175 lines
19 KiB
XML
175 lines
19 KiB
XML
<document id="C9EA3567D1AD8670DB17FFBEB5624291" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE350FFC9FF75FF3DBAB8FC68" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena tripartita Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="89" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="88" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="E6762D2F1634748CF7AB1559C5290EFE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="723ABB4600DAC134A6FBF99FA10BD8D5">
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<mods:title id="C3E209B25E8FD5CABEA285F6121EFA9C">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="5C2C98D1FFEF7C8B7CC5CE53B5ED7A88" type="personal">
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<mods:role id="C0F5520F73C407EBB4D1A94DD6D80AD7">
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<mods:roleTerm id="5FB4F359ACFC8D912C44B6ED772E4144">Author</mods:roleTerm>
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<mods:namePart id="0A316998A6C916C2C67F660ADAE4FA0C">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="5AA97DA0FDD357493CF3D959C3FF3E26">text</mods:typeOfResource>
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<mods:titleInfo id="32B69FE2367F5A4F43EB1AC88C86322E">
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<mods:title id="C950C59E78E26A252D3B8A59333EE1A9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="75ABE86DFECC5136DA79AFF6D2086A9A">
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<mods:date id="90420E0BF6CCEC0C602BECB634CFC54D">2017</mods:date>
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<mods:detail id="F93F11C91FCE6EA9A65588454B59A636" type="pubDate">
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<mods:number id="ED2E1D1D0D82F6D527128B833F54F23F">2017-11-03</mods:number>
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<mods:detail id="1B222B7CF26D92302484F0A746942898" type="volume">
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<mods:number id="3DB57C3994D3FA590BA93D34B7EA9EF1">4342</mods:number>
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</mods:detail>
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<mods:detail id="8286B25BC7F488B327D80B62A110605A" type="issue">
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<mods:number id="1E0FA7ABBF46CA89C0E061D09C8F9A1B">1</mods:number>
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<mods:start id="DE7FCAACEE9674D5DEBFBC4D006326C2">1</mods:start>
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<mods:end id="88D9E4856C48821F5EC0D3DC290BD35D">264</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="84B2E393D0A98DD2845DA14554AB1FAE">journal article</mods:classification>
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<mods:identifier id="1AAB651584DD0E15BE86CE1F80E14815" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="19F87D08EE0F6C12B10077FE37972756" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="74810B0DEB24220D7253250C28730631" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="476F6952130005296563CAC1C1CCE93E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BA4831F98F4ACCF25908B0CB322945D3" type="Zenodo-Dep">1041066</mods:identifier>
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<mods:identifier id="81022CBE3BE661F0DE1D638D8BB2B1CA" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
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</mods:mods>
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<treatment id="038587BBE350FFC9FF75FF3DBAB8FC68" ID-DOI="http://doi.org/10.5281/zenodo.5323566" ID-GBIF-Taxon="137233474" ID-Zenodo-Dep="5323566" LSID="urn:lsid:plazi:treatment:038587BBE350FFC9FF75FF3DBAB8FC68" httpUri="http://treatment.plazi.org/id/038587BBE350FFC9FF75FF3DBAB8FC68" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
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<subSubSection id="C3366526E350FFC6FF75FF3DBBB4FF56" pageId="87" pageNumber="88" type="nomenclature">
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<paragraph id="8B9336ADE350FFC6FF75FF3DBB1CFF37" blockId="87.[151,648,152,211]" box="[151,544,152,178]" pageId="87" pageNumber="88">
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<heading id="D0DB81C1E350FFC6FF75FF3DBB1CFF37" bold="true" box="[151,544,152,178]" fontSize="11" level="1" pageId="87" pageNumber="88" reason="1">
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<emphasis id="B958EABFE350FFC6FF75FF3DBB1CFF37" bold="true" box="[151,544,152,178]" pageId="87" pageNumber="88">
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<taxonomicName id="4C2C4D2EE350FFC6FF75FF3DB8BDFF37" ID-CoL="3N3MG" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,385,152,178]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tripartita" status="sp. nov.">
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<emphasis id="B958EABFE350FFC6FF75FF3DB8BDFF37" bold="true" box="[151,385,152,178]" italics="true" pageId="87" pageNumber="88">Hydraena tripartita</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26B57C4E350FFC6FE72FF3DBB1CFF37" box="[400,544,152,178]" pageId="87" pageNumber="88" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336ADE350FFC6FF75FF1FBBB4FF56" blockId="87.[151,648,152,211]" box="[151,648,186,211]" pageId="87" pageNumber="88">
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<figureCitation id="13172A28E350FFC6FF75FF1FB9CDFF57" box="[151,241,186,211]" captionStart="FIGURE 77" captionStartId="87.[151,250,1847,1869]" captionText="FIGURE 77. Hydraena tripartita, holotype, habitus dorsal, lateral, and ventral." pageId="87" pageNumber="88">Figs. 77</figureCitation>
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(habitus), 79 (aedeagus), 264 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E350FFC6FF75FEA7B813FE2E" pageId="87" pageNumber="88" type="materials_examined">
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<paragraph id="8B9336ADE350FFC6FF75FEA7B813FE2E" blockId="87.[151,1437,258,607]" pageId="87" pageNumber="88">
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<materialsCitation id="3B443CF0E350FFC6FF75FEA7BDAAFEBA" ID-GBIF-Occurrence="1697157584" collectingDate="2001-08-19" collectionCode="NMW" collectorName="Gerecke & Goldschmidt" elevation="950" latitude="-23.416666" location="Ranomena" longLatPrecision="1256" longitude="47.266666" pageId="87" pageNumber="88" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
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<emphasis id="B958EABFE350FFC6FF75FEA7B8FEFE9E" bold="true" box="[151,450,258,283]" pageId="87" pageNumber="88">
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<typeStatus id="5497880FE350FFC6FF75FEA7B9EEFE9E" box="[151,210,258,283]" pageId="87" pageNumber="88">Type</typeStatus>
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Material.
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<typeStatus id="5497880FE350FFC6FEB7FEA7B8FEFE9E" box="[341,450,258,283]" pageId="87" pageNumber="88" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingRegion id="49E8F84FE350FFC6FDCAFEA7BBFAFE9E" box="[552,710,258,283]" country="Madagascar" name="Fianarantsoa" pageId="87" pageNumber="88">
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||
<emphasis id="B958EABFE350FFC6FDCAFEA7BBFAFE9E" bold="true" box="[552,710,258,283]" pageId="87" pageNumber="88">Fianarantsoa</emphasis>
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||
</collectingRegion>
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,
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<location id="8EF36076E350FFC6FD35FEA6BA6EFE9F" LSID="urn:lsid:plazi:treatment:038587BBE350FFC9FF75FF3DBAB8FC68:8EF36076E350FFC6FD35FEA6BA6EFE9F" box="[727,850,259,282]" latitude="-23.416666" longLatPrecision="1256" longitude="47.266666" name="Ranomena" pageId="87" pageNumber="88" stateProvince="Fianarantsoa">Ranomena</location>
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, river NW from the 1.07 km-railway-tunnel, pool, elev.
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<elevation id="0001D19EE350FFC6FF30FE83B827FEBB" box="[210,283,294,319]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="87" pageNumber="88" unit="m" value="950.0">
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<quantity id="4CD49B48E350FFC6FF30FE83B827FEBB" box="[210,283,294,319]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="87" pageNumber="88" unit="m" value="950.0">950 m</quantity>
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</elevation>
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,
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<geoCoordinate id="EE18506AE350FFC6FEC5FE82B8B3FEBA" box="[295,399,294,319]" direction="south" orientation="latitude" pageId="87" pageNumber="88" precision="925" value="-23.416666">23° 25' S</geoCoordinate>
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,
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<geoCoordinate id="EE18506AE350FFC6FE7EFE82BB3BFEBB" box="[412,519,294,319]" direction="east" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="47.266666">47° 16' E</geoCoordinate>
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,
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<collectingDate id="EFD6E985E350FFC6FDF0FE82BBA1FEBA" box="[530,669,294,319]" pageId="87" pageNumber="88" value="2001-08-19">
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<date id="FF92106DE350FFC6FDF0FE82BBA1FEBA" box="[530,669,294,319]" pageId="87" pageNumber="88" value="2001-08-19">19 viii 2001</date>
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</collectingDate>
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,
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<collectorName id="26D9537BE350FFC6FD4AFE83BA39FEBA" box="[680,773,294,319]" pageId="87" pageNumber="88">Gerecke</collectorName>
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&
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<collectorName id="26D9537BE350FFC6FCC8FE83BA87FEBA" box="[810,955,294,319]" pageId="87" pageNumber="88">Goldschmidt</collectorName>
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(MD 34c) (
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<collectionCode id="ED3DAE68E350FFC6FBA3FE82BDB3FEBA" box="[1089,1167,295,319]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="87" pageNumber="88">NMW</collectionCode>
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)
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</materialsCitation>
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.
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<emphasis id="B958EABFE350FFC6FB43FE83BC25FEBA" bold="true" box="[1185,1305,294,319]" pageId="87" pageNumber="88">Paratypes</emphasis>
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(10): Same data as holotype (1 NMW).
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<materialsCitation id="3B443CF0E350FFC6FE15FEEFB813FE2E" ID-GBIF-Occurrence="1697157581" collectingDate="2014-11-02" collectionCode="MCZ" collectorName="Ranomafana NP & P. D. Perkins" elevation="1130" latitude="-21.2395" location="Sahamalaotra" longLatPrecision="7" longitude="47.3947" municipality="Matsiara Ambony" pageId="87" pageNumber="88" specimenCode="MAD14- 04" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
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||
<collectingRegion id="49E8F84FE350FFC6FE15FEEFBBAAFEE6" box="[503,662,330,355]" country="Madagascar" name="Fianarantsoa" pageId="87" pageNumber="88">
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||
<emphasis id="B958EABFE350FFC6FE15FEEFBBAAFEE6" bold="true" box="[503,662,330,355]" pageId="87" pageNumber="88">Fianarantsoa</emphasis>
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||
</collectingRegion>
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,
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<collectingMunicipality id="6BF7ACD7E350FFC6FD4EFEEFBA43FEE6" box="[684,895,330,355]" pageId="87" pageNumber="88">Matsiara Ambony</collectingMunicipality>
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,
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<collectorName id="26D9537BE350FFC6FC70FEEFBD6EFEE7" box="[914,1106,330,355]" pageId="87" pageNumber="88">Ranomafana NP</collectorName>
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, next to
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||
<location id="8EF36076E350FFC6FB2AFEEFBC5CFEE6" LSID="urn:lsid:plazi:treatment:038587BBE350FFC9FF75FF3DBAB8FC68:8EF36076E350FFC6FB2AFEEFBC5CFEE6" box="[1224,1376,330,355]" latitude="-21.2395" longLatPrecision="7" longitude="47.3947" municipality="Matsiara Ambony" name="Sahamalaotra" pageId="87" pageNumber="88" stateProvince="Fianarantsoa">Sahamalaotra</location>
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trail entrance; forest stream with sandy bottom, elev.
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<elevation id="0001D19EE350FFC6FD23FECABA24FE03" box="[705,792,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="87" pageNumber="88" unit="m" value="1130.0">
|
||
<quantity id="4CD49B48E350FFC6FD23FECABA24FE03" box="[705,792,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="87" pageNumber="88" unit="m" value="1130.0">1130 m</quantity>
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</elevation>
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,
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<geoCoordinate id="EE18506AE350FFC6FCC7FECBBAACFE02" box="[805,912,366,391]" direction="south" orientation="latitude" pageId="87" pageNumber="88" precision="5" value="-21.2395">21.2395S</geoCoordinate>
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<geoCoordinate id="EE18506AE350FFC6FC7AFECABD3BFE02" box="[920,1031,367,391]" direction="east" orientation="longitude" pageId="87" pageNumber="88" precision="5" value="47.3947">47.3947E</geoCoordinate>
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,
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<collectingDate id="EFD6E985E350FFC6FBF6FECABDB8FE02" box="[1044,1156,366,391]" pageId="87" pageNumber="88" value="2014-11-02">
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<date id="FF92106DE350FFC6FBF6FECABDB8FE02" box="[1044,1156,366,391]" pageId="87" pageNumber="88" value="2014-11-02">2 xi 2014</date>
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</collectingDate>
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,
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<collectorName id="26D9537BE350FFC6FB72FECABC18FE02" box="[1168,1316,366,391]" pageId="87" pageNumber="88">P. D. Perkins</collectorName>
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(
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<specimenCode id="DB8A9ED6E350FFC6FAD6FECAB988FE2E" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="87" pageNumber="88">MAD14- 04</specimenCode>
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) (9
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<collectionCode id="ED3DAE68E350FFC6FF03FE36B81FFE2E" box="[225,291,403,427]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="87" pageNumber="88">MCZ</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3366526E350FFC6FF25FE13B9DFFD92" pageId="87" pageNumber="88" type="diagnosis">
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<paragraph id="8B9336ADE350FFC6FF25FE13B9DFFD92" blockId="87.[151,1437,258,607]" pageId="87" pageNumber="88">
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<emphasis id="B958EABFE350FFC6FF25FE13B8EEFE4A" bold="true" box="[199,466,438,463]" pageId="87" pageNumber="88">Differential Diagnosis.</emphasis>
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Differentiated from other members of the group by the comparatively short elytra that have apices conjointly sharply rounded. The aedeagus, which has three distal processes, cannot be confused with others.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E350FFC9FF25FD87BB17FCD8" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
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<paragraph id="8B9336ADE350FFC6FF25FD87BA37FDDA" blockId="87.[151,1437,258,607]" pageId="87" pageNumber="88">
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<emphasis id="B958EABFE350FFC6FF25FD87B86DFDBE" bold="true" box="[199,337,546,571]" pageId="87" pageNumber="88">Description</emphasis>
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. Size:
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<typeStatus id="5497880FE350FFC6FE41FD87BB39FDBE" box="[419,517,546,571]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.72/0.73; head width 0.41; pronotum 0.39/0.55, PA 0.44, PB 0.45; elytra 0.94/0.73.
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</paragraph>
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<caption id="DF536625E350FFC6FF75F892BA82F8C8" box="[151,958,1847,1869]" pageId="87" pageNumber="88">
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<paragraph id="8B9336ADE350FFC6FF75F892BA82F8C8" blockId="87.[151,958,1847,1869]" box="[151,958,1847,1869]" pageId="87" pageNumber="88">
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<emphasis id="B958EABFE350FFC6FF75F892B823F8C9" bold="true" box="[151,287,1847,1869]" pageId="87" pageNumber="88">FIGURE 77.</emphasis>
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<taxonomicName id="4C2C4D2EE350FFC6FEC7F892B8D2F8C8" authorityName="Perkins" authorityYear="2017" box="[293,494,1847,1869]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tripartita">
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<emphasis id="B958EABFE350FFC6FEC7F892B8D2F8C8" box="[293,494,1847,1869]" italics="true" pageId="87" pageNumber="88">Hydraena tripartita</emphasis>
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</taxonomicName>
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, holotype, habitus dorsal, lateral, and ventral.
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</paragraph>
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</caption>
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<paragraph id="8B9336ADE350FFC6FF25F8D0BD13F808" blockId="87.[151,1437,1908,2005]" box="[199,1071,1908,1933]" pageId="87" pageNumber="88">Dorsum dark brown to piceous; legs dark brown; maxillary palpi light brown.</paragraph>
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<paragraph id="8B9336ADE350FFC9FF25F83DBD57FE98" blockId="87.[151,1437,1908,2005]" lastBlockId="88.[151,1437,151,1005]" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">Head with clypeus moderately finely and moderately sparsely punctate, lateral margins microreticulate; frons coarsely densely punctate near eyes, interstices narrow walls, weakly shining, disc less coarsely and less densely punctate, interstices 1–2xpd, shining. Pronotum moderately coarsely and moderately densely punctate, punctures in anterior and posterior larger than those on disc; interstices 1–2xpd, shining. Mentum moderately densely, moderately coarsely punctate, interstices shining; postmentum densely coarsely punctate, subrugose. Genae raised, non-punctate reliefs shining, posterior ridge well delimited by coarsely punctate sulcus.</paragraph>
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<paragraph id="8B9336ADE35FFFC9FF25FE8DBDD8FEE0" blockId="88.[151,1437,151,1005]" pageId="88" pageNumber="89">Pronotum sinuate laterally, distinctly narrowed basally, almost subcordate; anterior margin straight behind eyes, emarginate behind frons; PF1 and PF2 very shallow or absent; PF3 and PF4 deep, confluent.</paragraph>
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<paragraph id="8B9336ADE35FFFC9FF25FECABDCFFE70" blockId="88.[151,1437,151,1005]" pageId="88" pageNumber="89">Elytra arcuate laterally; posterior declivity steep, summit near midlength, elytra appearing slightly humped; elytra transversely very convex, especially at posterior 3/5; lateral explanate margins wide; serial punctures round, distinctly larger than punctures on pronotal disc, interstices 0.5–1xpd, shining. Apices in dorsal aspect conjointly sharply rounded, almost acuminate, in posterior aspect margins do not form angle with one another.</paragraph>
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<paragraph id="8B9336ADE35FFFC9FF25FE5ABB17FCD8" blockId="88.[151,1437,151,1005]" pageId="88" pageNumber="89">
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Ratios of P2 width and plaque shape (P2/w/l/s [posteriorly]) ca. 1/0.2/1.5/1. P1 slightly narrower than P2; median carina very weakly sinuate in profile. P2 l/w ca. 2/1, sides converging posteriorly, apex blunt and on same level as anteromedian carina of metaventrite. Plaques very narrow, strongly carinate lines, converging anteriorly and joining anteromedian carina, together forming inverted Y-shape, separated basally by ca.
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<date id="FF92106DE35FFFC9FB38FDC9BDC9FD01" box="[1242,1269,620,644]" pageId="88" pageNumber="89">4x</date>
|
||
plaque width, located at sides of deep median depression; area lateral to plaques concave. Metaventrite anteromedian carina strong, ca. same length as plaques; long longitudinal and slightly arcuate ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca. 1.5x P2. All tibiae straight, slender. Mesotibia medial margin near midlength with 2–3 very short sharp spines, margin shallowly notched between spines. Metafemur slender, arcuate on lateral margin, very weakly sinuate on medial margin. Abdominal apex with small apicomedian notch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366526E35FFFC9FF25FCC2B8FDFC20" pageId="88" pageNumber="89" type="etymology">
|
||
<paragraph id="8B9336ADE35FFFC9FF25FCC2B8FDFC20" blockId="88.[151,1437,151,1005]" pageId="88" pageNumber="89">
|
||
<emphasis id="B958EABFE35FFFC9FF25FCC2B872FC05" bold="true" box="[199,334,871,896]" pageId="88" pageNumber="89">Etymology.</emphasis>
|
||
Named in reference to the three processes of the aedeagus, and the three parts of the metaventral inverted Y-shaped carinae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3366526E35FFFC9FF25FC0ABAB8FC68" pageId="88" pageNumber="89" type="discussion">
|
||
<paragraph id="8B9336ADE35FFFC9FF25FC0ABAB8FC68" blockId="88.[151,1437,151,1005]" pageId="88" pageNumber="89">
|
||
<emphasis id="B958EABFE35FFFC9FF25FC0AB807FC4D" bold="true" box="[199,315,943,968]" pageId="88" pageNumber="89">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="5497880FE35FFFC9FE94FC0AB8E4FC4D" box="[374,472,943,968]" pageId="88" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
is solid black, but teneral specimens collected under decaying leaves on a cascade at Ranomafana N.P. have a distinct pronotal fascia, and lighter elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |