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<document id="7F184493CCE60CC9237D18C3B30C14B1" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFF784158CFBF9F5DF82FDC1" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes nigripes Kluge, 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="114" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="107" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="A84C156D5E086E42EAFEFAB18267894F">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:namePart id="AAC7540948FA890A057BAE52FB640003">Kluge, Nikita J.</mods:namePart>
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<mods:date id="102BAE8EEE9CBB6FE7B3788BF0B3B56E">2020</mods:date>
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<treatment id="039787A6FFF784158CFBF9F5DF82FDC1" ID-DOI="http://doi.org/10.5281/zenodo.3811806" ID-GBIF-Taxon="163420769" ID-Zenodo-Dep="3811806" LSID="urn:lsid:plazi:treatment:039787A6FFF784158CFBF9F5DF82FDC1" httpUri="http://treatment.plazi.org/id/039787A6FFF784158CFBF9F5DF82FDC1" lastPageId="113" lastPageNumber="114" pageId="106" pageNumber="107">
<subSubSection id="C324653BFFF7840E8CFBF9F5D93EF98F" box="[151,512,1554,1581]" pageId="106" pageNumber="107" type="nomenclature">
<paragraph id="8B8136B0FFF7840E8CFBF9F5D93EF98F" blockId="106.[151,512,1554,1617]" box="[151,512,1554,1581]" pageId="106" pageNumber="107">
<heading id="D0C981DCFFF7840E8CFBF9F5D93EF98F" bold="true" box="[151,512,1554,1581]" fontSize="10" level="2" pageId="106" pageNumber="107" reason="2">
<emphasis id="B94AEAA2FFF7840E8CFBF9F5D93EF98F" bold="true" box="[151,512,1554,1581]" pageId="106" pageNumber="107">
15.
<taxonomicName id="4C3E4D33FFF7840E8CAFF9F4DA89F98F" box="[195,439,1554,1581]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFF7840E8CAFF9F4DA89F98F" bold="true" box="[195,439,1554,1581]" italics="true" pageId="106" pageNumber="107">Thraulodes nigripes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957D9FFF7840E8DD2F9F5D93EF98F" box="[446,512,1555,1581]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C324653BFFF7840E8CFBF9D1DA7AF9F3" box="[151,324,1590,1617]" pageId="106" pageNumber="107" type="description">
<paragraph id="8B8136B0FFF7840E8CFBF9D1DA7AF9F3" blockId="106.[151,512,1554,1617]" box="[151,324,1590,1617]" pageId="106" pageNumber="107">
(
<figureCitation id="13052A35FFF7840E8CF3F9D1DA05F9F3" box="[159,315,1590,1617]" captionStart-0="FIGURES 479490" captionStart-1="FIGURES 491497" captionStart-2="FIGURES 498507" captionStart-3="FIGURES 508509" captionStartId-0="108.[151,264,1614,1639]" captionStartId-1="109.[151,264,1764,1789]" captionStartId-2="111.[151,264,1685,1710]" captionStartId-3="112.[151,264,2005,2030]" captionTargetBox-0="[151,1436,174,1595]" captionTargetBox-1="[151,1436,174,1744]" captionTargetBox-2="[157,1420,225,1620]" captionTargetBox-3="[193,1395,166,1980]" captionTargetId-0="figure@108.[151,1436,174,1595]" captionTargetId-1="figure@109.[151,1436,174,1744]" captionTargetId-2="figure@111.[151,1436,212,1639]" captionTargetId-3="figure@112.[193,1395,166,1980]" captionTargetPageId-0="108" captionTargetPageId-1="109" captionTargetPageId-2="111" captionTargetPageId-3="112" captionText-0="FIGURES 479490. Thraulodes nigripes sp. n. 479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype)." captionText-1="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." captionText-2="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." captionText-3="FIGURES 508509. Thraulodes nigripes sp. n., eggs." figureDoi-0="http://doi.org/10.5281/zenodo.3737028" figureDoi-1="http://doi.org/10.5281/zenodo.3737030" figureDoi-2="http://doi.org/10.5281/zenodo.3737032" figureDoi-3="http://doi.org/10.5281/zenodo.3737034" httpUri-0="https://zenodo.org/record/3737028/files/figure.png" httpUri-1="https://zenodo.org/record/3737030/files/figure.png" httpUri-2="https://zenodo.org/record/3737032/files/figure.png" httpUri-3="https://zenodo.org/record/3737034/files/figure.png" pageId="106" pageNumber="107">Figs 479509</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFF7840E8CFBF99EDF61F930" box="[151,1119,1656,1683]" pageId="106" pageNumber="107" type="etymology">
<paragraph id="8B8136B0FFF7840E8CFBF99EDF61F930" blockId="106.[151,1437,1656,2043]" box="[151,1119,1656,1683]" pageId="106" pageNumber="107">
<emphasis id="B94AEAA2FFF7840E8CFBF99EDA20F930" bold="true" box="[151,286,1656,1682]" pageId="106" pageNumber="107">Etymology.</emphasis>
Allusion to uniformly dark brown, nearly black color of all legs of imago.
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFF7840E8CABF97DDA38F95C" pageId="106" pageNumber="107" type="materials_examined">
<paragraph id="8B8136B0FFF7840E8CABF97DDA38F95C" blockId="106.[151,1437,1656,2043]" pageId="106" pageNumber="107">
<emphasis id="B94AEAA2FFF7840E8CABF97DDA8AF914" bold="true" box="[199,436,1691,1718]" pageId="106" pageNumber="107">Material examined.</emphasis>
<materialsCitation id="3B563CEDFFF7840E8DD3F97AD9F5F978" ID-GBIF-Occurrence="2594509896" collectingDate="2006-02-21" collectingDateMax="2006-05-14" collectingDateMin="2006-02-21" collectorName="N. Kluge." country="Peru" latitude="-9.066667" location="Pampa Yurac" longLatPrecision="1300" longitude="-75.95" municipality="Aquaytia" pageId="106" pageNumber="107" specimenCount="1" specimenCount-male="1" stateProvince="Huanuco" typeStatus="holotype">
<emphasis id="B94AEAA2FFF7840E8DD3F97AD90BF914" bold="true" box="[447,565,1692,1718]" pageId="106" pageNumber="107">
<typeStatus id="54858812FFF7840E8DD3F97AD90EF914" box="[447,560,1692,1718]" pageId="106" pageNumber="107" type="holotype">Holotype</typeStatus>
:
</emphasis>
L/S
<specimenCount id="9D38FD39FFF7840E8E06F97AD940F914" box="[618,638,1692,1718]" pageId="106" pageNumber="107" type="male"></specimenCount>
{slide
<date id="FF801070FFF7840E8EBAF97AD86AF915" box="[726,852,1692,1719]" pageId="106" pageNumber="107" value="2006-05-14">
<collectingDate id="EFC4E998FFF7840E8EBAF97AD86AF915" box="[726,852,1692,1719]" pageId="106" pageNumber="107" value="2006-05-14">14.V.2006</collectingDate>
</date>
-1},
<collectingCountry id="F3297620FFF7840E8F13F97BD8F7F915" box="[895,969,1693,1719]" name="Peru" pageId="106" pageNumber="107">PERU</collectingCountry>
, Reg.
<collectingRegion id="49FAF852FFF7840E887BF97BDF41F915" box="[1047,1151,1693,1719]" country="Peru" name="Huanuco" pageId="106" pageNumber="107">Huanuco</collectingRegion>
,
<collectingMunicipality id="6BE5ACCAFFF7840E88E0F97ADFC8F914" box="[1164,1270,1692,1718]" pageId="106" pageNumber="107">Aquaytia</collectingMunicipality>
,
<location id="8EE1606BFFF7840E8969F97BDEA2F915" LSID="urn:lsid:plazi:treatment:039787A6FFF784158CFBF9F5DF82FDC1:8EE1606BFFF7840E8969F97BDEA2F915" box="[1285,1436,1693,1719]" country="Peru" latitude="-9.066667" longLatPrecision="1300" longitude="-75.95" municipality="Aquaytia" name="Pampa Yurac" pageId="106" pageNumber="107" stateProvince="Huanuco">Pampa Yurac</location>
(
<geoCoordinate id="EE0A5077FFF7840E8CCCF926DBCFF978" box="[160,241,1728,1754]" degrees="9" direction="south" minutes="04" orientation="latitude" pageId="106" pageNumber="107" precision="925" value="-9.066667">9°04S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFF7840E8C90F959DA58F978" box="[252,358,1727,1754]" degrees="75" direction="west" minutes="57" orientation="longitude" pageId="106" pageNumber="107" precision="925" value="-75.95">75°57W</geoCoordinate>
),
<date id="FF801070FFF7840E8D16F926D925F978" box="[378,539,1728,1754]" pageId="106" pageNumber="107" value="2006-02-21" valueMax="2006-02-24" valueMin="2006-02-21">
<collectingDate id="EFC4E998FFF7840E8D16F926D925F978" box="[378,539,1728,1754]" pageId="106" pageNumber="107" value="2006-02-21" valueMax="2006-02-24" valueMin="2006-02-21">2124.II.2006</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFF7840E8E31F927D9F5F978" box="[605,715,1728,1755]" pageId="106" pageNumber="107">N. Kluge.</collectorName>
</materialsCitation>
<materialsCitation id="3B563CEDFFF7840E8EBDF927DA38F95C" ID-GBIF-Occurrence="2594509897" collectingDate="2006-02-25" collectingDateMax="2006-05-14" collectingDateMin="2006-02-21" collectorName="N. Kluge." country="Peru" latitude="-9.066667" location="Pampa Yurac" longLatPrecision="1300" longitude="-75.95" municipality="Aquaytia" pageId="106" pageNumber="107" specimenCount="19" specimenCount-female="1" specimenCount-larva="18" specimenCount-male="1" stateProvince="Huanuco" typeStatus="paratype">
<typeStatus id="54858812FFF7840E8EBDF927D87CF979" box="[721,834,1729,1755]" pageId="106" pageNumber="107" type="paratype">Paratypes</typeStatus>
: the same locality and collector,
<date id="FF801070FFF7840E88C0F959DE1BF978" box="[1196,1317,1727,1754]" pageId="106" pageNumber="107" value="2006-02-25">
<collectingDate id="EFC4E998FFF7840E88C0F959DE1BF978" box="[1196,1317,1727,1754]" pageId="106" pageNumber="107" value="2006-02-25">25.II.2006</collectingDate>
</date>
: 1 L-S-I
<specimenCount id="9D38FD39FFF7840E89E5F926DEA9F978" box="[1417,1431,1728,1754]" pageId="106" pageNumber="107" type="female"></specimenCount>
,
<specimenCount id="9D38FD39FFF7840E8CFBF902DA3CF95C" box="[151,258,1764,1790]" pageId="106" pageNumber="107" type="larva">18 larvae</specimenCount>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFF784158CABF8E1D8E7FEBD" lastPageId="113" lastPageNumber="114" pageId="106" pageNumber="107" type="description">
<paragraph id="8B8136B0FFF7840E8CABF8E1DA5AF880" blockId="106.[151,1437,1656,2043]" box="[199,356,1799,1826]" pageId="106" pageNumber="107">
<emphasis id="B94AEAA2FFF7840E8CABF8E1DA5AF880" bold="true" box="[199,356,1799,1826]" pageId="106" pageNumber="107">Descriptions.</emphasis>
</paragraph>
<paragraph id="8B8136B0FFF7840E8CABF8CADEB7F82C" blockId="106.[151,1437,1656,2043]" pageId="106" pageNumber="107">
<emphasis id="B94AEAA2FFF7840E8CABF8CADA2CF8E4" bold="true" box="[199,274,1836,1862]" italics="true" pageId="106" pageNumber="107">Larva.</emphasis>
CUTICULAR COLORATION (
<figureCitation id="13052A35FFF7840E8EEDF8CAD81FF8E5" box="[641,801,1836,1863]" captionStart="FIGURES 479490" captionStartId="108.[151,264,1614,1639]" captionTargetBox="[151,1436,174,1595]" captionTargetId="figure@108.[151,1436,174,1595]" captionTargetPageId="108" captionText="FIGURES 479490. Thraulodes nigripes sp. n. 479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737028" httpUri="https://zenodo.org/record/3737028/files/figure.png" pageId="106" pageNumber="107">Figs 479486</figureCitation>
). Dorsal side of head, thorax and abdomen mostly brown with few blanks: pronotum bordered by light laterally; abdominal terga with blanks on lateral parts, area mediad of tergalii attachments nearly unicolor brown. Femora mostly brown, with blanks; tibiae and tarsi uniformly brown.
</paragraph>
<paragraph id="8B8136B0FFF7840E8CABF87EDE74F859" blockId="106.[151,1437,1656,2043]" pageId="106" pageNumber="107">HYPODERMAL COLORATION. In last instar larva, tibiae more or less darkened with brown or blackish, darker than femora (this coloration at first appears on fore leg, than on other legs). In larvae of various instars, abdomen with more or less expressed transverse brown or gray stripes on posterior margins of all or some terga.</paragraph>
<caption id="DF416638FFF6840F8CFBF97FDF1BF958" ID-DOI="http://doi.org/10.5281/zenodo.3737026" ID-Zenodo-Dep="3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="107" pageNumber="108" startId="107.[151,264,1689,1714]" targetBox="[154,1422,186,1652]" targetPageId="107">
<paragraph id="8B8136B0FFF6840F8CFBF97FDF1BF958" blockId="107.[151,1436,1689,1786]" pageId="107" pageNumber="108">
<emphasis id="B94AEAA2FFF6840F8CFBF97FDA50F910" bold="true" box="[151,366,1689,1714]" pageId="107" pageNumber="108">FIGURES 474478.</emphasis>
<taxonomicName id="4C3E4D33FFF6840F8D1AF97FD9A6F913" box="[374,664,1689,1713]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
<emphasis id="B94AEAA2FFF6840F8D1AF97FD9A6F913" box="[374,664,1689,1713]" italics="true" pageId="107" pageNumber="108">Thraulodes nigrabdominalis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFF6840F8EF3F97CD9E7F910" bold="true" box="[671,729,1690,1714]" pageId="107" pageNumber="108">
<taxonomicNameLabel id="A27957D9FFF6840F8EF3F97CD9E7F910" box="[671,729,1690,1714]" pageId="107" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474476, holotype).
</paragraph>
</caption>
<paragraph id="8B8136B0FFF6840F8CABF8AAD90FF870" blockId="107.[151,1437,1867,2002]" pageId="107" pageNumber="108">
SHAPE AND SETATION. Clypeus parallel-sided; labrum 1.21.4 times wider than clypeus (
<figureCitation id="13052A35FFF6840F8882F8AADEAEF8C4" box="[1262,1424,1867,1894]" captionStart="FIGURES 479490" captionStartId="108.[151,264,1614,1639]" captionTargetBox="[151,1436,174,1595]" captionTargetId="figure@108.[151,1436,174,1595]" captionTargetPageId="108" captionText="FIGURES 479490. Thraulodes nigripes sp. n. 479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737028" httpUri="https://zenodo.org/record/3737028/files/figure.png" pageId="107" pageNumber="108">Figs 481, 485</figureCitation>
). Labrum widest at 0.4 length from base; initial fore margin (turned ventrally) without median emargination, with all 5 denticles wide; anterior transverse setal row regular (as in
<figureCitation id="13052A35FFF6840F8F54F872D8B4F80C" box="[824,906,1940,1966]" captionStart="FIGURES 8693" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 8693. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 9093, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="107" pageNumber="108">Fig. 88</figureCitation>
), as wide as all 5 denticles. Maxilla with 1617 pectinate setae in apical-ventral row.
</paragraph>
<caption id="DF416638FFF184088CFBF9A8DA53F955" ID-DOI="http://doi.org/10.5281/zenodo.3737028" ID-Zenodo-Dep="3737028" httpUri="https://zenodo.org/record/3737028/files/figure.png" pageId="108" pageNumber="109" startId="108.[151,264,1614,1639]" targetBox="[151,1436,174,1595]" targetPageId="108">
<paragraph id="8B8136B0FFF184088CFBF9A8DA53F955" blockId="108.[151,1437,1614,1783]" pageId="108" pageNumber="109">
<emphasis id="B94AEAA2FFF184088CFBF9A8DA50F9C5" bold="true" box="[151,366,1614,1639]" pageId="108" pageNumber="109">FIGURES 479490.</emphasis>
<taxonomicName id="4C3E4D33FFF184088D1BF9A9D97BF9C5" box="[375,581,1615,1639]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFF184088D1BF9A9D97BF9C5" box="[375,581,1615,1639]" italics="true" pageId="108" pageNumber="109">Thraulodes nigripes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957D9FFF184088E21F9A9D9B9F9C5" box="[589,647,1615,1639]" pageId="108" pageNumber="109" rank="species">
<emphasis id="B94AEAA2FFF184088E21F9A9D9B9F9C5" bold="true" box="[589,647,1615,1639]" pageId="108" pageNumber="109">sp. n.</emphasis>
</taxonomicNameLabel>
479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype).
</paragraph>
</caption>
<paragraph id="8B8136B0FFF184088CABF8C0DFF0F8C1" blockId="108.[151,1436,1829,2036]" pageId="108" pageNumber="109">
Femora: Stout setae on anterior surface narrowed distally or parallel-sided, apically truncated (
<figureCitation id="13052A35FFF18408896BF8C3DE55F89D" box="[1287,1387,1829,1855]" captionStart="FIGURES 479490" captionStartId="108.[151,264,1614,1639]" captionTargetBox="[151,1436,174,1595]" captionTargetId="figure@108.[151,1436,174,1595]" captionTargetPageId="108" captionText="FIGURES 479490. Thraulodes nigripes sp. n. 479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737028" httpUri="https://zenodo.org/record/3737028/files/figure.png" pageId="108" pageNumber="109">Fig. 487</figureCitation>
). Irregular row of hairs near inner margin absent on fore femur, present on middle and hind femora.
</paragraph>
<paragraph id="8B8136B0FFF184088CABF888D9AEF856" blockId="108.[151,1436,1829,2036]" pageId="108" pageNumber="109">
Fore tibia (
<figureCitation id="13052A35FFF184088D2DF88BDA9AF825" box="[321,420,1901,1927]" captionStart="FIGURES 479490" captionStartId="108.[151,264,1614,1639]" captionTargetBox="[151,1436,174,1595]" captionTargetId="figure@108.[151,1436,174,1595]" captionTargetPageId="108" captionText="FIGURES 479490. Thraulodes nigripes sp. n. 479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737028" httpUri="https://zenodo.org/record/3737028/files/figure.png" pageId="108" pageNumber="109">Fig. 488</figureCitation>
): outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inner-anterior row of stout setae represented by few (23) blunt stout setae near tibia base; inner field of stout pointed setae dense (i.e. setae longer than distances between them), consists of pointed bipectinate and smooth setae, situated irregularly (about 23 setae in cross section).
</paragraph>
<caption id="DF416638FFF084098CFBF902DF57F8E7" ID-DOI="http://doi.org/10.5281/zenodo.3737030" ID-Zenodo-Dep="3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="109" pageNumber="110" startId="109.[151,264,1764,1789]" targetBox="[151,1436,174,1744]" targetPageId="109">
<paragraph id="8B8136B0FFF084098CFBF902DF57F8E7" blockId="109.[151,1437,1764,1861]" pageId="109" pageNumber="110">
<emphasis id="B94AEAA2FFF084098CFBF902DA51F95F" bold="true" box="[151,367,1764,1789]" pageId="109" pageNumber="110">FIGURES 491497.</emphasis>
<taxonomicName id="4C3E4D33FFF084098D1BF902D97BF95E" box="[375,581,1764,1788]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFF084098D1BF902D97BF95E" box="[375,581,1764,1788]" italics="true" pageId="109" pageNumber="110">Thraulodes nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFF084098E22F903D9B6F95F" bold="true" box="[590,648,1765,1789]" pageId="109" pageNumber="110">
<taxonomicNameLabel id="A27957D9FFF084098E22F903D9B6F95F" box="[590,648,1765,1789]" pageId="109" pageNumber="110" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype).
</paragraph>
</caption>
<paragraph id="8B8136B0FFF0840A8CABF871DA47FF75" blockId="109.[151,1437,1943,2041]" lastBlockId="110.[151,1437,153,1979]" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
Hind tibia (
<figureCitation id="13052A35FFF084098D29F871DADBF813" box="[325,485,1943,1969]" captionStart="FIGURES 479490" captionStartId="108.[151,264,1614,1639]" captionTargetBox="[151,1436,174,1595]" captionTargetId="figure@108.[151,1436,174,1595]" captionTargetPageId="108" captionText="FIGURES 479490. Thraulodes nigripes sp. n. 479483 and 484486, larval exuviae of two specimens at the same magnification: 479, abdominal terga; 480, left half of pronotum and mesonotum; 481 and 484486, parts of head with labrum and mandibles; 482483, fore and hind legs; 487, anterior side of hind femur; 488, anterior side of fore tibia; 489, hind tibia with focus on outer-posterior row of stout setae; 490, the same with focus on outer-anterior and inner-anterior rows of stout setae (484486, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737028" httpUri="https://zenodo.org/record/3737028/files/figure.png" pageId="109" pageNumber="110">Figs 489490</figureCitation>
): outer-anterior row of stout setae consists of two clearly different
<typeStatus id="54858812FFF0840988AEF871DFC3F813" box="[1218,1277,1943,1969]" pageId="109" pageNumber="110">types</typeStatus>
of stout setae: numerous shorter blunt ones and several long spoon-like ones; outer-posterior row of stout setae consists of blunt stout setae of various lengths; hairs located between these rows, numerous and form more than one row (besides row of hairs posteriad of outer-posterior row of stout setae); stout setae of inner-anterior row moderately long, parallelsided and truncated.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFF07DE23FF59" blockId="110.[151,1437,153,1979]" box="[199,1309,225,251]" pageId="110" pageNumber="111">Claws with 67 denticles on rigid portion, with or without minute denticles on articulatory portion.</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFEE3D82EFE29" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
Tergalii (
<figureCitation id="13052A35FFF3840A8D40FEE3DAF3FEBD" box="[300,461,260,287]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="110" pageNumber="111">Figs 503506</figureCitation>
): moderately wide; on both lamellae main trachea with branches directed toward costal and anal margins; dorsal lamella with costal margin most convex in proximal part and anal margin most convex in distal part, gradually narrowed toward apex, with slender apical filament; ventral lamella widest near base, gradually narrowed toward apex, with slender apical filament.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFE73D826FE55" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
Male genitalia in last larval instar (
<figureCitation id="13052A35FFF3840A8E2BFE73D994FE0D" box="[583,682,405,431]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="110" pageNumber="111">Fig. 498</figureCitation>
): protogonostyli long and separated one from another by deep emargination. Each protopenis lobe apically wide, with gonopore-bearing process sharply projected caudally and located near median margin; gonopore opened caudally-laterally.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFDE6DFA6FD6D" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
<emphasis id="B94AEAA2FFF3840A8CABFDE6DA7DFDB9" bold="true" box="[199,323,512,539]" italics="true" pageId="110" pageNumber="111">Subimago.</emphasis>
CUTICULAR COLORATION. Cuticle mostly light ocher, nearly colorless. Flagellum of antenna darker, light brown. Mesonotum partly brown, with dark brown antelateroparapsidal suture, colorless area on posterior scutal protuberance and colorless achromozones; medioscutal chromozone somewhat darker than submedioscutal chromozone (
<figureCitation id="13052A35FFF3840A8D1FFD8BDAE9FD25" box="[371,471,621,647]" captionStart="FIGURES 491497" captionStartId="109.[151,264,1764,1789]" captionTargetBox="[151,1436,174,1744]" captionTargetId="figure@109.[151,1436,174,1744]" captionTargetPageId="109" captionText="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="110" pageNumber="111">Fig. 497</figureCitation>
). Thoracic pleura with some sclerites light brownish. On all legs femur mostly light ocher, bordered by darker brown on outer and inner margins and apically; tibia and tarsus light brown. Cuticle of wings colorless, microtrichia brown. Abdominal terga very light brownish, sterna colorless.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFD3FDA1CFC99" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd4th tarsomeres of middle and hind legs (as in
<figureCitation id="13052A35FFF3840A8CD8FCC7DA2BFC99" box="[180,277,801,827]" captionStart="FIGURES 464473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="110" pageNumber="111">Fig. 473</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFCA2DA20FC21" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
<emphasis id="B94AEAA2FFF3840A8CABFCA2DA69FCFD" bold="true" box="[199,343,836,863]" italics="true" pageId="110" pageNumber="111">Male imago.</emphasis>
Unknown. Based on male subimago extracted from mature larva, following imaginal features can be reported.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFC6BDA19FC69" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">Head brown. Antenna with scapus and flagellum ocher, pedicellus dark brown. Dorsal eyes contiguous medially, grayish.</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFC33D8F0FBB1" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">Legs of all pairs with femur and tibia uniformly dark brown; tarsus of fore leg with each segment proximally dark brown, distally ocher; tarsus of middle and hind legs light brownish.</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFBFBD90EFA91" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
Abdominal hypodermal coloration (
<figureCitation id="13052A35FFF3840A8E09FBFBD9F2FB95" box="[613,716,1052,1079]" captionStart="FIGURES 491497" captionStartId="109.[151,264,1764,1789]" captionTargetBox="[151,1436,174,1744]" captionTargetId="figure@109.[151,1436,174,1744]" captionTargetPageId="109" captionText="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="110" pageNumber="111">Fig. 495</figureCitation>
): Terga with posterior margins narrowly bordered by grayishbrown; tergum I mostly brown; terga IIVI mostly ocher, each with pair of small, diffusive, transverse reddish-andbrown marks laterally; terga VIIX medially reddish-brown, laterally whitish; tergum VIII with pair of longitudinal lateral brown stripes, laterally narrowly bordered by brown; tergum IX with pair of smaller longitudinal lateral brown stripes. Abdominal sterna ocher with grayish nerve ganglia; sternum VIII with pair of longitudinal reddish stripes and brown maculae anteriad and posteriad of them; sternum IX tinged with reddish antero-laterally. Caudalii in proximal part dark brown with each 4th segment lighter, in middle part with alternated darker and lighter brown segments; all joinings darker brown.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFADBD8BAFA61" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
Genitalia (
<figureCitation id="13052A35FFF3840A8D2DFADBDA94FAF5" box="[321,426,1341,1367]" captionStart="FIGURES 491497" captionStartId="109.[151,264,1764,1789]" captionTargetBox="[151,1436,174,1744]" captionTargetId="figure@109.[151,1436,174,1744]" captionTargetPageId="109" captionText="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="110" pageNumber="111">Figs 496</figureCitation>
,
<figureCitation id="13052A35FFF3840A8DD5FADAD91EFAF5" box="[441,544,1340,1367]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="110" pageNumber="111">499501</figureCitation>
). Long segments of gonostyli (initial 1st+2nd segments) with intensive dark brown hypodermal coloration, both apical segments without it. Dorsal extension of styliger very long, parallelsided, truncated apically. Telopenes in form of spear-like rolls, long and straight; judging by projection laterad of spear-like roll (
<figureCitation id="13052A35FFF3840A8D2EFA4FDA9AFA61" box="[322,420,1448,1475]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="110" pageNumber="111">Fig. 501</figureCitation>
), imaginal penis with well-developed ears.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABFA2AD832F9F1" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
<emphasis id="B94AEAA2FFF3840A8CABFA2ADA53FA45" bold="true" box="[199,365,1484,1511]" italics="true" pageId="110" pageNumber="111">Female imago</emphasis>
(
<figureCitation id="13052A35FFF3840A8D17FA2BD925FA45" box="[379,539,1485,1511]" captionStart="FIGURES 491497" captionStartId="109.[151,264,1764,1789]" captionTargetBox="[151,1436,174,1744]" captionTargetId="figure@109.[151,1436,174,1744]" captionTargetPageId="109" captionText="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="110" pageNumber="111">Figs 493494</figureCitation>
). Head brown. Antenna with scapus and flagellum ocher, pedicellus dark brown. Pronotum brown with darker and lighter areas, propleura and prosternum ocher with brown. Mesonotum light brown; mesothoracic pleural sclerites, basisternum, furcasternal median impression and furcasternal protuberances brown, other areas ocher. Metathorax brown with ocher.
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABF9BBDF56F91D" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">Legs of all pairs with femur and tibia uniformly dark brown, tarsus light brownish. In specimen reared from larva, relation of fore, middle and hind femora to fore wing length 70:67:85:310; proportions femur/tibia/tarsomeres on fore leg 70:77:2:7:6:5:9; on middle leg 67:72:2:3:3:2:8; on hind leg 85:82:2:3:3:2:8.</paragraph>
<paragraph id="8B8136B0FFF3840A8CABF92FD8B2F8ED" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">
Fore wing with longitudinal and cross veins ocher-brownish; costal brace and adjacent parts of C and Sc dark brown. Costal cross veins proximad of bulla few, light ocher-brownish and very thin, so that visible only under high magnification (as in
<figureCitation id="13052A35FFF3840A8DD8F8F7D92BF889" box="[436,533,1809,1835]" captionStart="FIGURES 133142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133142. Thraulodes panamensis sp. n. 133, pterostigma; 134136, male imago in lateral in ventral and dorsal view; 137138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="110" pageNumber="111">Fig. 142</figureCitation>
). Pterostigmatic cross veins dense, oblique, non-branched. Hind wing with costal projection prominent; proximal part of hind wing brown (
<figureCitation id="13052A35FFF3840A8F71F8D3D8BEF8ED" box="[797,896,1844,1871]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="110" pageNumber="111">Fig. 507</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFF3840A8CABF8BFD999F819" blockId="110.[151,1437,153,1979]" pageId="110" pageNumber="111">Abdominal terga dorsally light brown, laterally with ocher; sterna mostly ocher; uromere VII with posterior part of sternum dark brown; uromere VIII with lateral parts of tergum and sternum brown; uromere IX with lateral parts of tergum and posterior part of sternum brown.</paragraph>
<caption id="DF416638FFF2840B8CFBF973DAEDF8B8" ID-DOI="http://doi.org/10.5281/zenodo.3737032" ID-Zenodo-Dep="3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="111" pageNumber="112" startId="111.[151,264,1685,1710]" targetBox="[157,1420,225,1620]" targetPageId="111">
<paragraph id="8B8136B0FFF2840B8CFBF973DAEDF8B8" blockId="111.[151,1437,1685,1818]" pageId="111" pageNumber="112">
<emphasis id="B94AEAA2FFF2840B8CFBF973DA4FF90C" bold="true" box="[151,369,1685,1710]" pageId="111" pageNumber="112">FIGURES 498507.</emphasis>
<taxonomicName id="4C3E4D33FFF2840B8D10F970D973F90F" box="[380,589,1686,1710]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFF2840B8D10F970D973F90F" box="[380,589,1686,1710]" italics="true" pageId="111" pageNumber="112">Thraulodes nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFF2840B8E3BF970D9AAF90C" bold="true" box="[599,660,1686,1710]" pageId="111" pageNumber="112">
<taxonomicNameLabel id="A27957D9FFF2840B8E3BF970D9AAF90C" box="[599,660,1686,1710]" pageId="111" pageNumber="112" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype).
</paragraph>
</caption>
<caption id="DF416638FFED84148CFBF833D9FCF84D" ID-DOI="http://doi.org/10.5281/zenodo.3737034" ID-Zenodo-Dep="3737034" box="[151,706,2005,2031]" httpUri="https://zenodo.org/record/3737034/files/figure.png" pageId="112" pageNumber="113" startId="112.[151,264,2005,2030]" targetBox="[193,1395,166,1980]" targetPageId="112">
<paragraph id="8B8136B0FFED84148CFBF833D9FCF84D" blockId="112.[151,706,2005,2031]" box="[151,706,2005,2031]" pageId="112" pageNumber="113">
<emphasis id="B94AEAA2FFED84148CFBF833DA53F84C" bold="true" box="[151,365,2005,2030]" pageId="112" pageNumber="113">FIGURES 508509.</emphasis>
<taxonomicName id="4C3E4D33FFED84148D1FF830D97EF84C" box="[371,576,2006,2030]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFED84148D1FF830D97EF84C" box="[371,576,2006,2030]" italics="true" pageId="112" pageNumber="113">Thraulodes nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFED84148E2BF830D9BEF84C" bold="true" box="[583,640,2006,2030]" pageId="112" pageNumber="113">
<taxonomicNameLabel id="A27957D9FFED84148E2BF830D9BEF84C" box="[583,640,2006,2030]" pageId="112" pageNumber="113" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, eggs.
</paragraph>
</caption>
<paragraph id="8B8136B0FFEC84158CABFF7FD900FF59" blockId="113.[151,1437,152,611]" pageId="113" pageNumber="114">
<emphasis id="B94AEAA2FFEC84158CABFF7FDA3FFF11" bold="true" box="[199,257,153,179]" italics="true" pageId="113" pageNumber="114">Eggs</emphasis>
(
<figureCitation id="13052A35FFEC84158D7CFF7FDA8EFF11" box="[272,432,152,179]" captionStart="FIGURES 508509" captionStartId="112.[151,264,2005,2030]" captionTargetBox="[193,1395,166,1980]" captionTargetId="figure@112.[193,1395,166,1980]" captionTargetPageId="112" captionText="FIGURES 508509. Thraulodes nigripes sp. n., eggs." figureDoi="http://doi.org/10.5281/zenodo.3737034" httpUri="https://zenodo.org/record/3737034/files/figure.png" pageId="113" pageNumber="114">Figs 508509</figureCitation>
). Mostly barrel-shaped. Each KTC surrounded by ring-like cover with smooth surface; in some areas of egg ring-like covers smoothed out, being indistinguishable from surrounding surface. Other chorion rugose, nearly without protuberances.
</paragraph>
<paragraph id="8B8136B0FFEC84158CABFEE2D8E7FEBD" blockId="113.[151,1437,152,611]" box="[199,985,260,288]" pageId="113" pageNumber="114">
<emphasis id="B94AEAA2FFEC84158CABFEE2DA70FEBD" bold="true" box="[199,334,260,287]" pageId="113" pageNumber="114">Dimension.</emphasis>
Fore wing length (and approximate body length)
<quantity id="4CC69B55FFEC84158F10FEE2D8EBFEBD" box="[892,981,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="113" pageNumber="114" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFEC84158CABFECEDF82FDC1" pageId="113" pageNumber="114" type="discussion">
<paragraph id="8B8136B0FFEC84158CABFECEDF82FDC1" blockId="113.[151,1437,152,611]" pageId="113" pageNumber="114">
<emphasis id="B94AEAA2FFEC84158CABFECEDA5CFEE1" bold="true" box="[199,354,296,323]" pageId="113" pageNumber="114">Comparison.</emphasis>
Besides the new species
<taxonomicName id="4C3E4D33FFEC84158E16FECFD83FFEE1" box="[634,769,297,323]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158E16FECFD83FFEE1" box="[634,769,297,323]" italics="true" pageId="113" pageNumber="114">Th. nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158F6BFECFD878FEE1" bold="true" box="[775,838,297,323]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158F6BFECFD878FEE1" box="[775,838,297,323]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, femora and tibiae of all legs in imaginal and subimaginal stages are uniformly dark brown only in
<taxonomicName id="4C3E4D33FFEC84158EF0FEABD83CFEC5" box="[668,770,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158EF0FEABD83CFEC5" box="[668,770,333,359]" italics="true" pageId="113" pageNumber="114">Th. niger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158F64FEABD876FEC5" bold="true" box="[776,840,333,359]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158F64FEABD876FEC5" box="[776,840,333,359]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3E4D33FFEC84158F11FEABDF5FFEC5" box="[893,1121,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158F11FEABDF5FFEC5" box="[893,1121,333,359]" italics="true" pageId="113" pageNumber="114">Th. nigrabdominalis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158804FEABDF99FEC5" bold="true" box="[1128,1191,333,359]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158804FEABDF99FEC5" box="[1128,1191,333,359]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Male
<taxonomicName id="4C3E4D33FFEC84158881FEABDE4AFEC5" box="[1261,1396,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158881FEABDE4AFEC5" box="[1261,1396,333,359]" italics="true" pageId="113" pageNumber="114">Th. nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158917FEABDB90FE29" bold="true" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158917FEABDB90FE29" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from both of them by the longer and wider dorsal extension of styliger (
<figureCitation id="13052A35FFEC84158857FE97DFA0FE29" box="[1083,1182,369,395]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="113" pageNumber="114">Fig. 499</figureCitation>
); this difference is expressed beginning from the early stage of development in last-instar larva—compare
<figureCitation id="13052A35FFEC84158826FE73DF98FE0D" box="[1098,1190,404,431]" captionStart="FIGURES 498507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503506, tergalii I, III, VI and VII (498501, 503506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="113" pageNumber="114">Fig. 502</figureCitation>
and
<figureCitation id="13052A35FFEC841588B7FE73DE05FE0D" box="[1243,1339,405,431]" captionStart="FIGURES 474478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="113" pageNumber="114">Fig. 477</figureCitation>
. Besides this, male of
<taxonomicName id="4C3E4D33FFEC84158D42FE5FDA89FE71" box="[302,439,441,467]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158D42FE5FDA89FE71" box="[302,439,441,467]" italics="true" pageId="113" pageNumber="114">Th. nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158DD3FE5FD93EFE71" bold="true" box="[447,512,441,467]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158DD3FE5FD93EFE71" box="[447,512,441,467]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C3E4D33FFEC84158EF4FE5FD83EFE71" box="[664,768,441,467]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158EF4FE5FD83EFE71" box="[664,768,441,467]" italics="true" pageId="113" pageNumber="114">Th. niger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158F65FE5FD874FE71" bold="true" box="[777,842,441,467]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158F65FE5FD874FE71" box="[777,842,441,467]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3E4D33FFEC84158FEFFE5FDF57FE71" box="[899,1129,441,467]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158FEFFE5FDF57FE71" box="[899,1129,441,467]" italics="true" pageId="113" pageNumber="114">Th. nigrabdominalis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC8415881EFE5FDF8DFE71" bold="true" box="[1138,1203,441,467]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC8415881EFE5FDF8DFE71" box="[1138,1203,441,467]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the contrastingly different coloration of abdominal terga IIVI and VIIX (
<figureCitation id="13052A35FFEC84158F71FE3BD8BEFE55" box="[797,896,476,503]" captionStart="FIGURES 491497" captionStartId="109.[151,264,1764,1789]" captionTargetBox="[151,1436,174,1744]" captionTargetId="figure@109.[151,1436,174,1744]" captionTargetPageId="109" captionText="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="113" pageNumber="114">Fig. 495</figureCitation>
). Male and female imagines and subimagines of
<taxonomicName id="4C3E4D33FFEC84158CFBFDE7DA20FDB9" box="[151,286,513,539]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158CFBFDE7DA20FDB9" box="[151,286,513,539]" italics="true" pageId="113" pageNumber="114">Th. nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158D49FDE7DA5AFDB9" bold="true" box="[293,356,513,539]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158D49FDE7DA5AFDB9" box="[293,356,513,539]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
also differ from
<taxonomicName id="4C3E4D33FFEC84158E73FDE7D9BBFDB9" box="[543,645,513,539]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158E73FDE7D9BBFDB9" box="[543,645,513,539]" italics="true" pageId="113" pageNumber="114">Th. niger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158EE0FDE7D9F5FDB9" bold="true" box="[652,715,513,539]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158EE0FDE7D9F5FDB9" box="[652,715,513,539]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by presence of dark macula close to costal brace and by absence of coloration in costal and subcostal fields of fore wing (
<figureCitation id="13052A35FFEC84158F62FDC3D84FFD9D" box="[782,881,549,575]" captionStart="FIGURES 491497" captionStartId="109.[151,264,1764,1789]" captionTargetBox="[151,1436,174,1744]" captionTargetId="figure@109.[151,1436,174,1744]" captionTargetPageId="109" captionText="FIGURES 491497. Thraulodes nigripes sp. n. 491492, fore and middle legs of female imago; 493494, female imago in dorsal and lateral view; 495, abdominal terga and sterna of male subimago extracted from larva; 496, genitalia of male subimago extracted from larva; 497, right half of subimaginal exuviae of mesonotum (495496, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737030" httpUri="https://zenodo.org/record/3737030/files/figure.png" pageId="113" pageNumber="114">Fig. 493</figureCitation>
). Larva of
<taxonomicName id="4C3E4D33FFEC84158F86FDC3DF4FFD9D" box="[1002,1137,549,575]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158F86FDC3DF4FFD9D" box="[1002,1137,549,575]" italics="true" pageId="113" pageNumber="114">Th. nigripes</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158814FDC3DF89FD9D" bold="true" box="[1144,1207,549,575]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158814FDC3DF89FD9D" box="[1144,1207,549,575]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differ both from
<taxonomicName id="4C3E4D33FFEC84158914FDC3DBECFDC1" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158914FDC3DBECFDC1" italics="true" pageId="113" pageNumber="114">Th. niger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158CB5FDAFDA27FDC1" bold="true" box="[217,281,585,611]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158CB5FDAFDA27FDC1" box="[217,281,585,611]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3E4D33FFEC84158D23FDAFD90DFDC1" box="[335,563,585,611]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
<emphasis id="B94AEAA2FFEC84158D23FDAFD90DFDC1" box="[335,563,585,611]" italics="true" pageId="113" pageNumber="114">Th. nigrabdominalis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEC84158E56FDAFD944FDC1" bold="true" box="[570,634,585,611]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="A27957D9FFEC84158E56FDAFD944FDC1" box="[570,634,585,611]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by uniformly brown coloration of abdominal terga.
</paragraph>
</subSubSection>
</treatment>
</document>