210 lines
19 KiB
XML
210 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.259.2970" ID-GBIF-Dataset="37fcf036-b7b9-407b-a090-85e2cea21080" ID-PMC="PMC3591765" ID-Pensoft-Pub="1313-2970-259-1" ID-PubMed="23653501" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-259-1" ModsDocOrigin="ZooKeys 259" ModsDocTitle="A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U.S.A." checkinTime="1451247765886" checkinUser="pensoft" docAuthor="Hunting, Wesley M." docDate="2013" docId="ED22791744779F5B842D68C25CC2CF58" docLanguage="en" docName="ZooKeys 259: 1-73" docOrigin="ZooKeys 259" docSource="http://dx.doi.org/10.3897/zookeys.259.2970" docTitle="Cymindis punctigera subsp. sulcipennis, stat. n." docType="treatment" docVersion="4" lastPageNumber="45" masterDocId="FF90904E7A57FFE9CD13EE7858701A20" masterDocTitle="A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U. S. A." masterLastPageNumber="73" masterPageNumber="1" pageNumber="42" updateTime="1668155161763" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U. S. A.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunting, Wesley M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>259</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>73</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.259.2970</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.259.2970</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-259-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152039695" LSID="urn:lsid:plazi:treatment:ED22791744779F5B842D68C25CC2CF58" httpUri="http://treatment.plazi.org/id/ED22791744779F5B842D68C25CC2CF58" lastPageId="44" lastPageNumber="45" pageId="41" pageNumber="42">
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<subSubSection pageId="41" pageNumber="42" type="nomenclature">
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName LSID="http://species-id.net/wiki/Cymindis_punctigera_sulcipennis" authority="Horn" class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="sulcipennis">Cymindis punctigera sulcipennis (Horn)</taxonomicName>
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<taxonomicNameLabel pageId="41" pageNumber="42">stat. n.</taxonomicNameLabel>
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Figs 23-24, 25B, 29
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="reference_group">
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera sulcipennis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulcipennis">Pinacodera sulcipennis</taxonomicName>
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<bibRefCitation pageId="41" pageNumber="42">Horn 1881</bibRefCitation>
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: 40. HOLOTYPE male, labeled:
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<normalizedToken originalValue="“Cal”">"Cal"</normalizedToken>
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; [small yellow paper rectangle]; "HOLOTYPE 2932" [red paper]; "
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera sulcipennis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulcipennis">Pinacodera sulcipennis</taxonomicName>
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" Horn [handwritten]. [MCZC]. -
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<bibRefCitation pageId="41" pageNumber="42">Horn 1882</bibRefCitation>
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: 147-148. - 1894: 310. TYPE AREA - Baja California, Mexico syn. n.
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</paragraph>
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera semisulcata" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="semisulcata">Pinacodera semisulcata</taxonomicName>
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<bibRefCitation pageId="41" pageNumber="42">Horn 1881</bibRefCitation>
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: 40. HOLOTYPE, male labeled:
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<normalizedToken originalValue="“Cal”">"Cal"</normalizedToken>
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; [small yellow paper rectangle]; "
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera semisulcata" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="semisulcata">Pinacodera semisulcata</taxonomicName>
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" Horn [handwritten]. [MCZC]. TYPE AREA - Baja California Sur -
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<bibRefCitation pageId="41" pageNumber="42">Horn 1882</bibRefCitation>
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: 147-149. - 1894: 310. syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="type locality">
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<paragraph pageId="41" pageNumber="42">Type locality.</paragraph>
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<paragraph pageId="41" pageNumber="42">La Paz, Baja California Sur, Mexico, here designated.</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="notes about synonymy">
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<paragraph pageId="41" pageNumber="42">Notes about synonymy.</paragraph>
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<paragraph pageId="41" pageNumber="42">
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Horn (1881:40) distinguished
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera sulcipennis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulcipennis">Pinacodera sulcipennis</taxonomicName>
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from
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera semisulcata" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="semisulcata">Pinacodera semisulcata</taxonomicName>
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through the former having raised elytral intervals. Careful examination of more than 80 specimens revealed that some members of this subspecies (32%) have raised elytral intervals, especially in the basal third. This is the rationale for combining
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera semisulcata" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="semisulcata">Pinacodera semisulcata</taxonomicName>
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with
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera sulcipennis" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulcipennis">Pinacodera sulcipennis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="diagnosis">
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<paragraph pageId="41" pageNumber="42">Diagnosis.</paragraph>
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<paragraph pageId="41" pageNumber="42">Most specimens of this subspecies (94.2%) have two to several rugulose transverse lines on dorsal surface of head between eyes with rugosity in some extended to clypeus (Fig. 25B).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
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<paragraph pageId="41" pageNumber="42">Description.</paragraph>
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<paragraph lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
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With character states of subgenus
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">Pinacodera</taxonomicName>
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and species
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="punctigera">Cymindis punctigera</taxonomicName>
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restricted as follows: OBL 8.26 - 10.67 mm. Length (n= 20 males, 20 females):
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<pageBreakToken pageId="42" pageNumber="43" start="start">head</pageBreakToken>
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0.80 - 1.02, pronotum 1.72 - 2.20, elytra 4.50 - 6.12, metepisternum 0.94 - 1.48 mm; width: head 1.72 - 2.20, pronotum 2.00 - 2.64, elytra 3.12- 4.33, metepisternum 0.45 - 0.86 mm.
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</paragraph>
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<paragraph pageId="42" pageNumber="43">
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<taxonomicName genus="Body" lsidName="Body proportions" pageId="42" pageNumber="43" rank="species" species="proportions">Body proportions</taxonomicName>
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. HW/HL 1.91 - 2.30; PWM/PL 1.08 - 1.25; EL/EW 1.26 - 1.49; ML/MW 1.40 - 1.88.
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</paragraph>
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<paragraph pageId="42" pageNumber="43">Color (Fig. 23). Dorsum of head rufous to rufo-piceous; dorsum of pronotum and elytra rufous to rufo-piceous; antennae rufo- testaceous to rufo- piceous; palpi rufo-testaceous to rufo-piceous; elytral epipleura rufo-testaceous to rufo-piceous; abdominal sterna and other thoracic sclerites rufo-testaceous to rufo-piceous.</paragraph>
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<paragraph pageId="42" pageNumber="43">Elytra (Fig. 23). Humeri narrowed.</paragraph>
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<caption pageId="42" pageNumber="43">
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<paragraph pageId="42" pageNumber="43">
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Figure 23. Dorsal habitus and color pattern of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. sulcipennis" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="sulcipennis">Cymindis punctigera sulcipennis</taxonomicName>
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(Horn) (OBL 10.83 mm).
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</paragraph>
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</caption>
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<paragraph pageId="42" pageNumber="43">Hind wings. Brachypterous.</paragraph>
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<paragraph pageId="42" pageNumber="43">Male Genitalia Phallus (cf. Fig. 24A-C) length 1.82-2.00 mm.</paragraph>
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<caption pageId="42" pageNumber="43">
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<paragraph pageId="42" pageNumber="43">
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Figure 24. Pronotum, dorsal aspect, of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. sulcipennis" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="sulcipennis">Cymindis punctigera sulcipennis</taxonomicName>
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(Horn).
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</paragraph>
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</caption>
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<caption pageId="42" pageNumber="43">
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<paragraph pageId="42" pageNumber="43">
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Figure 25. Head capsule, dorsal aspect, showing frontal macrosculpture: A,
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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LeConte; B,
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. sulcipennis" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="sulcipennis">Cymindis punctigera sulcipennis</taxonomicName>
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(Horn), note transverse lines between eyes.
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</paragraph>
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</caption>
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<caption pageId="42" pageNumber="43">
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<paragraph pageId="42" pageNumber="43">
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Figure 26. Structural features of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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LeConte: A, phallus and everted endophallus, right lateral aspect; B, female reproductive tract and ovipositor, ventral aspect. Legend: aa, apical area; bc, bursa copulatrix; bl, basal lobe; co, common oviduct; ep, endophallic plate; gc1, gonocoxite 1; gc2, gonocoxite 2; lt, lateral tergite; s, shaft; sd, spermathecal diverticulum; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca.
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</paragraph>
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</caption>
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<caption pageId="42" pageNumber="43">
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<paragraph pageId="42" pageNumber="43">
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Figure 27. Male genitalia of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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LeConte: A, phallus and everted endophallus, dorsal aspect, showing right-hand placement of endophallus when everted; B-C, left lateral aspect of phallus, endophallus inverted, showing intrapopulation variation (Jeff Davis Co., Texas, U.S.A.) in form of phallic apex; texture and distinctive striations extended from mid-apical area toward shaft. Legend: aa, apical area; bl, basal lobe; ep, endophallic plate; s, shaft.
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</paragraph>
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</caption>
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<caption pageId="42" pageNumber="43">
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<paragraph pageId="42" pageNumber="43">
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Figure 28. Right hindwings of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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LeConte, dorsal aspect: A-B, extremes of intraspecific variation from functional (A) to markedly atrophied (B); C-D, showing intrapopulation variation (Cloudcroft, New Mexico, U.S.A.) from functional (C) to somewhat atrophied (D). Legend: (from
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<bibRefCitation author="Kukalova-Peck, J" journalOrPublisher="European Journal of Entomology" pageId="57" pageNumber="58" pagination="95 - 144" title="Relationships among coleopteran suborders and major neopteran lineages: Evidence from hind wing characters." volume="101" year="2004">Kukalova-Peck and Lawrence 2004</bibRefCitation>
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) AA3, anterior anal 3; AA4, anterior anal 4; AP3+4, posterior anal 3+4; MP3, posterior median 3; MP4+CuA1+2, posterior median 4 + anterior cubitus 1+2; O, Oblongum cell; RA1+2, anterior radius 1+2; RA3+4, anterior radius 3+4; RP2, posterior radius 2; RP3+4, posterior radius 3+4; W, wedge cell.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="43" type="collection notes and habitat">
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<paragraph pageId="42" pageNumber="43">Collection notes and habitat.</paragraph>
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<paragraph pageId="42" pageNumber="43">
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The known elevational range of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. sulcipennis" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="sulcipennis">Cymindis punctigera sulcipennis</taxonomicName>
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extends from 140 to 1850 m. Specimens have been collected from stands of yucca and on shrubs of the species
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<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euphorbia misera" order="Malpighiales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="misera">Euphorbia misera</taxonomicName>
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Benth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="44" type="geographical distribution">
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<paragraph pageId="43" pageNumber="44">
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<pageBreakToken pageId="43" pageNumber="44" start="start">Geographical</pageBreakToken>
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distribution.
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</paragraph>
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<paragraph pageId="43" pageNumber="44">This subspecies is restricted to Baja California in Mexico (Fig. 29), ranging from the southern tip of the peninsula, to as far north as San Quentin, Baja California Norte.</paragraph>
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<caption pageId="43" pageNumber="44">
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<paragraph pageId="43" pageNumber="44">
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Figure 29. Map of southwestern U.S.A. and northern Mexico, showing position of localities of the subspecies of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Cymindis punctigera</taxonomicName>
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LeConte, and distribution of macropterous and brachypterous individuals of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="44" type="morphological affinities">
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<paragraph pageId="43" pageNumber="44">Morphological affinities.</paragraph>
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<paragraph pageId="43" pageNumber="44">
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This subspecies is by definition the closest relative of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="45" type="chorological affinities">
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<paragraph pageId="44" pageNumber="45">
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<pageBreakToken pageId="44" pageNumber="45" start="start">Chorological</pageBreakToken>
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affinities.
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</paragraph>
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<paragraph pageId="44" pageNumber="45">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. sulcipennis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="sulcipennis">Cymindis punctigera sulcipennis</taxonomicName>
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is allopatric with
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
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and with all other taxa of the limbata group.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="45" type="material examined">
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<paragraph pageId="44" pageNumber="45">Material examined.</paragraph>
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<paragraph pageId="44" pageNumber="45">
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I have examined 84 specimens; 10 males and 10 females were dissected. For details see University of Alberta Strickland Virtual Entomology Museum Database (
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<bibRefCitation pageId="44" pageNumber="45">University of Alberta 2009</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |