treatments-xml/data/D7/32/40/D7324026FFF1FFC792ACF9E6FE3F7416.xml

<document id="61BC9D21F4A42CCC61EE2F86A84D539B" ID-CLB-Dataset="263449" ID-DOI="10.11646/phytotaxa.607.1.7" ID-GBIF-Dataset="0c8c007a-9892-47b6-9740-7811adbe9909" ID-ISSN="1179-3163" ID-Zenodo-Dep="8212241" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1691078460740" checkinUser="plazi" docAuthor="Lu, Ngan Thi, Zhang, Liang, Thanh, Nguyen Trung, Toan, Le Chi &amp; Zhang, Li-Bing" docDate="2023" docId="D7324026FFF1FFC792ACF9E6FE3F7416" docLanguage="en" docName="phytotaxa.607.1.7.pdf" docOrigin="Phytotaxa 607 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.607.1.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Polystichum unicum Li Bing Zhang, N. T. Lu &amp; Liang Zhang 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="89" masterDocId="2B0B385EFFF3FFC39224FFDFFFB7700E" masterDocTitle="Three new species and one new record of the fern genus Polystichum (Dryopteridaceae) from Vietnam" masterLastPageNumber="94" masterPageNumber="85" pageNumber="87" updateTime="1699550717687" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="79F6BF98A1A82789485AC3ACF54E17A9">Three new species and one new record of the fern genus Polystichum (Dryopteridaceae) from Vietnam</mods:title>
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<mods:namePart id="C9193DC113BD7FD0365330D728FDA49C">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="2DBA40963A70146D8C78D411876C0CD5">vnmngan @ gmail. com; https: // orcid. org / 0000 - 0001 - 9823 - 2018</mods:affiliation>
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<mods:namePart id="31A5EFB4F9515728AB8AC3971654AF67">Zhang, Liang</mods:namePart>
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<mods:affiliation id="57308C35ECA92714E81EF176F9A92A3A">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China &amp; zhangliang @ mail. kib. ac. cn; https: // orcid. org / 0000 - 0003 - 3784 - 3135</mods:affiliation>
<mods:nameIdentifier id="55D48E7231A84515D52B9580B4B806F5" type="email">zhangliang@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="AE7B83BC29B62D750AF7A8A7143CA7F4">Thanh, Nguyen Trung</mods:namePart>
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<mods:affiliation id="7BBC25B3B8C91C55764A6D0827F56CE8">Faculty of Biology, VNU University of Science, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam &amp; thanhntsh @ gmail. com; https: // orcid. org / 0000 - 0003 - 4734 - 3775</mods:affiliation>
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<mods:namePart id="3AB024E6733A6AC0EDD9519A907948BA">Toan, Le Chi</mods:namePart>
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<mods:affiliation id="448933A35F926229A066D7F711B7EDEC">Department of Botany, Hanoi Pedagogical University 2, 32 Nguyen Van Linh, Xuan Hoa, Phuc Yen, Vietnam &amp; lechitoan @ hpu 2. edu. vn; https: // orcid. org / 0000 - 0002 - 5339 - 4440</mods:affiliation>
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<mods:namePart id="78503EFF668755A81F091B92CE31F074">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="986235CAFB3A58B2158282A37AFE0B15">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Sichuan 610041, China &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<taxonomicName id="989B8AB3FFF1FFC192ACF9E6FC80765D" ID-CoL="BSTY3" authority="Li Bing Zhang, N.T.Lu &amp; Liang Zhang" authorityName="Li Bing Zhang, N. T. Lu &amp; Liang Zhang" authorityYear="2023" box="[136,823,1593,1620]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="87" phylum="Tracheophyta" rank="species" species="unicum" status="sp. nov.">
<emphasis id="6DEF2D22FFF1FFC192ACF9E6FEC0765D" bold="true" box="[136,375,1593,1620]" italics="true" pageId="2" pageNumber="87">Polystichum unicum</emphasis>
Li Bing Zhang, N.T.Lu &amp; Liang Zhang
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, 
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</emphasis>
(
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).
</heading>
</paragraph>
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<emphasis id="6DEF2D22FFF1FFC192ACF95AFF6D7694" bold="true" box="[136,218,1669,1690]" pageId="2" pageNumber="87">Type:—</emphasis>
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<collectingCountry id="278CB1A0FFF1FFC192FEF95AFEFB7694" box="[218,332,1669,1690]" name="Vietnam" pageId="2" pageNumber="87">VIETNAM</collectingCountry>
. 
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: 
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, 
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, 
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(
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, 
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, 
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, elev. ca. 
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, at the top of a limestone mountain, 
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<collectorName id="F26E94E6FFF1FFC19733F975FA1F76B0" box="[1303,1448,1705,1727]" pageId="2" pageNumber="87">Li Bing Zhang</collectorName>
, 
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&amp; 
<collectorName id="F26E94E6FFF1FFC19373F911FE1776ED" box="[343,416,1742,1763]" pageId="2" pageNumber="87">Ngan T</collectorName>
. Lu 6933
</emphasis>
(
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!;
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!, 
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!)
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</paragraph>
</subSubSection>
<subSubSection id="1781A2BBFFF1FFC192ACF92EFE2D7741" pageId="2" pageNumber="87" type="diagnosis">
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<emphasis id="6DEF2D22FFF1FFC192ACF92EFEBC7708" bold="true" box="[136,267,1777,1798]" italics="true" pageId="2" pageNumber="87">Diagnosis:—</emphasis>
This is a very distinct species and does not seem to be similar to any species known so far in having the combination of crenulate pinna margins, distal basiscopic pinna margins forming a ca. 140° angle with the rachis, acute pinna apex, and auricles overlapping the rachis.
</paragraph>
</subSubSection>
<subSubSection id="1781A2BBFFF1FFC792ACF8A2FBE572BF" lastPageId="4" lastPageNumber="89" pageId="2" pageNumber="87" type="description">
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Plants medium-sized, evergreen, 
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tall. Rhizomes erect, ca. 
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in diam., scales similar to those of the stipe base, but softer, crushed; roots blackish when dried, up to 
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long, ca. 
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in diam., densely covered with red iridescent hairs. Fronds in clusters but few, 5 or more per rhizome; stipe brown, 
<quantity id="98635CD5FFF1FFC19604F819FBC277EE" box="[1056,1141,1990,2016]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="2" pageNumber="87" unit="cm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 cm</quantity>
long, ca. 
<quantity id="98635CD5FFF1FFC196C3F81AFA8877EE" box="[1255,1343,1989,2016]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="87" unit="mm" value="0.6">0.6 mm</quantity>
in diam., scales subulate, lanceolate or ovate-lanceolate, 0.5–3.0 × 
<quantity id="98635CD5FFF1FFC19120F835FC3B780A" box="[772,908,2026,2052]" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="2" pageNumber="87" unit="mm" value="0.6" valueMax="0.8" valueMin="0.4">0.4–0.8 mm</quantity>
, brown-blackish on abaxial side, glossy-blackish on adaxial side, thick papery, margins entire, often light brown, apex acuminate or long twisted; distal petiole scales long subulate, narrow lanceolate, 0.3–0.5 × 
<quantity id="98635CD5FFF7FFC7905BFF41FCBD70B7" box="[639,778,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="3.2" metricValueMin="1.2" pageId="4" pageNumber="89" unit="mm" value="2.2" valueMax="3.2" valueMin="1.2">1.2–3.2 mm</quantity>
, thick membranous, margins subentire to irregular dentate, apex long-acuminate. Laminae 1-pinnate, lanceolate, 2.7–3.7 × 
<quantity id="98635CD5FFF7FFC79144FF1DFC6470D3" box="[864,979,194,221]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="89" unit="cm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 cm</quantity>
, apex long acuminate; rachises ca. 
<quantity id="98635CD5FFF7FFC7974DFF1DFA1B70D3" box="[1385,1452,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="89" unit="mm" value="1.0">1 mm</quantity>
in diam., stramineous, adaxially sulcate, sparsely scaly, abaxially densely scaly; scales appressed, brown, subulate with rounded base, thick membranous, long acuminate, up to 
<quantity id="98635CD5FFF7FFC79160FED5FC30712B" box="[836,903,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="89" unit="mm" value="2.0">2 mm</quantity>
long, margins dentate at base, irregular sparsely dentate at distal part. Pinnae ca. 22 pairs, alternate, distal basiscopic margins forming a ca. 140° angle with rachis, proximal margins attached at 75° (–90°) angles to rachis, separate, 
<quantity id="98635CD5FFF7FFC7914EFE8DFC517163" box="[874,998,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="4" pageNumber="89" unit="cm" value="0.85" valueMax="1.1" valueMin="0.6">0.6–1.1 cm</quantity>
distant (from attached point), acroscopic auriculate; basal pinnae obliquely oblong, 0.5–1.2 × 2.0–6.0 mm, short stalked, apices acute, sometimes obtuse, bases obviously asymmetrical, acroscopic base with deltate or semicircular auricles, and proximal margins of auricles often overlapping rachis, margins both sides toothed, teeth mostly long spinulose apically; middle pinnae largest, oblong to obliquely oblong, 0.4–0.7 × 
<quantity id="98635CD5FFF7FFC793C1FE3DFDD271F3" box="[485,613,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="4" pageNumber="89" unit="cm" value="1.9" valueMax="2.5" valueMin="1.3">1.3–2.5 cm</quantity>
, short stalked, bases acute to obtuse, margins long spinulose, apices acute; adaxially glabrous, abaxially sparsely covered with appressed, subulate, filiform, light brown microscales; frond thickly papery texture; venation distinct abaxially, slightly distinct adaxially, lateral veins simple, nearly reaching pinna margin. Sori small, often on distal part of pinnae, lower pinnae not seen sori, often broken with debris on the leaves, terminal on shorter veinlets, 
<quantity id="98635CD5FFF7FFC79038FDADFD147283" box="[540,675,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="4" pageNumber="89" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
in diam., close to pinna margin than midvein, 0–6 on acroscopic side and 0–3 on basiscopic side, centers ca. 
<quantity id="98635CD5FFF7FFC7901BFD49FD3772BF" box="[575,640,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="89" unit="mm" value="1.0">1 mm</quantity>
apart from one another; indusia not seen.
</paragraph>
</subSubSection>
<caption id="0BE4A1B8FFF0FFC092ACF8E8FB75779B" ID-DOI="http://doi.org/10.5281/zenodo.8212247" ID-Zenodo-Dep="8212247" httpUri="https://zenodo.org/record/8212247/files/figure.png" pageId="3" pageNumber="88" startId="3.[136,234,1847,1869]" targetBox="[194,1401,202,1813]" targetPageId="3" targetType="figure">
<paragraph id="5F24F130FFF0FFC092ACF8E8FB75779B" blockId="3.[136,1452,1847,1941]" pageId="3" pageNumber="88">
<emphasis id="6DEF2D22FFF0FFC092ACF8E8FEB67742" bold="true" box="[136,257,1847,1869]" pageId="3" pageNumber="88">FIGURE 2.</emphasis>
<taxonomicName id="989B8AB3FFF0FFC09322F8E8FCF17743" authority="Li Bing Zhang, N. T. Lu &amp; Liang Zhang." authorityName="Li Bing Zhang, N. T. Lu &amp; Liang Zhang." box="[262,838,1847,1869]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="88" phylum="Tracheophyta" rank="species" species="unicum">
<emphasis id="6DEF2D22FFF0FFC09322F8E8FE7E7743" box="[262,457,1847,1869]" italics="true" pageId="3" pageNumber="88">Polystichum unicum</emphasis>
Li Bing Zhang, N.T.Lu &amp; Liang Zhang.
</taxonomicName>
—A.Adaxial view of upper portion of lamina.—B. Abaxial view of upper portion of lamina.—C. Fiddlehead.—D. Adaxial view of middle portion of lamina.—E. Portion of lamina showing pinnae with microscales.—F. Portion of petiole.—G, H &amp; L. Scales from petiole base, distal petiole, and rachis, respectively.
</paragraph>
</caption>
<subSubSection id="1781A2BBFFF7FFC79299FD65FA1072DB" box="[189,1447,698,725]" pageId="4" pageNumber="89" type="etymology">
<paragraph id="5F24F130FFF7FFC79299FD65FA1072DB" blockId="4.[136,1452,158,1048]" box="[189,1447,698,725]" pageId="4" pageNumber="89">
<emphasis id="6DEF2D22FFF7FFC79299FD65FED372DA" bold="true" box="[189,356,698,724]" pageId="4" pageNumber="89">Etymology:—</emphasis>
From the Latin 
<taxonomicName id="989B8AB3FFF7FFC79037FD64FDD272DA" box="[531,613,699,724]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="unicum">
<emphasis id="6DEF2D22FFF7FFC79037FD64FDD272DA" box="[531,613,699,724]" italics="true" pageId="4" pageNumber="89">unicum</emphasis>
</taxonomicName>
, unique, referring the unique morphology of the new species in the genus.
</paragraph>
</subSubSection>
<subSubSection id="1781A2BBFFF7FFC79299FD01FB6D72F6" box="[189,1242,734,761]" pageId="4" pageNumber="89" type="distribution">
<paragraph id="5F24F130FFF7FFC79299FD01FB6D72F6" blockId="4.[136,1452,158,1048]" box="[189,1242,734,761]" pageId="4" pageNumber="89">
<emphasis id="6DEF2D22FFF7FFC79299FD01FEC272F6" bold="true" box="[189,373,734,760]" pageId="4" pageNumber="89">Distribution:—</emphasis>
This species is currently found only in 
<collectingRegion id="9D5F3FD2FFF7FFC7910DFD00FBB572F6" box="[809,1026,734,761]" country="Vietnam" name="Ha Giang" pageId="4" pageNumber="89">Ha Giang Province</collectingRegion>
, northern 
<collectingCountry id="278CB1A0FFF7FFC79657FD01FB6172F6" box="[1139,1238,734,760]" name="Vietnam" pageId="4" pageNumber="89">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1781A2BBFFF7FFC79299FCDDFE3F7416" pageId="4" pageNumber="89" type="discussion">
<paragraph id="5F24F130FFF7FFC79299FCDDFB9B73DF" blockId="4.[136,1452,158,1048]" pageId="4" pageNumber="89">
<emphasis id="6DEF2D22FFF7FFC79299FCDDFEB77312" bold="true" box="[189,256,770,796]" pageId="4" pageNumber="89">Notes</emphasis>
:— 
<taxonomicName id="989B8AB3FFF7FFC79300FCDDFDBF7312" box="[292,520,770,796]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="unicum">
<emphasis id="6DEF2D22FFF7FFC79300FCDDFDBF7312" box="[292,520,770,796]" italics="true" pageId="4" pageNumber="89">Polystichum unicum</emphasis>
</taxonomicName>
is somehow similar to 
<taxonomicName id="989B8AB3FFF7FFC7912AFCDCFAAE7312" authority="Li Bing Zhang, M. Q. Han &amp; Yan Liu" authorityName="Li Bing Zhang, M. Q. Han &amp; Yan Liu" box="[782,1305,770,797]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="rectum">
<emphasis id="6DEF2D22FFF7FFC7912AFCDCFCC07312" box="[782,887,771,796]" italics="true" pageId="4" pageNumber="89">P. rectum</emphasis>
Li Bing Zhang, M.Q.Han &amp; Yan Liu
</taxonomicName>
in 
<bibRefCitation id="3B0A8CC1FFF7FFC79719FCDCFEBC734E" author="Han, M. - Q. &amp; Liu, Y. &amp; Zhang, L. - B." pageId="4" pageNumber="89" pagination="145 - 168" refId="ref4604" refString="Han, M. - Q., Liu, Y. &amp; Zhang, L. - B. (2018) Eight new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from limestone caves in Guangdong and Yunnan, China, with reference to species diversity in the karst terrains at high elevations in subtropical areas. Phytotaxa 365 (2): 145 - 168. https: // doi. org / 10.11646 / phytotaxa. 365.2.3" type="journal article" year="2018">
Han 
<emphasis id="6DEF2D22FFF7FFC79757FCDCFA1B7312" box="[1395,1452,770,796]" italics="true" pageId="4" pageNumber="89">et al.</emphasis>
(2018: 163)
</bibRefCitation>
in having crenulate acroscopic pinna margins, but the former has acute pinna apex, distal basiscopic pinna margins forming a ca. 140° angle with the rachis, and auricles overlapping the rachis, whereas the latter has rounded pinna apex, basiscopic distal pinna margins forming a (60–)70–90° angle with rachis, and auricles not overlapping the rachis. In addition, the new species occurs in an open mountain top with an elevation of ca. 
<quantity id="98635CD5FFF7FFC796A2FC4DFB6C73A3" box="[1158,1243,914,941]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="89" unit="m" value="1150.0">1150 m</quantity>
, whereas the latter grows in a limestone cave in southern 
<collectingRegion id="9D5F3FD2FFF7FFC79011FC68FD3973DF" box="[565,654,951,977]" country="China" name="Yunnan" pageId="4" pageNumber="89">Yunnan</collectingRegion>
, 
<collectingCountry id="278CB1A0FFF7FFC790BDFC69FD6873DE" box="[665,735,950,976]" name="China" pageId="4" pageNumber="89">China</collectingCountry>
, with an elevation of 
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.
</paragraph>
<paragraph id="5F24F130FFF7FFC79299FC05FE3F7416" blockId="4.[136,1452,158,1048]" pageId="4" pageNumber="89">
<taxonomicName id="989B8AB3FFF7FFC79299FC05FE1773FA" box="[189,416,986,1012]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="unicum">
<emphasis id="6DEF2D22FFF7FFC79299FC05FE1773FA" box="[189,416,986,1012]" italics="true" pageId="4" pageNumber="89">Polystichum unicum</emphasis>
</taxonomicName>
has auricles that overlap the rachis, which is less common (18 out of 48 species) than auricles that parallel the rachis.
</paragraph>
</subSubSection>
</treatment>
</document>