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<document id="CB1F77B17B8871D228F90D4441F5EB9B" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3BCFF2FFF75F9B1BD6FFE4A" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madagaster procarina Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="191" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="188" updateTime="1698480684898" updateUser="ExternalLinkService">
<mods:mods id="2F24F0C3D84D63B2961B657C4B18FA3B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C90E2B7F56AC561E33433231015E36EB">
<mods:title id="B0A90EAFA2CCAFA7CA23D82C085E88FA">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="51CDC0455D110CBEB88DD891828E847C" type="personal">
<mods:role id="C70E5F093FB10696D6EC19092D540688">
<mods:roleTerm id="F79C866381B22A8DB3CD6C2AAD1D8FC0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31C796D3548A3C68D21A067147BA9203">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="05B29B370DBE1CEB05ADFD7203A0D97E">text</mods:typeOfResource>
<mods:relatedItem id="3633E7FB8A54959B00544810A2D45481" type="host">
<mods:titleInfo id="AFE23EC09DE7D9AD9F74F7DF38A4C0BB">
<mods:title id="98D6562CEBABC37390DC8CFE29662FD8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B5A122DC5BBEB4CB738957123BF0E898">
<mods:date id="927123B67B87885CAF4BCF2F0477E4D6">2017</mods:date>
<mods:detail id="197941167C0B30B5DCAAB7D359091024" type="pubDate">
<mods:number id="820583882D75797EBC719941AFD975E8">2017-11-03</mods:number>
</mods:detail>
<mods:detail id="CDD6040F4B31DE6A6CB7E1E43FFCC120" type="volume">
<mods:number id="5CB1FD39A85F928BBF598B8D7D70CB3D">4342</mods:number>
</mods:detail>
<mods:detail id="7839D16D508AA239542CA7A311677AB8" type="issue">
<mods:number id="1C7E30327CCC90B85F719DA0C046DD56">1</mods:number>
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<mods:extent id="ABD9417051D9F23AD3393882C6CE3D33" unit="page">
<mods:start id="AF07E2DBB7AB49F7F8D7D8F51859EB12">1</mods:start>
<mods:end id="3C96910B3FABE37A6582435A902C9B90">264</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="25098AEDFAA275569BC88A1198348025">journal article</mods:classification>
<mods:identifier id="8B611B9C53CE1ADD997E8584FFCCB820" type="CLB-Dataset">31639</mods:identifier>
<mods:identifier id="23D7BBC882DF999D083624EDBD48510A" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
<mods:identifier id="ED00E931EAC1F6AEC481A3D31D6AD10D" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
<mods:identifier id="D9AE27ECC76D171BF67B59B3740B3164" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8C359E68E12E85EE04626A5AB98B41D7" type="Zenodo-Dep">1041066</mods:identifier>
<mods:identifier id="F5F52279980E81AB3E3C16D4A0DBA9D7" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
</mods:mods>
<treatment id="038587BBE3BCFF2FFF75F9B1BD6FFE4A" ID-DOI="http://doi.org/10.5281/zenodo.5323736" ID-GBIF-Taxon="137233384" ID-Zenodo-Dep="5323736" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A" httpUri="http://treatment.plazi.org/id/038587BBE3BCFF2FFF75F9B1BD6FFE4A" lastPageId="190" lastPageNumber="191" pageId="187" pageNumber="188">
<subSubSection id="C3366526E3BCFF2AFF75F9B1BB98F9CA" pageId="187" pageNumber="188" type="nomenclature">
<paragraph id="8B9336ADE3BCFF2AFF75F9B1BB7FF9AB" blockId="187.[151,676,1556,1615]" box="[151,579,1556,1582]" pageId="187" pageNumber="188">
<heading id="D0DB81C1E3BCFF2AFF75F9B1BB7FF9AB" bold="true" box="[151,579,1556,1582]" fontSize="11" level="1" pageId="187" pageNumber="188" reason="1">
<emphasis id="B958EABFE3BCFF2AFF75F9B1BB7FF9AB" bold="true" box="[151,579,1556,1582]" pageId="187" pageNumber="188">
<emphasis id="B958EABFE3BCFF2AFF75F9B1B890F9AB" bold="true" box="[151,428,1556,1582]" italics="true" pageId="187" pageNumber="188">
<taxonomicName id="4C2C4D2EE3BCFF2AFF75F9B1B899F9AB" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,421,1556,1582]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="187" pageNumber="188" phylum="Arthropoda" rank="species" species="procarina" status="sp. nov.">Madagaster procarina</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26B57C4E3BCFF2AFE51F9B1BB7FF9AB" box="[435,579,1556,1582]" pageId="187" pageNumber="188" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE3BCFF2AFF75F993BB98F9CA" blockId="187.[151,676,1556,1615]" box="[151,676,1590,1615]" pageId="187" pageNumber="188">
<figureCitation id="13172A28E3BCFF2AFF75F993B9C3F9CA" box="[151,255,1590,1615]" captionStart="FIGURE 180" captionStartId="188.[151,250,1601,1623]" captionText="FIGURE 180. Madagaster procarina, holotype, habitus dorsal, lateral, and ventral." pageId="187" pageNumber="188">Figs. 180</figureCitation>
(habitus), 182 (aedeagus), 301 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3BCFF2DFF75F9DBBB29FEC5" lastPageId="188" lastPageNumber="189" pageId="187" pageNumber="188" type="materials_examined">
<paragraph id="8B9336ADE3BCFF2AFF75F9DBBCA1F85E" blockId="187.[151,1437,1662,2011]" pageId="187" pageNumber="188">
<materialsCitation id="3B443CF0E3BCFF2AFF75F9DBBC26F93E" ID-GBIF-Occurrence="1697157764" collectingDate="1990-03-25" collectionCode="USNM" collectorName="W. E. Steiner &amp; C. Kremen &amp; V. Razafimahatratra" elevation="1400" latitude="-20.716667" location="Ambositra" longLatPrecision="1266" longitude="47.266666" pageId="187" pageNumber="188" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B958EABFE3BCFF2AFF75F9DBB882F912" bold="true" box="[151,446,1662,1687]" pageId="187" pageNumber="188">
<typeStatus id="5497880FE3BCFF2AFF75F9DBB9EEF912" box="[151,210,1662,1687]" pageId="187" pageNumber="188">Type</typeStatus>
Material.
<typeStatus id="5497880FE3BCFF2AFEB3F9DBB882F912" box="[337,446,1662,1687]" pageId="187" pageNumber="188" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE3BCFF2AFDC2F9DBBB83F912" box="[544,703,1662,1687]" country="Madagascar" name="Fianarantsoa" pageId="187" pageNumber="188">
<emphasis id="B958EABFE3BCFF2AFDC2F9DBBB83F912" bold="true" box="[544,703,1662,1687]" pageId="187" pageNumber="188">Fianarantsoa</emphasis>
</collectingRegion>
,
<quantity id="4CD49B48E3BCFF2AFD2CF9DABA2AF912" box="[718,790,1662,1687]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="187" pageNumber="188" unit="km" value="32.0">32 km</quantity>
S
<location id="8EF36076E3BCFF2AFCD4F9DBBA93F912" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BCFF2AFCD4F9DBBA93F912" box="[822,943,1662,1687]" latitude="-20.716667" longLatPrecision="1266" longitude="47.266666" name="Ambositra" pageId="187" pageNumber="188" stateProvince="Fianarantsoa">Ambositra</location>
, in seepage over exposed granite cliff face, elev.
<elevation id="0001D19EE3BCFF2AFF30F906B814F93F" box="[210,296,1699,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="187" pageNumber="188" unit="m" value="1400.0">
<quantity id="4CD49B48E3BCFF2AFF30F906B814F93F" box="[210,296,1699,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="187" pageNumber="188" unit="m" value="1400.0">1400 m</quantity>
</elevation>
,
<geoCoordinate id="EE18506AE3BCFF2AFED6F906B8A7F93E" box="[308,411,1699,1723]" direction="south" orientation="latitude" pageId="187" pageNumber="188" precision="925" value="-20.716667">20° 43' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE3BCFF2AFE4AF906BB2EF93F" box="[424,530,1698,1723]" direction="east" orientation="longitude" pageId="187" pageNumber="188" precision="925" value="47.266666">47° 16' E</geoCoordinate>
,
<collectingDate id="EFD6E985E3BCFF2AFDFFF907BBA5F93E" box="[541,665,1698,1723]" pageId="187" pageNumber="188" value="1990-03-25">
<date id="FF92106DE3BCFF2AFDFFF907BBA5F93E" box="[541,665,1698,1723]" pageId="187" pageNumber="188" value="1990-03-25">25 iii 1990</date>
</collectingDate>
,
<collectorName id="26D9537BE3BCFF2AFD46F906BA05F93E" box="[676,825,1698,1723]" pageId="187" pageNumber="188">W. E. Steiner</collectorName>
,
<collectorName id="26D9537BE3BCFF2AFCA6F906BAFEF93F" box="[836,962,1699,1723]" pageId="187" pageNumber="188">C. Kremen</collectorName>
,
<collectorName id="26D9537BE3BCFF2AFC2FF906BD90F93E" box="[973,1196,1698,1723]" pageId="187" pageNumber="188">V. Razafimahatratra</collectorName>
(
<collectionCode id="ED3DAE68E3BCFF2AFB59F906BC2EF93E" box="[1211,1298,1699,1723]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="187" pageNumber="188">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B443CF0E3BCFF2AFAC6F907BDB6F8A2" ID-GBIF-Occurrence="1697157744" collectingDate="2012-11-28" collectionCode="USNM, NHRS" collectorName="Anstaba, S &amp; J. Bergsten &amp; R. Bukontaite &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" county="Same" elevation="296" location="Antsaba" longitude="13.60931" municipality="Diana" pageId="187" pageNumber="188" specimenCode="MAD12-31" specimenCount="2" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="5497880FE3BCFF2AFAC6F907BCA0F93E" box="[1316,1436,1698,1723]" pageId="187" pageNumber="188" type="paratype">
<emphasis id="B958EABFE3BCFF2AFAC6F907BCA0F93E" bold="true" box="[1316,1436,1698,1723]" pageId="187" pageNumber="188">Paratypes</emphasis>
</typeStatus>
(26):
<collectingCounty id="62F24E21E3BCFF2AFF3BF962B82BF95A" box="[217,279,1735,1759]" pageId="187" pageNumber="188">Same</collectingCounty>
data as
<typeStatus id="5497880FE3BCFF2AFE9DF963B8DDF95A" box="[383,481,1734,1759]" pageId="187" pageNumber="188" type="holotype">holotype</typeStatus>
(2
<collectionCode id="ED3DAE68E3BCFF2AFDEDF962BB59F95A" box="[527,613,1735,1759]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="187" pageNumber="188">USNM</collectionCode>
).
<collectingRegion id="49E8F84FE3BCFF2AFD9EF963BA2DF95A" box="[636,785,1734,1759]" country="Madagascar" name="Antsiranana" pageId="187" pageNumber="188">
<emphasis id="B958EABFE3BCFF2AFD9EF963BA2DF95A" bold="true" box="[636,785,1734,1759]" pageId="187" pageNumber="188">Antsiranana</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BF7ACD7E3BCFF2AFCC6F963BA57F95A" box="[804,875,1734,1759]" pageId="187" pageNumber="188">Diana</collectingMunicipality>
:
<location id="8EF36076E3BCFF2AFC98F963BAE6F95A" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BCFF2AFC98F963BAE6F95A" box="[890,986,1734,1759]" county="Same" longitude="13.60931" municipality="Diana" name="Antsaba" pageId="187" pageNumber="188" stateProvince="Antsiranana">Antsaba</location>
:
<location id="8EF36076E3BCFF2AFC0BF963BD82F95A" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BCFF2AFC0BF963BD82F95A" box="[1001,1214,1734,1759]" county="Same" longitude="13.60931" municipality="Diana" name="Galoko mountains" pageId="187" pageNumber="188" stateProvince="Antsiranana">Galoko mountains</location>
,
<quantity id="4CD49B48E3BCFF2AFB2CF962BC1CF95A" box="[1230,1312,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="187" pageNumber="188" unit="km" value="3.4">3.4 km</quantity>
NW from
<collectorName id="26D9537BE3BCFF2AFF75F94FB833F886" box="[151,271,1770,1795]" pageId="187" pageNumber="188">Anstaba, S</collectorName>
<geoCoordinate id="EE18506AE3BCFF2AFEECF94FB8AEF886" box="[270,402,1770,1795]" direction="east" orientation="longitude" pageId="187" pageNumber="188" precision="1" value="13.60931">13.60931 E</geoCoordinate>
48.72129, aspirator, forceps, sieves: hygropetric rocks and pools, elev.
<elevation id="0001D19EE3BCFF2AFB4CF94FBDC4F887" box="[1198,1272,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="187" pageNumber="188" unit="m" value="296.0">
<quantity id="4CD49B48E3BCFF2AFB4CF94FBDC4F887" box="[1198,1272,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="187" pageNumber="188" unit="m" value="296.0">296 m</quantity>
</elevation>
,
<collectingDate id="EFD6E985E3BCFF2AFAE6F94EBC43F886" box="[1284,1407,1770,1795]" pageId="187" pageNumber="188" value="2012-11-28">
<date id="FF92106DE3BCFF2AFAE6F94EBC43F886" box="[1284,1407,1770,1795]" pageId="187" pageNumber="188" value="2012-11-28">28 xi 2012</date>
</collectingDate>
,
<collectorName id="26D9537BE3BCFF2AFA68F94EB9C1F8A2" pageId="187" pageNumber="188">J. Bergsten</collectorName>
,
<collectorName id="26D9537BE3BCFF2AFEE5F8AAB89AF8A2" box="[263,422,1806,1831]" pageId="187" pageNumber="188">R. Bukontaite</collectorName>
,
<collectorName id="26D9537BE3BCFF2AFE52F8AABBB5F8A2" box="[432,649,1806,1831]" pageId="187" pageNumber="188">J.H. Randriamihaja</collectorName>
&amp;
<collectorName id="26D9537BE3BCFF2AFD4EF8AABA4FF8A2" box="[684,883,1806,1831]" pageId="187" pageNumber="188">T. Ranarilalatiana</collectorName>
(
<specimenCode id="DB8A9ED6E3BCFF2AFC63F8AABD3AF8A2" box="[897,1030,1807,1831]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="187" pageNumber="188">MAD12-31</specimenCode>
) (6
<collectionCode id="ED3DAE68E3BCFF2AFBD1F8AABDBFF8A2" box="[1075,1155,1807,1831]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="187" pageNumber="188">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B443CF0E3BCFF2AFB77F8ABBCA1F85E" ID-GBIF-Occurrence="1697157784" collectingDate="2011-04-11" collectingDateMax="2014-11-01" collectingDateMin="2011-04-11" collectionCode="NHRS, NMW" collectorName="J. Bergsten &amp; T. Ranarilalatiana &amp; S. Holmgren &amp; Andringitra NP &amp; R. Gerecke" county="Amoron'i Mania" elevation="1700" latitude="-22.133333" location="Ankazomivady forest" longLatPrecision="1261" longitude="46.883335" municipality="Ambalamanakana" pageId="187" pageNumber="188" specimenCode="MAD14-02" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingRegion id="49E8F84FE3BCFF2AFB77F8ABBC1BF8A2" box="[1173,1319,1806,1831]" country="Madagascar" name="Fianarantsoa" pageId="187" pageNumber="188">Fianarantsoa</collectingRegion>
,
<collectingCounty id="62F24E21E3BCFF2AFAD0F8ABB9DEF8CE" pageId="187" pageNumber="188">Amoron'i Mania</collectingCounty>
,
<quantity id="4CD49B48E3BCFF2AFF17F897B81BF8CE" box="[245,295,1842,1867]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="187" pageNumber="188" unit="km" value="3.0">3km</quantity>
south of
<collectingMunicipality id="6BF7ACD7E3BCFF2AFE49F897BB49F8CE" box="[427,629,1842,1867]" pageId="187" pageNumber="188">Ambalamanakana</collectingMunicipality>
next to RN7,
<location id="8EF36076E3BCFF2AFCDAF897BD0AF8CE" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BCFF2AFCDAF897BD0AF8CE" box="[824,1078,1842,1867]" county="Amoron'i Mania" latitude="-22.133333" longLatPrecision="1261" longitude="46.883335" municipality="Ambalamanakana" name="Ankazomivady forest" pageId="187" pageNumber="188" stateProvince="Fianarantsoa">Ankazomivady forest</location>
,
<geoCoordinate id="EE18506AE3BCFF2AFBA8F896BD88F8CE" box="[1098,1204,1843,1867]" direction="south" orientation="latitude" pageId="187" pageNumber="188" precision="5" value="-20.7722">20.7722S</geoCoordinate>
<geoCoordinate id="EE18506AE3BCFF2AFB26F896BC0FF8CE" box="[1220,1331,1843,1867]" direction="east" orientation="longitude" pageId="187" pageNumber="188" precision="5" value="47.1809">47.1809E</geoCoordinate>
,
<elevation id="0001D19EE3BCFF2AFAA5F896BCA4F8CE" box="[1351,1432,1843,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="187" pageNumber="188" unit="m" value="1700.0">
<quantity id="4CD49B48E3BCFF2AFAA5F896BCA4F8CE" box="[1351,1432,1843,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="187" pageNumber="188" unit="m" value="1700.0">1700m</quantity>
</elevation>
, hygropetric rocks and marsh with vegetation, elev.
<elevation id="0001D19EE3BCFF2AFD1FF8F2BA65F8EB" box="[765,857,1879,1903]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="187" pageNumber="188" unit="m" value="1700.0">
<quantity id="4CD49B48E3BCFF2AFD1FF8F2BA65F8EB" box="[765,857,1879,1903]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="187" pageNumber="188" unit="m" value="1700.0">1700 m</quantity>
</elevation>
,
<collectingDate id="EFD6E985E3BCFF2AFC89F8F2BAD8F8EA" box="[875,996,1878,1903]" pageId="187" pageNumber="188" value="2014-11-01">
<date id="FF92106DE3BCFF2AFC89F8F2BAD8F8EA" box="[875,996,1878,1903]" pageId="187" pageNumber="188" value="2014-11-01">1 xi 2014</date>
</collectingDate>
,
<collectorName id="26D9537BE3BCFF2AFC16F8F2BD45F8EA" box="[1012,1145,1879,1903]" pageId="187" pageNumber="188">J. Bergsten</collectorName>
,
<collectorName id="26D9537BE3BCFF2AFB68F8F2BC6AF8EA" box="[1162,1366,1878,1903]" pageId="187" pageNumber="188">T. Ranarilalatiana</collectorName>
&amp;
<collectorName id="26D9537BE3BCFF2AFA64F8F2B835F816" pageId="187" pageNumber="188">S. Holmgren</collectorName>
(
<specimenCode id="DB8A9ED6E3BCFF2AFEFAF8DEB8A1F816" box="[280,413,1915,1939]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="187" pageNumber="188">MAD14-02</specimenCode>
) (1
<collectionCode id="ED3DAE68E3BCFF2AFE2EF8DEBB27F816" box="[460,539,1915,1939]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="187" pageNumber="188">NHRS</collectionCode>
);
<collectorName id="26D9537BE3BCFF2AFDCDF8DFBBE1F817" box="[559,733,1914,1939]" pageId="187" pageNumber="188">Andringitra NP</collectorName>
hygropetric cascade stream crossing way to cascades, 16.2°C, elev.
<elevation id="0001D19EE3BCFF2AFF37F83AB812F832" box="[213,302,1951,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="187" pageNumber="188" unit="m" value="1722.0">
<quantity id="4CD49B48E3BCFF2AFF37F83AB812F832" box="[213,302,1951,1975]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="187" pageNumber="188" unit="m" value="1722.0">1722 m</quantity>
</elevation>
,
<geoCoordinate id="EE18506AE3BCFF2AFEDFF83AB8A7F832" box="[317,411,1951,1975]" direction="south" orientation="latitude" pageId="187" pageNumber="188" precision="925" value="-22.133333">22° 8' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE3BCFF2AFE49F83BBB26F832" box="[427,538,1950,1975]" direction="east" orientation="longitude" pageId="187" pageNumber="188" precision="925" value="46.883335">46° 53' E</geoCoordinate>
,
<collectingDate id="EFD6E985E3BCFF2AFDCAF83ABB9AF832" box="[552,678,1950,1975]" pageId="187" pageNumber="188" value="2011-04-11">
<date id="FF92106DE3BCFF2AFDCAF83ABB9AF832" box="[552,678,1950,1975]" pageId="187" pageNumber="188" value="2011-04-11">11 iv 2011</date>
</collectingDate>
,
<collectorName id="26D9537BE3BCFF2AFD56F805BA08F832" box="[692,820,1950,1975]" pageId="187" pageNumber="188">R. Gerecke</collectorName>
(MD 198) (7
<collectionCode id="ED3DAE68E3BCFF2AFC3FF83ABD16F832" box="[989,1066,1951,1975]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="187" pageNumber="188">NMW</collectionCode>
);
<location id="8EF36076E3BCFF2AFBDDF83BBC71F832" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BCFF2AFBDDF83BBC71F832" box="[1087,1357,1950,1975]" county="Amoron'i Mania" latitude="-22.133333" longLatPrecision="1261" longitude="46.883335" municipality="Ambalamanakana" name="Isalo Canyon de Makis" pageId="187" pageNumber="188" stateProvince="Fianarantsoa">Isalo Canyon de Makis</location>
, sandy bottom of river, with side pools and hygropetric sections at sides, wood in water, N-22° 32.922, E45° 24.064, elev.
</materialsCitation>
</paragraph>
<paragraph id="8B9336ADE3BBFF2DFF75FF3DBB29FEC5" blockId="188.[151,1436,151,320]" pageId="188" pageNumber="189">
<quantity id="4CD49B48E3BBFF2DFF75FF3DB9DFFF2A" box="[151,227,152,176]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="188" pageNumber="189" unit="m" value="780.0">780 m</quantity>
, 22° 33' S, 45° 24' E,
<date id="FF92106DE3BBFF2DFE09FF3DBB5EFF35" box="[491,610,151,176]" pageId="188" pageNumber="189" value="2006-05-11">11 v 2006</date>
, Bergsten
<emphasis id="B958EABFE3BBFF2DFD3FFF3CBA24FF35" box="[733,792,151,176]" italics="true" pageId="188" pageNumber="189">et al.</emphasis>
(P41E) (8 NHRS);
<materialsCitation id="3B443CF0E3BBFF2DFBE3FF32BDE0FF7D" ID-GBIF-Occurrence="1697157734" collectingDate="2012-11-13" collectionCode="NHRS" collectorName="Isalo NP &amp; Canyon de Makis, GB &amp; R. Bukontaite &amp; J. H. Randriamihaja" elevation="50" latitude="-22.55" location="Ihorombe" longLatPrecision="1259" longitude="45.4" municipality="Canyon de Makis" pageId="188" pageNumber="189" specimenCode="MAD12-05" specimenCount="1">
<location id="8EF36076E3BBFF2DFBE3FF32BD4DFF35" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BBFF2DFBE3FF32BD4DFF35" box="[1025,1137,151,176]" latitude="-22.55" longLatPrecision="1259" longitude="45.4" municipality="Canyon de Makis" name="Ihorombe" pageId="188" pageNumber="189">Ihorombe</location>
,
<collectorName id="26D9537BE3BBFF2DFB9DFF32BDDFFF2A" box="[1151,1251,151,176]" pageId="188" pageNumber="189">Isalo NP</collectorName>
<elevation id="0001D19EE3BBFF2DFB0EFF32BC22FF35" box="[1260,1310,151,176]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="188" pageNumber="189" unit="m" value="50.0">
<quantity id="4CD49B48E3BBFF2DFB0EFF32BC22FF35" box="[1260,1310,151,176]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="188" pageNumber="189" unit="m" value="50.0">50m</quantity>
</elevation>
before the entrance of
<collectorName id="26D9537BE3BBFF2DFEFEFF19BB25FF51" box="[284,537,188,213]" pageId="188" pageNumber="189">
<collectingMunicipality id="6BF7ACD7E3BBFF2DFEFEFF19B8DAFF50" box="[284,486,188,213]" pageId="188" pageNumber="189">Canyon de Makis</collectingMunicipality>
, GB
</collectorName>
nets and sieves, canyon river with side pools, S22.48660, E45.37954, elev.
<elevation id="0001D19EE3BBFF2DFA90FF19B993FF72" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="188" pageNumber="189" unit="m" value="752.0">
<quantity id="4CD49B48E3BBFF2DFA90FF19B993FF72" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="188" pageNumber="189" unit="m" value="752.0">752 m</quantity>
</elevation>
,
<geoCoordinate id="EE18506AE3BBFF2DFF58FF45B81DFF7D" box="[186,289,224,248]" direction="south" orientation="latitude" pageId="188" pageNumber="189" precision="925" value="-22.55">22° 33' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE3BBFF2DFECFFF7AB8AAFF72" box="[301,406,223,248]" direction="east" orientation="longitude" pageId="188" pageNumber="189" precision="925" value="45.4">45° 24' E</geoCoordinate>
,
<collectingDate id="EFD6E985E3BBFF2DFE43FF7ABB27FF7D" box="[417,539,223,248]" pageId="188" pageNumber="189" value="2012-11-13">
<date id="FF92106DE3BBFF2DFE43FF7ABB27FF7D" box="[417,539,223,248]" pageId="188" pageNumber="189" value="2012-11-13">13.xi.2012</date>
</collectingDate>
,
<collectorName id="26D9537BE3BBFF2DFDC4FF45BBFEFF7D" box="[550,706,223,248]" pageId="188" pageNumber="189">R. Bukontaite</collectorName>
&amp;
<collectorName id="26D9537BE3BBFF2DFD07FF45BAF8FF7D" box="[741,964,223,248]" pageId="188" pageNumber="189">J. H. Randriamihaja</collectorName>
(
<specimenCode id="DB8A9ED6E3BBFF2DFC31FF7ABD64FF7D" box="[979,1112,223,248]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="188" pageNumber="189">MAD12-05</specimenCode>
) (1
<collectionCode id="ED3DAE68E3BBFF2DFB67FF45BDE8FF7D" box="[1157,1236,224,248]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="188" pageNumber="189">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B443CF0E3BBFF2DFB05FF7ABB29FEC5" ID-GBIF-Occurrence="1697157750" collectingDate="2004-11-01" collectingDateMax="2004-12-29" collectingDateMin="2004-11-01" collectionCode="MNCN, DNA" collectorName="M. Balke" elevation="303" latitude="-13.65" location="Antsaba" longLatPrecision="1290" longitude="48.75" municipality="Galoko Mountains" pageId="188" pageNumber="189" specimenCode="AI546" specimenCount="1">
<location id="8EF36076E3BBFF2DFB05FF7ABC79FF7D" LSID="urn:lsid:plazi:treatment:038587BBE3BCFF2FFF75F9B1BD6FFE4A:8EF36076E3BBFF2DFB05FF7ABC79FF7D" box="[1255,1349,223,248]" latitude="-13.65" longLatPrecision="1290" longitude="48.75" municipality="Galoko Mountains" name="Antsaba" pageId="188" pageNumber="189">Antsaba</location>
,
<collectingMunicipality id="6BF7ACD7E3BBFF2DFAABFF7AB82EFE98" pageId="188" pageNumber="189">Galoko Mountains</collectingMunicipality>
, hygropetric cascade, elev.
<elevation id="0001D19EE3BBFF2DFDA0FEA1BBB7FE99" box="[578,651,260,284]" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="188" pageNumber="189" unit="m" value="303.0">
<quantity id="4CD49B48E3BBFF2DFDA0FEA1BBB7FE99" box="[578,651,260,284]" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="188" pageNumber="189" unit="m" value="303.0">303 m</quantity>
</elevation>
,
<geoCoordinate id="EE18506AE3BBFF2DFD74FEA1BBC0FE99" box="[662,764,260,284]" direction="south" orientation="latitude" pageId="188" pageNumber="189" precision="925" value="-13.65">13° 39' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE3BBFF2DFCEAFEA1BA4CFE99" box="[776,880,260,285]" direction="east" orientation="longitude" pageId="188" pageNumber="189" precision="925" value="48.75">48° 45' E</geoCoordinate>
,
<collectingDate id="EFD6E985E3BBFF2DFC9EFEA1BD09FE99" box="[892,1077,260,285]" pageId="188" pageNumber="189" value="2004-11-01" valueMax="2004-12-29" valueMin="2004-11-01">
<date id="FF92106DE3BBFF2DFC9EFEA1BD09FE99" box="[892,1077,260,285]" pageId="188" pageNumber="189" value="2004-11-01" valueMax="2004-12-29" valueMin="2004-11-01">1 xi–29 xii 2004</date>
</collectingDate>
,
<collectorName id="26D9537BE3BBFF2DFBA2FEA0BD9BFE98" box="[1088,1191,260,285]" pageId="188" pageNumber="189">M. Balke</collectorName>
(MD 016) (on female &quot;
<collectionCode id="ED3DAE68E3BBFF2DFF7DFE8DB9CAFEC5" box="[159,246,296,320]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="188" pageNumber="189">MNCN</collectionCode>
<specimenCode id="DB8A9ED6E3BBFF2DFF1FFE82B879FEC5" box="[253,325,295,320]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="188" pageNumber="189">AI546</specimenCode>
<collectionCode id="ED3DAE68E3BBFF2DFEAEFE8DB8B5FEC5" box="[332,393,296,320]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="188" pageNumber="189">DNA</collectionCode>
specimen&quot;).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF536625E3BBFF2DFF75F9E4BADBF9D2" box="[151,999,1601,1623]" pageId="188" pageNumber="189">
<paragraph id="8B9336ADE3BBFF2DFF75F9E4BADBF9D2" blockId="188.[151,999,1601,1623]" box="[151,999,1601,1623]" pageId="188" pageNumber="189">
<emphasis id="B958EABFE3BBFF2DFF75F9E4B810F9D3" bold="true" box="[151,300,1601,1623]" pageId="188" pageNumber="189">FIGURE 180.</emphasis>
<taxonomicName id="4C2C4D2EE3BBFF2DFED0F9E4BB2BF9D2" authorityName="Perkins" authorityYear="2017" box="[306,535,1601,1623]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="procarina">
<emphasis id="B958EABFE3BBFF2DFED0F9E4BB2BF9D2" box="[306,535,1601,1623]" italics="true" pageId="188" pageNumber="189">Madagaster procarina</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<subSubSection id="C3366526E3BBFF2DFF25F9DBBB83F8A2" pageId="188" pageNumber="189" type="diagnosis">
<paragraph id="8B9336ADE3BBFF2DFF25F9DBBB83F8A2" blockId="188.[151,1436,1662,2011]" pageId="188" pageNumber="189">
<emphasis id="B958EABFE3BBFF2DFF25F9DBB8EEF912" bold="true" box="[199,466,1662,1687]" pageId="188" pageNumber="189">Differential Diagnosis.</emphasis>
Similar in habitus to
<taxonomicName id="4C2C4D2EE3BBFF2DFD20F9DABA7CF912" authorityName="Perkins" authorityYear="2017" box="[706,832,1662,1687]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="B958EABFE3BBFF2DFD20F9DABA7CF912" box="[706,832,1662,1687]" italics="true" pageId="188" pageNumber="189">M. barbata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D2EE3BBFF2DFC97F9DABD3DF912" authorityName="Perkins" authorityYear="2017" box="[885,1025,1662,1687]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B958EABFE3BBFF2DFC97F9DABD3DF912" box="[885,1025,1662,1687]" italics="true" pageId="188" pageNumber="189">M. bergsteni</emphasis>
</taxonomicName>
(
<figureCitation id="13172A28E3BBFF2DFBF2F9DBBD47F912" box="[1040,1147,1662,1687]" captionStart="FIGURE 179" captionStartId="187.[151,250,1464,1486]" captionText="FIGURE 179. Madagaster bergsteni, holotype, habitus dorsal, lateral, and ventral." pageId="188" pageNumber="189">Figs. 179</figureCitation>
,
<figureCitation id="13172A28E3BBFF2DFB65F9DABD8FF912" box="[1159,1203,1663,1687]" captionStart="FIGURE 180" captionStartId="188.[151,250,1601,1623]" captionText="FIGURE 180. Madagaster procarina, holotype, habitus dorsal, lateral, and ventral." pageId="188" pageNumber="189">180</figureCitation>
,
<figureCitation id="13172A28E3BBFF2DFB5CF9DABDD0F912" box="[1214,1260,1663,1687]" captionStart="FIGURE 183" captionStartId="191.[151,250,1440,1462]" captionTargetBox="[340,1247,195,1408]" captionTargetId="figure@191.[340,1247,195,1408]" captionTargetPageId="191" captionText="FIGURE 183. Madagaster barbata, holotype, frontal view, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041124/files/figure.png" pageId="188" pageNumber="189">183</figureCitation>
). Differentiated from both species by the low midlongitudinal carina on the prosternum, the straight meso- and metatibiae, and the densely setose medial margin of the metafemur. The very complex male genitalia of the three species share a hookshaped process on the ventral surface of the main piece, but the parameres and processes of the distal part of the main piece differ markedly (
<figureCitation id="13172A28E3BBFF2DFE34F8ABBB45F8A2" box="[470,633,1806,1831]" captionStart="FIGURES 181 – 182" captionStartId="189.[151,264,1764,1787]" captionText="FIGURES 181 – 182. Madagaster, aedeagi of holotypes." pageId="188" pageNumber="189">Figs. 181, 182</figureCitation>
,
<figureCitation id="13172A28E3BBFF2DFD67F8ABBB8FF8A2" box="[645,691,1806,1831]" captionStart="FIGURES 185 – 186" captionStartId="193.[151,264,1869,1892]" captionText="FIGURES 185 – 186. Madagaster, aedeagi of holotypes." pageId="188" pageNumber="189">185</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3BBFF2FFF25F897BC2CFE28" lastPageId="190" lastPageNumber="191" pageId="188" pageNumber="189" type="description">
<paragraph id="8B9336ADE3BBFF2DFF25F897BA37F8EA" blockId="188.[151,1436,1662,2011]" pageId="188" pageNumber="189">
<emphasis id="B958EABFE3BBFF2DFF25F897B86DF8CE" bold="true" box="[199,337,1842,1867]" pageId="188" pageNumber="189">Description</emphasis>
. Size:
<typeStatus id="5497880FE3BBFF2DFE41F897BB39F8CE" box="[419,517,1842,1867]" pageId="188" pageNumber="189" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.23/1.02; head width 0.60; pronotum 0.55/0.86, PA 0.69, PB 0.68; elytra 1.38/1.02.
</paragraph>
<paragraph id="8B9336ADE3BBFF2DFF75F8DEBADBF817" blockId="188.[151,1436,1662,2011]" box="[151,999,1914,1939]" pageId="188" pageNumber="189">Dorsum and venter black, legs dark brown, ocelli light brown to testaceous.</paragraph>
<paragraph id="8B9336ADE3BBFF2CFF25F83BBB2DFE98" blockId="188.[151,1436,1662,2011]" lastBlockId="189.[151,1436,151,357]" lastPageId="189" lastPageNumber="190" pageId="188" pageNumber="189">Entire dorsum dull, with very small, closely spaced tubercles, each with a short recumbent seta. Labrum bilobed, anterior margin slightly reflexed, sparsely granulate and with sparse small setae except with longer setae along anterior margin. Labrum originating at anterior, slightly overhanging, margin of clypeus. Labroclypeal suture moderately deep. Short, deep sulcus in front of each ocellus. Frons with deep concavity between ocelli. Maxillary palpi (male) ca. as along as antenna, with last two palpomeres slightly enlarged. Antenna with penultimate antennomere larger than ultimate.</paragraph>
<paragraph id="8B9336ADE3BAFF2CFF25FE8DBDD0FEE0" blockId="189.[151,1436,151,357]" pageId="189" pageNumber="190">Pronotum with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; transverse fovea along base; lateral margin rounded.</paragraph>
<caption id="DF536625E3BAFF2CFF75F941BBEEF97E" box="[151,722,1764,1787]" pageId="189" pageNumber="190">
<paragraph id="8B9336ADE3BAFF2CFF75F941BBEEF97E" blockId="189.[151,722,1764,1787]" box="[151,722,1764,1787]" pageId="189" pageNumber="190">
<emphasis id="B958EABFE3BAFF2CFF75F941B851F97F" bold="true" box="[151,365,1764,1787]" pageId="189" pageNumber="190">FIGURES 181–182.</emphasis>
<emphasis id="B958EABFE3BAFF2CFE91F940B8D0F97E" box="[371,492,1765,1787]" italics="true" pageId="189" pageNumber="190">Madagaster</emphasis>
, aedeagi of holotypes.
</paragraph>
</caption>
<paragraph id="8B9336ADE3BAFF2CFF25F887BBC3F807" blockId="189.[151,1437,1826,2031]" pageId="189" pageNumber="190">Elytral suture costate in posterior. Each elytron with four non-interrupted costae; first costa (from suture) slightly curved in ca. middle ½ of length. Each costa with unilinear row of granules. Areas between costae with four irregular rows of granules. Elytral apices truncate.</paragraph>
<paragraph id="8B9336ADE3BAFF2CFF25F82ABA8CF86A" blockId="189.[151,1437,1826,2031]" pageId="189" pageNumber="190">Mentum and postmentum densely micropunctulate, sparsely pubescent; Genae micropunctulate, dull, concave behind cardo, with very strong, high posterior ridge, obliquely oriented on each side of midline. Subocular groove for antenna moderately deep and wide. Postocular margin not carinate.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FF3DBB21FEC5" blockId="190.[151,1437,151,464]" pageId="190" pageNumber="191">
Prosternum with midlongitudinal carina. Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short and very dense vestiture; midlongitudinal depression moderately deep (
<figureCitation id="13172A28E3B9FF2FFF7DFF7AB839FF7D" box="[159,261,223,248]" captionStart="FIGURE 180" captionStartId="188.[151,250,1601,1623]" captionText="FIGURE 180. Madagaster procarina, holotype, habitus dorsal, lateral, and ventral." pageId="190" pageNumber="191">Fig. 180</figureCitation>
). First four abdominal ventrites and anterior margin of fifth with very dense, recumbent pubescence (plastron?); setae sparser in midline than laterally. Last ventrite (male only) oval, and with dense pubescence, setae longer at margin forming oval rim.
</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FEE8BBA9FE0D" blockId="190.[151,1437,151,464]" pageId="190" pageNumber="191">Pro- and metatibiae slightly arcuate, toward midline. All tibiae with brush of setae apically. Metafemur with brush of setae along lower margin in basal ½.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FE31BC2CFE28" blockId="190.[151,1437,151,464]" box="[199,1296,404,429]" pageId="190" pageNumber="191">
Female length (ca.
<quantity id="4CD49B48E3B9FF2FFE7CFE31BB7EFE29" box="[414,578,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" metricValueMax="2.39" metricValueMin="2.09" pageId="190" pageNumber="191" unit="mm" value="2.24" valueMax="2.39" valueMin="2.09">2.09–2.39 mm</quantity>
) sometimes greater than males, but form narrower (less robust).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B9FF2FFF25FE12BD6FFE4A" box="[199,1107,439,464]" pageId="190" pageNumber="191" type="etymology">
<paragraph id="8B9336ADE3B9FF2FFF25FE12BD6FFE4A" blockId="190.[151,1437,151,464]" box="[199,1107,439,464]" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFF25FE12B872FE55" bold="true" box="[199,334,439,464]" pageId="190" pageNumber="191">Etymology.</emphasis>
Named in reference to the midlongitudinal carina of the prosternum.
</paragraph>
</subSubSection>
</treatment>
</document>
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