289 lines
29 KiB
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289 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1134.87853" ID-Pensoft-Pub="1313-2970-1134-129" ID-Pensoft-UUID="0973AEF60C1D54EBA960C4E252F447B0" ID-ZooBank="C79C5E40D9C64E3B816F0201713DBA77" ModsDocID="1313-2970-1134-129" checkinTime="1670590531721" checkinUser="pensoft" docAuthor="Brunke, Adam J." docDate="2022" docId="CE4046D5DAF15D83AC30F5043F1172D7" docLanguage="en" docName="ZooKeys 1134: 129-170" docOrigin="ZooKeys 1134" docPubDate="2022-12-08" docSource="http://dx.doi.org/10.3897/zookeys.1134.87853" docTitle="Iratiquedius Brunke 2022, gen. nov." docType="treatment" docUuid="9DF5C736-97AD-48AD-9BDA-A49203711E06" docUuidSource="ZooBank" docVersion="1" id="0973AEF60C1D54EBA960C4E252F447B0" lastPageNumber="129" masterDocId="0973AEF60C1D54EBA960C4E252F447B0" masterDocTitle="A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages" masterLastPageNumber="170" masterPageNumber="129" pageNumber="129" updateTime="1670590531721" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Brunke, Adam J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1158-936X</mods:nameIdentifier>
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<mods:affiliation>Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, 960 Carling Avenue, Ottawa, Ontario, Canada</mods:affiliation>
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<mods:nameIdentifier type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-08</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1134</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>170</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1134.87853</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1134.87853</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1134-129</mods:identifier>
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<mods:identifier type="ZooBank">C79C5E40D9C64E3B816F0201713DBA77</mods:identifier>
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<mods:identifier type="Pensoft-UUID">0973AEF60C1D54EBA960C4E252F447B0</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:9DF5C736-97AD-48AD-9BDA-A49203711E06" httpUri="http://treatment.plazi.org/id/CE4046D5DAF15D83AC30F5043F1172D7" lastPageNumber="129" pageId="0" pageNumber="129">
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<subSubSection pageId="0" pageNumber="129" type="nomenclature">
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<paragraph pageId="0" pageNumber="129">
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<taxonomicName LSID="https://zoobank.org/9DF5C736-97AD-48AD-9BDA-A49203711E06" authority="Brunke, 2022" authorityName="Brunke" authorityYear="2022" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" rank="genus" status="gen. nov.">Iratiquedius</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="129">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="129" type="description">
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<paragraph pageId="0" pageNumber="129">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">Figs 2A-F</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">, 3A, C-F</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">, 4A-D</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">, 6A-P</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-K median lobe of aedeagus A-C in ventral view D-K in lateral view H, J internal sac partly everted, showing paired ventral sclerites I, K completely everted L-N underside of paramere showing peg setae A, B, D, E, H, I, L, M Iratiquedius seriatus (Horn) C, F, G, J, K, N I. uncifer sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure7" httpUri="https://binary.pensoft.net/fig/779765" pageId="0" pageNumber="129">, 7A-N</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9A-E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="129" type="type species">
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<paragraph pageId="0" pageNumber="129">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="129">
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<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
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<emphasis italics="true" pageId="0" pageNumber="129">Quedius amabilis</emphasis>
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</taxonomicName>
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Smetana, 1971.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="129" type="etymology">
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<paragraph pageId="0" pageNumber="129">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="129">
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The generic name is a combination of the Latin adjective
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<normalizedToken originalValue="‘iratus’">'iratus'</normalizedToken>
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and
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<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="129">Quedius</emphasis>
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</taxonomicName>
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. It refers to the characteristic shape of the eyes, which are strongly convergent anteriad and create a comical, angry appearance.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="129" type="diagnosis">
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<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="129">
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Within
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<taxonomicName lsidName="" pageId="0" pageNumber="129" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
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,
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<taxonomicName authorityName="Brunke" authorityYear="2022" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
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</taxonomicName>
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can be distinguished from all other genera of the tribe by the distinctive eyes, which occupy nearly the entire lateral head margin, and are so convergent anteriad that their inner margin forms an obtuse angle with the suprantennal ridge (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3A</figureCitation>
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). The global diagnosis is the same as the regional Nearctic diagnosis.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="129" type="description">
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<paragraph pageId="0" pageNumber="129">Description.</paragraph>
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<paragraph pageId="0" pageNumber="129">
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Medium to small rove beetles, with variable coloration (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2A-F</figureCitation>
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). With the character states of
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<taxonomicName lsidName="" pageId="0" pageNumber="129" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
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(see
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<bibRefCitation DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. (2021)</bibRefCitation>
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) and the following: antennomere 3 longer than 2, without dense setation; outer antennomeres (8-10) about as long as wide or shorter; antennae inserted close to inner margin of eye, separated by about the width of the antennal sclerite or less; head with eyes large, strongly convex, bulging from and nearly occupying entire lateral head outline, convergent anteriad and with inner margin forming obtuse angle with suprantennal ridge (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3A</figureCitation>
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); with basal puncture doubled (at least one side), interocular punctures present in some individuals of some species (
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<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
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<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
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<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
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</taxonomicName>
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) or absent, paraocular punctures absent, genal puncture absent (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3C, D</figureCitation>
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); frons not well-developed anterolaterad of antennal insertions; labrum notched medially, creating two lobes; apical maxillary and labial palpi fusiform and glabrous; infraorbital ridge complete to mandibles; gular sutures converging towards neck and narrowly spaced posteriad; mandibles with dorsal lateral groove absent or rudimentary, right mandible with single proximal tooth, tooth simple (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3D</figureCitation>
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) or bifid (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">3C</figureCitation>
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); pronotum transverse to elongate, non-explanate, with three punctures in dorsal row, sublateral row at most reaching large lateral puncture, not extended posteriad; with only single large lateral puncture (e.g., Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4A-D</figureCitation>
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); hypomeron strongly inflexed, not visible in lateral view; basisternum with pair of macrosetae (reduced in
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<taxonomicName lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
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<emphasis italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
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<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
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</taxonomicName>
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) and well-developed longitudinal carina; scutellum impunctate; elytron with subbasal ridge complete and forming scutellar collar, disc without microsculpture between punctures; row of humeral spines present and well-developed; elytral punctation evenly distributed or in serial rows (
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<taxonomicName lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
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<emphasis italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
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</taxonomicName>
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); foretibia with lateral spines (reduced in
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<taxonomicName lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
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<emphasis italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
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</taxonomicName>
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, absent in
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<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
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<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
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</taxonomicName>
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) and apical spurs; metatarsomeres with disc setose; metatibia with at least three spines on outer face; abdominal tergite I with prototergal glands developed as moderately deep impressions, outer margin with row of setae; abdominal tergites not deeply impressed at base but some species with paired median impressions causing a
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<normalizedToken originalValue="‘pinched’">'pinched'</normalizedToken>
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appearance; abdominal sternite III with basal transverse line forming obtuse angle at middle, not produced posteriad; aedeagus with well developed paramere bearing peg setae; at least some species with discrete, paired internal sac sclerites that may be homologous with the ventral paired sclerites described by
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<bibRefCitation DOI="https://doi.org/10.1111/cla.12139" author="Brunke, AJ" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="427 - 451" refId="B2" refString="Brunke, AJ, Chatzimanolis, S, Schillhammer, H, Solodovnikov, A, 2016. Early evolution of the hyperdiverse rove beetle tribe Staphylinini (Coleoptera: Staphylinidae: Staphylininae) and a revision of its higher classification. Cladistics 32 (4): 427 - 451, DOI: https://doi.org/10.1111/cla.12139" title="Early evolution of the hyperdiverse rove beetle tribe Staphylinini (Coleoptera: Staphylinidae: Staphylininae) and a revision of its higher classification." url="https://doi.org/10.1111/cla.12139" volume="32" year="2016">Brunke et al. (2016)</bibRefCitation>
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(e.g.,
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<taxonomicName lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
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<emphasis italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="I. uncifer" pageId="0" pageNumber="129" rank="species" species="uncifer">
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<emphasis italics="true" pageId="0" pageNumber="129">I. uncifer</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A-K median lobe of aedeagus A-C in ventral view D-K in lateral view H, J internal sac partly everted, showing paired ventral sclerites I, K completely everted L-N underside of paramere showing peg setae A, B, D, E, H, I, L, M Iratiquedius seriatus (Horn) C, F, G, J, K, N I. uncifer sp. nov. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure7" httpUri="https://binary.pensoft.net/fig/779765" pageId="0" pageNumber="129">7H-K</figureCitation>
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)).
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</paragraph>
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<caption doi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="129">
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<emphasis bold="true" pageId="0" pageNumber="129">Figure 2.</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="129">A-H</emphasis>
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dorsal habitus of
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<emphasis bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius amabilis</emphasis>
|
||
</taxonomicName>
|
||
(Smetana)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
|
||
male holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
|
||
female non-type
|
||
<emphasis bold="true" pageId="0" pageNumber="129">C, D</emphasis>
|
||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||
</taxonomicName>
|
||
(Smetana)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
|
||
male non-type
|
||
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
|
||
female holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
|
||
<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
|
||
<taxonomicName lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">G</emphasis>
|
||
<taxonomicName authorityName="Horn" authorityYear="1878" class="Insecta" family="Staphylinidae" genus="Paraquedius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paraquedius puncticeps" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="puncticeps">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">Paraquedius puncticeps</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">H</emphasis>
|
||
<taxonomicName family="Staphylinidae" kingdom="Animalia" lsidName="P. marginicollis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="marginicollis">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">P. marginicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="129">
|
||
<emphasis bold="true" pageId="0" pageNumber="129">Figure 3.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="129">A-D</emphasis>
|
||
dorsal head
|
||
<emphasis bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||
showing confluence of inner eye margin and supra-antennal carina (arrow)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">E, F</emphasis>
|
||
abdominal tergites
|
||
<emphasis bold="true" pageId="0" pageNumber="129">A, D</emphasis>
|
||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Horn" baseAuthorityYear="1878" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius seriatus" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius seriatus</emphasis>
|
||
</taxonomicName>
|
||
(Horn)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
|
||
<taxonomicName baseAuthorityName="Casey" baseAuthorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) probus" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="probus" subGenus="Raphirus">Quedius (Raphirus) probus</taxonomicName>
|
||
(Casey)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">C, E</emphasis>
|
||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||
</taxonomicName>
|
||
(Smetana)
|
||
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
|
||
<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
(Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="129">C-F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="129" type="distribution">
|
||
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="129">
|
||
<taxonomicName authorityName="Brunke" authorityYear="2022" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
|
||
</taxonomicName>
|
||
is endemic to western North America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="129" type="bionomics">
|
||
<paragraph pageId="0" pageNumber="129">Bionomics.</paragraph>
|
||
<paragraph pageId="0" pageNumber="129">
|
||
Species of
|
||
<taxonomicName authorityName="Brunke" authorityYear="2022" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
|
||
</taxonomicName>
|
||
are most often found in wet moss, though
|
||
<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
is more of a generalist and can be collected from a variety of wet debris along running water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="129" type="comments">
|
||
<paragraph pageId="0" pageNumber="129">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="129">
|
||
The three included species of
|
||
<taxonomicName authorityName="Brunke" authorityYear="2022" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius" order="Coleoptera" pageId="0" pageNumber="129" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="I. seriatus" pageId="0" pageNumber="129" rank="species" species="seriatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="129">I. seriatus</emphasis>
|
||
</taxonomicName>
|
||
) were resolved together in 'Clade
|
||
<normalizedToken originalValue="A’">A'</normalizedToken>
|
||
using a phylogenomic dataset (
|
||
<bibRefCitation DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. 2021</bibRefCitation>
|
||
). Although Clade A was recovered by the concatenated but not coalescent analyses, the morphological configuration of the eyes, unique in
|
||
<taxonomicName lsidName="" pageId="0" pageNumber="129" rank="tribe" tribe="Quediini">Quediini</taxonomicName>
|
||
, provides strong further evidence for its monophyly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |