</quantity>
; tempora in dorsal view about twice as long as eyes; vertex and frons confluent, together weakly convex; each eye small, strongly convex, bean-shaped, with distinct posteromedian emargination, composed of 13 facets of equal diameters. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; tempora and posterior margin of vertex densely covered with thick bristles; additionally posterior margin of vertex with a pair of long lateral setae. Antennae slender, AnL
<quantity id="1D0E9B3FFF8BFF9F9FF5FE54FB2CFE26" box="[1167,1269,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="357" unit="mm" value="0.35">0.35 mm</quantity>
, antennomeres I and II each strongly elongate, IIIVIII each about as long as broad or indistinctly transverse, IX distinctly larger than VIII, slightly transverse, X much larger than IX, strongly transverse, XI indistinctly narrower and slightly longer than X, about as long as broad.
</paragraph>
<paragraph id="DA4936DAFF8BFF9F9BBDFDA5FAF2FD5F" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
Pronotum subconical with strongly rounded sides, broadest shortly in front of base; PL
<quantity id="1D0E9B3FFF8BFF9F9FE8FDA4FB2EFD96" box="[1170,1271,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="357" unit="mm" value="0.23">0.23 mm</quantity>
, PW
<quantity id="1D0E9B3FFF8BFF9F9E48FDA4FA40FD96" box="[1330,1433,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="357" unit="mm" value="0.2">0.20 mm</quantity>
. Anterior and posterior margins weakly convex, base with two pairs of small, distinct pits. Punctures on pronotal disc fine, inconspicuous; setae sparse, short, suberect; sides, especially posteriorly, with sparse, thick bristles.
</paragraph>
<paragraph id="DA4936DAFF8BFF9F9BBDFD50FC68FCE3" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
Elytra together oval, broadest indistinctly in front of middle; EL
<quantity id="1D0E9B3FFF8BFF9F98EDFD50FC25FD3A" box="[919,1020,690,717]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="357" unit="mm" value="0.45">0.45 mm</quantity>
, EW
<quantity id="1D0E9B3FFF8BFF9F9F43FD50FB47FD3A" box="[1081,1182,690,717]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="23" pageNumber="357" unit="mm" value="0.33">0.33 mm</quantity>
, EI 1.38; humeral calli weakly elevated, basal impressions large, deep, nearly circular; apices separately rounded. Punctures inconspicuous; setae slightly longer and denser than those on pronotum, suberect.
</paragraph>
<paragraph id="DA4936DAFF8BFF9F9BBDFCFDFD01FCCF" blockId="23.[151,1437,150,1293]" box="[199,728,798,825]" pageId="23" pageNumber="357">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="DA4936DAFF8BFF9F9BBDFCA0FF13FC3E" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
Aedeagus (
<figureCitation id="42CD2A5FFF8BFF9F9A39FCA0FE1AFCAB" box="[323,451,834,860]" captionStart="FIGURES 7784" captionStartId="21.[151,264,1915,1939]" captionTargetBox="[164,1432,189,1882]" captionTargetId="figure-19@21.[151,1436,181,1892]" captionTargetPageId="21" captionText="FIGURES 7784. Aedeagus in ventral (77, 79, 81, 83) and lateral (78, 80, 82, 84) views. Nogunius kerri sp. n. (7780); and N. fukuuzanus sp. n. (8184)." figureDoi="http://doi.org/10.5281/zenodo.4401643" httpUri="https://zenodo.org/record/4401643/files/figure.png" pageId="23" pageNumber="357">Figs 7780</figureCitation>
) elongate, in ventral view about three times as long as broad; AeL
<quantity id="1D0E9B3FFF8BFF9F9FD9FCA0FAD0FCAA" box="[1187,1289,834,861]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="357" unit="mm" value="0.13">0.13 mm</quantity>
; median lobe in ventral view nearly parallel-sided in proximal 2/3, then slightly narrowing toward truncate apex, apex at middle with large, elongate subtrapezoidal projection; parameres slender, each with one long apical and one long subapical seta.
</paragraph>
<paragraph id="DA4936DAFF8BFF9F9BBDFC30FE48FC1B" blockId="23.[151,1437,150,1293]" box="[199,401,978,1004]" pageId="23" pageNumber="357">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8BFF9F9BBDFC14FBBDFBE6" box="[199,1124,1014,1041]" pageId="23" pageNumber="357" type="distribution">
<paragraph id="DA4936DAFF8BFF9F9BBDFC14FBBDFBE6" blockId="23.[151,1437,150,1293]" box="[199,1124,1014,1041]" pageId="23" pageNumber="357">
<emphasis id="E882EAC8FF8BFF9F9BBDFC14FEB8FBE7" bold="true" box="[199,353,1014,1040]" pageId="23" pageNumber="357">Distribution.</emphasis>
Ishigaki-jima island of the Yaeyama group of the Ryukyus (
<figureCitation id="42CD2A5FFF8BFF9F9F7EFC14FB8EFBE6" box="[1028,1111,1014,1041]" captionStart="FIGURE 85" captionStartId="24.[151,250,1259,1283]" captionTargetBox="[151,1429,181,1230]" captionTargetId="figure-15@24.[151,1436,181,1236]" captionTargetPageId="24" captionText="FIGURE 85. Distribution of Nogunius." figureDoi="http://doi.org/10.5281/zenodo.4401647" httpUri="https://zenodo.org/record/4401647/files/figure.png" pageId="23" pageNumber="357">Fig. 85</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8BFF9F9BBDFBF8FB70FBAF" pageId="23" pageNumber="357" type="etymology">
<paragraph id="DA4936DAFF8BFF9F9BBDFBF8FB70FBAF" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
<emphasis id="E882EAC8FF8BFF9F9BBDFBF8FE97FBC3" bold="true" box="[199,334,1050,1076]" pageId="23" pageNumber="357">Etymology.</emphasis>
This species is dedicated to George H. Kerr, author of “
<collectingRegion id="1832F838FF8BFF9F98CEFBF8FBF9FBC3" box="[948,1056,1050,1076]" country="Japan" name="Okinawa" pageId="23" pageNumber="357">Okinawa</collectingRegion>
: The History of an Island People”, a book that introduced me into the fascinating history and rich culture of the Ryukyu Islands.
</paragraph>
</subSubSection>
<subSubSection id="92EC6551FF8BFF9F9BBDFB80FC16FAFA" pageId="23" pageNumber="357" type="discussion">
<paragraph id="DA4936DAFF8BFF9F9BBDFB80FC16FAFA" blockId="23.[151,1437,150,1293]" pageId="23" pageNumber="357">
<emphasis id="E882EAC8FF8BFF9F9BBDFB80FEE2FB8B" bold="true" box="[199,315,1122,1148]" pageId="23" pageNumber="357">Remarks.</emphasis>
<taxonomicName id="1DF64D59FF8BFF9F9A3BFB81FE30FB8B" authorityName="Jałoszyński" authorityYear="2020" box="[321,489,1122,1148]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="357" phylum="Arthropoda" rank="species" species="kerri">
<emphasis id="E882EAC8FF8BFF9F9A3BFB81FE30FB8B" box="[321,489,1122,1148]" italics="true" pageId="23" pageNumber="357">Nogunius kerri</emphasis>
</taxonomicName>
is the only species of this genus known to occur in the Yaeyama Islands. The body of male is stouter than that of all remaining species, and the eyes are conspicuously small, similar only to those of
<taxonomicName id="1DF64D59FF8BFF9F9EF9FB65FECCFB33" authorityName="Jałoszyński" authorityYear="2020" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="357" phylum="Arthropoda" rank="species" species="fukuuzanus">
<emphasis id="E882EAC8FF8BFF9F9EF9FB65FECCFB33" italics="true" pageId="23" pageNumber="357">N. fukuuzanus</emphasis>
</taxonomicName>
. The latter species occurs on Okinawa-jima,
<emphasis id="E882EAC8FF8BFF9F9874FB49FCC4FB32" box="[782,797,1195,1221]" italics="true" pageId="23" pageNumber="357"></emphasis>
<quantity id="1D0E9B3FFF8BFF9F985FFB48FCA3FB33" box="[805,890,1194,1220]" metricMagnitude="5" metricUnit="m" metricValue="4.7" pageId="23" pageNumber="357" unit="km" value="470.0">470 km</quantity>
NE of Ishigaki-jima, and males of the latter species are distinctly slenderer, including a narrower head, and have conspicuously elongate aedeagus, which is over five times as long as broad (only three times as long as broad in
<taxonomicName id="1DF64D59FF8BFF9F981DFB11FC67FAFB" authorityName="Jałoszyński" authorityYear="2020" box="[871,958,1266,1292]" class="Insecta" family="Scydmaenidae" genus="Nogunius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="357" phylum="Arthropoda" rank="species" species="kerri">
<emphasis id="E882EAC8FF8BFF9F981DFB11FC67FAFB" box="[871,958,1266,1292]" italics="true" pageId="23" pageNumber="357">N. kerri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>