144 lines
16 KiB
XML
144 lines
16 KiB
XML
<document id="0FE9B8AE0F6A04B02E40196FC42CD117" ID="10.11646/zootaxa.4169.2.2" ID-DOI="10.11646/zootaxa.4169.2.2" ID-GBIF-Dataset="5509767a-7e3a-4919-bb01-6b025bb293e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="260910" ID-ZooBank="53B27ED7-8CFA-4058-A47E-3A340ED322FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474353234057" checkinUser="plazi" docAuthor="Márquez, Juan & Asiain, Julieta" docDate="2016" docId="0E256050FF84CC2EFF34F918FAD2FE64" docLanguage="en" docName="zootaxa.4169.2.2.pdf" docOrigin="Zootaxa 4169 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyreocephalus puncticeps" docType="treatment" docVersion="8" lastPageNumber="264" masterDocId="F21C1828FF88CC23FFA3FF81FFDBFFEC" masterDocTitle="Taxonomy of the Mexican species of Thyreocephalus Guérin-Méneville (Coleoptera: Staphylinidae, Xantholinini)" masterLastPageNumber="285" masterPageNumber="251" pageNumber="263" updateTime="1698386383488" updateUser="plazi">
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<mods:title id="F73E9E439C35369ED132C400CABAB2E1">Taxonomy of the Mexican species of Thyreocephalus Guérin-Méneville (Coleoptera: Staphylinidae, Xantholinini)</mods:title>
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<mods:namePart id="C4F52AFC75B6316671B3EC44F3E03C94">Márquez, Juan</mods:namePart>
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<mods:namePart id="9B7E37FBAA84F451FBAE11FF594D76E3">Asiain, Julieta</mods:namePart>
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<mods:title id="A815F47AF75ADF71C1331752841FD0AD">Zootaxa</mods:title>
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<mods:date id="27A01FECBCA386AF9A43A6875385AAEB">2016</mods:date>
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<mods:number id="9200A3DDE4D3B9A5F7EB64E0C267784E">4169</mods:number>
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<treatment id="0E256050FF84CC2EFF34F918FAD2FE64" ID-DOI="http://doi.org/10.5281/zenodo.5623711" ID-GBIF-Taxon="124552125" ID-Zenodo-Dep="5623711" LSID="urn:lsid:plazi:treatment:0E256050FF84CC2EFF34F918FAD2FE64" httpUri="http://treatment.plazi.org/id/0E256050FF84CC2EFF34F918FAD2FE64" lastPageId="13" lastPageNumber="264" pageId="12" pageNumber="263">
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<subSubSection id="CE9682CDFF84CC2FFF34F918FE0CF840" pageId="12" pageNumber="263" type="nomenclature">
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<paragraph id="8633D146FF84CC2FFF34F918FE24F95F" blockId="12.[151,511,1689,1715]" box="[151,511,1689,1715]" pageId="12" pageNumber="263">
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<heading id="DD7B662AFF84CC2FFF34F918FE24F95F" bold="true" box="[151,511,1689,1715]" fontSize="11" level="1" pageId="12" pageNumber="263" reason="1">
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<emphasis id="B4F80D54FF84CC2FFF34F918FE24F95F" bold="true" box="[151,511,1689,1715]" pageId="12" pageNumber="263">
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The
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<taxonomicName id="418CAAC5FF84CC2FFF6CF918FE94F95F" ID-CoL="56RY7" box="[207,335,1689,1715]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="puncticeps">
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<emphasis id="B4F80D54FF84CC2FFF6CF918FE94F95F" bold="true" box="[207,335,1689,1715]" italics="true" pageId="12" pageNumber="263">puncticeps</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8633D146FF84CC2FFF34F961FE0CF840" blockId="12.[151,1437,1759,2037]" pageId="12" pageNumber="263">
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For the analyses of the following species, we propose the preliminary formation of the species group "
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<taxonomicName id="418CAAC5FF84CC2FFAB1F960FA51F914" box="[1298,1418,1761,1784]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="puncticeps">
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<emphasis id="B4F80D54FF84CC2FFAB1F960FA51F914" box="[1298,1418,1761,1784]" italics="true" pageId="12" pageNumber="263">puncticeps</emphasis>
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</taxonomicName>
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", that includes species with similar body size (
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<quantity id="41747CA3FF84CC2FFD24F885FCEBF8F0" box="[647,816,1796,1821]" metricMagnitude="-2" metricUnit="m" metricValue="1.785" metricValueMax="2.12" metricValueMin="1.45" pageId="12" pageNumber="263" unit="mm" value="17.85" valueMax="21.2" valueMin="14.5">14.5–21.2 mm</quantity>
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), pattern color (body black, with reddish elytra), having conspicuous umbilicate punctures (variably dense) on the dorsal and ventral surfaces, dense punctation on the elytra almost homogeneously distributed, and the prosternum with developed posteromedian carina. The
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<taxonomicName id="418CAAC5FF84CC2FFF34F8F0FED4F864" box="[151,271,1905,1928]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="puncticeps">
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<emphasis id="B4F80D54FF84CC2FFF34F8F0FED4F864" box="[151,271,1905,1928]" italics="true" pageId="12" pageNumber="263">puncticeps</emphasis>
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</taxonomicName>
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species group can include
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<taxonomicName id="418CAAC5FF84CC2FFDE1F8F0FD0DF864" box="[578,726,1905,1928]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="puncticeps">
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<emphasis id="B4F80D54FF84CC2FFDE1F8F0FD0DF864" box="[578,726,1905,1928]" italics="true" pageId="12" pageNumber="263">T. puncticeps</emphasis>
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</taxonomicName>
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(from which the name is taken),
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<taxonomicName id="418CAAC5FF84CC2FFBF3F8F0FB3BF864" box="[1104,1248,1905,1928]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="arizonicus">
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<emphasis id="B4F80D54FF84CC2FFBF3F8F0FB3BF864" box="[1104,1248,1905,1928]" italics="true" pageId="12" pageNumber="263">T. arizonicus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="418CAAC5FF84CC2FFB53F8F0FA4EF864" box="[1264,1429,1903,1928]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="cribripennis">
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<emphasis id="B4F80D54FF84CC2FFB53F8F0FA4EF864" box="[1264,1429,1903,1928]" italics="true" pageId="12" pageNumber="263">T. cribripennis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="418CAAC5FF84CC2FFF34F814FED7F841" box="[151,268,1940,1965]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="halffteri">
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<emphasis id="B4F80D54FF84CC2FFF34F814FED7F841" box="[151,268,1940,1965]" italics="true" pageId="12" pageNumber="263">T. halffteri</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="418CAAC5FF84CC2FFEE0F814FE0BF840" box="[323,464,1940,1964]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF84CC2FFEE0F814FE0BF840" box="[323,464,1940,1964]" italics="true" pageId="12" pageNumber="263">T. rufipennis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="CE9682CDFF84CC2EFF64F839FAD2FE64" lastPageId="13" lastPageNumber="264" pageId="12" pageNumber="263" type="description">
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<paragraph id="8633D146FF84CC2EFF64F839FED8FEF1" blockId="12.[151,1437,1759,2037]" lastBlockId="13.[151,1436,151,392]" lastPageId="13" lastPageNumber="264" pageId="12" pageNumber="263">
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Another aspect that can support the formation of this species group is their geographic distribution, that includes southern
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<collectingCountry id="FE9B91D6FF84CC2FFECAF85DFD55F819" box="[361,654,2012,2037]" name="United States of America" pageId="12" pageNumber="263">United States of America</collectingCountry>
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(
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<collectingRegion id="44481FA4FF84CC2FFD3CF85DFD20F819" box="[671,763,2012,2037]" country="United States of America" name="Arizona" pageId="12" pageNumber="263">Arizona</collectingRegion>
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,
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<taxonomicName id="418CAAC5FF84CC2FFCAFF85CFC45F818" box="[780,926,2013,2036]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="arizonicus">
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<emphasis id="B4F80D54FF84CC2FFCAFF85CFC45F818" box="[780,926,2013,2036]" italics="true" pageId="12" pageNumber="263">T. arizonicus</emphasis>
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</taxonomicName>
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) to
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<collectingRegion id="44481FA4FF84CC2FFC71F85DFB90F819" box="[978,1099,2012,2037]" country="Guatemala" name="Guatemala" pageId="12" pageNumber="263">
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<collectingCountry id="FE9B91D6FF84CC2FFC71F85DFB90F819" box="[978,1099,2012,2037]" name="Guatemala" pageId="12" pageNumber="263">Guatemala</collectingCountry>
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</collectingRegion>
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(
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<taxonomicName id="418CAAC5FF84CC2FFBFDF85CFB34F818" box="[1118,1263,2012,2036]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF84CC2FFBFDF85CFB34F818" box="[1118,1263,2012,2036]" italics="true" pageId="12" pageNumber="263">T. rufipennis</emphasis>
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</taxonomicName>
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), a geographic area known as the Mexican transition zone (
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<bibRefCitation id="E21DACB7FF85CC2EFD2AFF16FCFEFF5C" author="Halffter" box="[649,805,151,176]" pageId="13" pageNumber="264" refString="Halffter, G. (1987) Biogeography of the montane entomofauna of Mexico and Central America. Annual Review of Entomology, 32, 95 - 114. http: // dx. doi. org / 10.1146 / annurev. en. 32.010187.000523" type="journal article" year="1987">Halffter 1987</bibRefCitation>
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,
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<bibRefCitation id="E21DACB7FF85CC2EFC92FF19FB85FF5C" author="Morrone" box="[817,1118,152,176]" pageId="13" pageNumber="264" refString="Morrone, J. J. & Marquez, J. (2001) Halffter's Mexican Transition Zone, beetle generalized tracks, and geographical homology. Journal of Biogeography, 28 (5), 635 - 650. http: // dx. doi. org / 10.1046 / j. 1365 - 2699.2001.00571. x" type="journal article" year="2001">Morrone & Márquez 2001</bibRefCitation>
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,
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<bibRefCitation id="E21DACB7FF85CC2EFBC9FF19FAD4FF5C" author="Morrone" box="[1130,1295,151,176]" pageId="13" pageNumber="264" refString="Morrone, J. J. (2006) Biogeographic areas and transition zones of Latin America and the Caribbean Islands based on panbiogeographic and cladistic analyses of the entomofauna. Annual Review of Entomology, 51, 467 - 494. http: // dx. doi. org / 10.1146 / annurev. ento. 50.071803.130447" type="journal article" year="2006">Morrone 2006</bibRefCitation>
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,
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<bibRefCitation id="E21DACB7FF85CC2EFAB9FF16FA8CFF5C" author="Morrone" box="[1306,1367,151,176]" pageId="13" pageNumber="264" refString="Morrone, J. J. (2015) Halffter's Mexican transition zone (1962 - 2014), cenocrons and evolutionary biogeography. Journal of Zoological Systematics and Evolutionary Research, 53 (3), 249 - 257. http: // dx. doi. org / 10.1111 / jzs. 12098" type="journal article" year="2015">2015</bibRefCitation>
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). This distributional pattern has been observed in diverse biological groups, including some species of
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<taxonomicName id="418CAAC5FF85CC2EFB70FF3DFAB7FF39" box="[1235,1388,188,213]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
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and other coleopteran groups (
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<bibRefCitation id="E21DACB7FF85CC2EFE1EFF61FD09FF14" author="Marquez" box="[445,722,223,248]" pageId="13" pageNumber="264" refString="Marquez, J. & Asiain, J. (2006) Patrones de distribucion de la familia Staphylinidae (Coleoptera). In: Morrone, J. J. & Llorente Bousquets, J. (Eds.), Componentes bioticos principales de la Entomofauna Mexicana. Las Prensas de Ciencias, UNAM, Mexico, D. F., pp. 157 - 236." type="book chapter" year="2006">Márquez & Asiain 2006</bibRefCitation>
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, 2012,
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<bibRefCitation id="E21DACB7FF85CC2EFC87FF61FBE0FF14" author="Moron" box="[804,1083,224,248]" pageId="13" pageNumber="264" refString="Moron, M. A. & Marquez, J. (2012) Nuevos registros estatales y nacionales de escarabajos (Coleoptera: Scarabaeoidea) y comentarios sobre su distribucion. Revista Mexicana de Biodiversidad, 83 (3), 698 - 711. http: // dx. doi. org / 10.7550 / rmb. 28386" type="journal article" year="2012">Morón & Márquez 2012</bibRefCitation>
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||
,
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<bibRefCitation id="E21DACB7FF85CC2EFBE5FF5EFACAFF14" author="Asiain" box="[1094,1297,223,248]" pageId="13" pageNumber="264" refString="Asiain, J., Marquez, J. & Irmler, U. (2015) New national and state records of Neotropical Staphylinidae (Insecta: Coleoptera). Zootaxa, 3974 (1), 76 - 92. http: // dx. doi. org / 10.11646 / zootaxa. 3974.1.5" type="journal article" year="2015">
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Asiain
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<emphasis id="B4F80D54FF85CC2EFB35FF60FB1CFF14" box="[1174,1223,223,248]" italics="true" pageId="13" pageNumber="264">et al</emphasis>
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||
. 2015
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||
</bibRefCitation>
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, Márquez
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<emphasis id="B4F80D54FF85CC2EFA2BFF60FF76FEF1" italics="true" pageId="13" pageNumber="264">et al</emphasis>
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. 2015).
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</paragraph>
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<paragraph id="8633D146FF85CC2EFF64FEA9FAD2FE64" blockId="13.[151,1436,151,392]" pageId="13" pageNumber="264">
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To avoid repeating information, the species of the
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<taxonomicName id="418CAAC5FF85CC2EFCB0FEA8FC50FEAC" box="[787,907,297,320]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="species" species="puncticeps">
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<emphasis id="B4F80D54FF85CC2EFCB0FEA8FC50FEAC" box="[787,907,297,320]" italics="true" pageId="13" pageNumber="264">puncticeps</emphasis>
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</taxonomicName>
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group will not be redescribed, but their most relevant characters for their identification are summarized in the
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<tableCitation id="CB0EE4FDFF85CC2EFC29FECDFC39FE88" box="[906,994,332,357]" captionStart="TABLE 3" captionStartId="29.[125,204,557,578]" captionText="TABLE 3. Principal taxonomic characters for the iđentification of the Mexican species of Thyreocephalus of the punticeps group." pageId="13" pageNumber="264">Table 3</tableCitation>
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. In addition, a comparative diagnosis, discussion, natural history, geographic distribution and material studied are included for each species.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |