470 lines
66 KiB
XML
470 lines
66 KiB
XML
<document id="3ED2077BDC2473DB57DAB03A24852ED3" ID-DOI="10.11646/zootaxa.3981.4.1" ID-GBIF-Dataset="15c3bf52-4481-4a75-8643-7c919bdde6c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="235846" ID-ZooBank="EF71D83B-17B4-49CA-826E-D3A8E7979750" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461248924452" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2015" docId="5C5BE52CFF93BC732CB5FD3C96B60B1B" docLanguage="en" docName="zt03981p490.pdf" docOrigin="Zootaxa 3981 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cygnocyphon ibex Zwick, 2015, n. gen." docType="treatment" docVersion="8" lastPageNumber="459" masterDocId="A0629D54FF91BC7B2C22FFE99334080F" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts" masterLastPageNumber="490" masterPageNumber="451" pageNumber="453" updateTime="1698607092210" updateUser="plazi">
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<mods:mods id="70CC4999AB9C79899573385E638AD43D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0312AC1E12DEDACA5567D8FCE47404DD">
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<mods:title id="09B3D0D1E7A2B4D1BABA7AA9D03E1726">Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts</mods:title>
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<mods:name id="567F65DB822367F5AE33C179C50EB893" type="personal">
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<mods:roleTerm id="71C77F26208028E8385184061DAC5D21">Author</mods:roleTerm>
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<mods:namePart id="657E15ECB1F8CEBE955924C2F2CAC414">Zwick, Peter</mods:namePart>
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<mods:typeOfResource id="94B885A707607440BE0559C61C1531DC">text</mods:typeOfResource>
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<mods:title id="67BC140642B5937F025F3079BC002654">Zootaxa</mods:title>
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<mods:part id="C98CF0C768D89D9954F3BA83E73BE532">
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<mods:date id="2F13576C8DBBF745A9F178DB3E2DA3B5">2015</mods:date>
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<mods:detail id="A15BA1B4539F811B69F3CA599C5F3E53" type="volume">
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<mods:number id="CBD41BCF317976052C95D380422DDB2F">3981</mods:number>
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<mods:detail id="5A3281E9C907425715EE0A11E2375F52" type="issue">
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<mods:number id="1744D8A6BD587FA8C224E5CA18B78D78">4</mods:number>
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<mods:start id="004AEE1D30E4F0B3EA046C7DB3F3627F">451</mods:start>
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<mods:classification id="71F484E3A8C6B658348F750E56644D72">journal article</mods:classification>
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<mods:identifier id="CF54910978566769E0FEFC712A171F7B" type="DOI">10.11646/zootaxa.3981.4.1</mods:identifier>
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<mods:identifier id="BFFB7B94D7CF585CAA8A0F94CBA73669" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5C5BE52CFF93BC732CB5FD3C96B60B1B" ID-DOI="http://doi.org/10.5281/zenodo.6110530" ID-GBIF-Taxon="119635588" ID-Zenodo-Dep="6110530" LSID="urn:lsid:plazi:treatment:5C5BE52CFF93BC732CB5FD3C96B60B1B" httpUri="http://treatment.plazi.org/id/5C5BE52CFF93BC732CB5FD3C96B60B1B" lastPageId="8" lastPageNumber="459" pageId="2" pageNumber="453">
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<subSubSection id="9CE807B1FF93BC792CB5FD3C92250B1F" pageId="2" pageNumber="453" type="nomenclature">
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<paragraph id="D44D543AFF93BC792CB5FD3C910B0AE0" blockId="2.[151,575,725,784]" box="[151,575,725,751]" pageId="2" pageNumber="453">
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<heading id="8F05E356FF93BC792CB5FD3C910B0AE0" bold="true" box="[151,575,725,751]" fontSize="11" level="1" pageId="2" pageNumber="453" reason="1">
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<emphasis id="E6868828FF93BC792CB5FD3C910B0AE0" bold="true" box="[151,575,725,751]" pageId="2" pageNumber="453">
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||
<emphasis id="E6868828FF93BC792CB5FD3C924A0AE0" bold="true" box="[151,382,725,751]" italics="true" pageId="2" pageNumber="453">
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||
<taxonomicName id="13F22FB9FF93BC792CB5FD3C924C0AE0" box="[151,376,725,751]" class="Insecta" family="Scirtidae" genus="Cygnocyphon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="ibex" status="gen. nov.">Cygnocyphon ibex</taxonomicName>
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,
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||
</emphasis>
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||
<taxonomicNameLabel id="FDB53553FF93BC792DA7FD3C92E30AE0" box="[389,471,725,751]" pageId="2" pageNumber="453" rank="genus">n. gen.</taxonomicNameLabel>
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et n. sp.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D44D543AFF93BC792CB5FD1E92250B1F" blockId="2.[151,575,725,784]" box="[151,273,759,784]" pageId="2" pageNumber="453">
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||
(
|
||
<figureCitation id="4CC948BFFF93BC792CBDFD1E923C0B1F" box="[159,264,759,784]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 8" captionStartId-0="3.[151,264,1943,1966]" captionStartId-1="4.[151,264,1267,1290]" captionTargetBox-0="[241,1349,193,1902]" captionTargetBox-1="[175,1411,201,1234]" captionTargetId-0="figure@3.[235,1349,193,1920]" captionTargetId-1="figure@4.[156,1431,193,1246]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 5. Cygnocyphon ibex, n. gen., n. sp., male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2 – 4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium." captionText-1="FIGURES 6 – 8. Cygnocyphon ibex, n. gen., n. sp., male. 6, T 9 and S 9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus." httpUri-0="https://zenodo.org/record/235847/files/figure.png" httpUri-1="https://zenodo.org/record/235848/files/figure.png" pageId="2" pageNumber="453">Figs. 1–8</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9CE807B1FF93BC792CB5FCD6901E0BAF" pageId="2" pageNumber="453" type="materials_examined">
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||
<paragraph id="D44D543AFF93BC792CB5FCD691F20B57" blockId="2.[151,1437,831,2009]" box="[151,710,831,856]" pageId="2" pageNumber="453">
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||
<emphasis id="E6868828FF93BC792CB5FCD692960B57" bold="true" box="[151,418,831,856]" pageId="2" pageNumber="453">
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||
<typeStatus id="0B49EA98FF93BC792CB5FCD693E60B57" box="[151,210,831,856]" pageId="2" pageNumber="453">Type</typeStatus>
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and only species:
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||
</emphasis>
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||
<emphasis id="E6868828FF93BC792D88FCD691B40B57" box="[426,640,831,856]" italics="true" pageId="2" pageNumber="453">
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||
<taxonomicName id="13F22FB9FF93BC792D88FCD691480B57" box="[426,636,831,856]" class="Insecta" family="Scirtidae" genus="Cygnocyphon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="ibex" status="sp. nov.">Cygnocyphon ibex</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="FDB53553FF93BC792EA5FCD691F20B57" box="[647,710,831,856]" pageId="2" pageNumber="453" rank="species">
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<emphasis id="E6868828FF93BC792EA5FCD691F20B57" bold="true" box="[647,710,831,856]" pageId="2" pageNumber="453">n. sp.</emphasis>
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</taxonomicNameLabel>
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||
</paragraph>
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<paragraph id="D44D543AFF93BC792CE5FC8D901E0BAF" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
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||
<emphasis id="E6868828FF93BC792CE5FC8D92DD0B72" bold="true" box="[199,489,868,893]" pageId="2" pageNumber="453">
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||
<typeStatus id="0B49EA98FF93BC792CE5FC8D92360B72" box="[199,258,868,893]" pageId="2" pageNumber="453">Type</typeStatus>
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||
material.
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<typeStatus id="0B49EA98FF93BC792D5EFC8D92DD0B72" box="[380,489,868,893]" pageId="2" pageNumber="453" type="holotype">Holotype</typeStatus>
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</emphasis>
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♂: Swan R. Lea
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<emphasis id="E6868828FF93BC792E8EFC8D91D80B72" bold="true" box="[684,748,868,893]" pageId="2" pageNumber="453">[WA]</emphasis>
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(
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<collectionCode id="B2E3CCFFFF93BC792ED8FC8D90640B73" LSID="urn:lsid:biocol.org:col:34982" box="[762,848,868,892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="2" pageNumber="453">SAMA</collectionCode>
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. In poor condition, thorax and elytra on a card, head and abdomen on plastic slides in Euparal on the same pin).
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</paragraph>
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</subSubSection>
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<subSubSection id="9CE807B1FF93BC792CE5FC4592B10C93" pageId="2" pageNumber="453" type="diagnosis">
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<paragraph id="D44D543AFF93BC792CE5FC4592B10C93" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
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<emphasis id="E6868828FF93BC792CE5FC45920E0BCA" bold="true" box="[199,314,940,965]" pageId="2" pageNumber="453">Diagnosis</emphasis>
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. About
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<quantity id="130AF9DFFF93BC792DBEFC4592E10BCB" box="[412,469,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="453" unit="mm" value="2.0">2mm</quantity>
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long, elongate, pronotum transverse, barely narrower than the elytra, with rounded angles and slightly explanate sides. Head with buttonhole configuration (
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||
<bibRefCitation id="B06329CBFF93BC792FF9FC26974C0BE7" author="Zwick" box="[987,1144,975,1000]" pageId="2" pageNumber="453" refString="Zwick, P. (2013 b) Australian Marsh Beetles (Coleoptera: Scirtidae). 3. A restricted concept of genus Cyphon, Australian species of Cyphon s. str., and the new Australasian genus Nanocyphon. Genus, 24 (2), 163 - 189." type="journal article" year="2013" yearSuffix="b">Zwick 2013b</bibRefCitation>
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||
). Mandibles with 3 teeth, maxillary palpus with very long terminal segment, labial palpus straight. Antennal base unmodified (flagellum missing). Unmodified ambulatory legs. Prosternal process very small, receiving rhomboid metaventral groove poorly developed. Penis with large bilobed pala, small cap-like trigonium and long horn-like parameroids. Tegmen and parameres flat, without major spines but with elongate invaginations armed with fine spinules. A rudimentary lateral stylus present.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9CE807B1FF93BC7F2CE5FB4E90C20FDD" lastPageId="4" lastPageNumber="455" pageId="2" pageNumber="453" type="description">
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||
<paragraph id="D44D543AFF93BC792CE5FB4E974E0D07" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
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||
<emphasis id="E6868828FF93BC792CE5FB4E92FC0CCF" bold="true" box="[199,456,1191,1216]" pageId="2" pageNumber="453">Description. Habitus.</emphasis>
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||
BL
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||
<quantity id="130AF9DFFF93BC792DDDFB4191650CCF" box="[511,593,1192,1216]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="453" unit="mm" value="1.9">1.9mm</quantity>
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, BL/BW 2.0. Head blackish, remainder brown. Head and pronotum with fine sharply defined dense punctures. Punctures on elytra much larger and deep, growing finer towards apex. Pubescence decumbent, blond. Long and slender, elytra parallel-sided, body flat (
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||
<figureCitation id="4CC948BFFF93BC792805FB06975C0D07" box="[1063,1128,1263,1288]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1943,1966]" captionTargetBox="[241,1349,193,1902]" captionTargetId="figure@3.[235,1349,193,1920]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cygnocyphon ibex, n. gen., n. sp., male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2 – 4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium." httpUri="https://zenodo.org/record/235847/files/figure.png" pageId="2" pageNumber="453">Fig. 1</figureCitation>
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||
).
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||
</paragraph>
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<paragraph id="D44D543AFF93BC792CE5FAFD97290DB3" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
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Head short, a little wider than long. Eyes projecting, small, in side view almost 1/3 of their diameter above the subgenal ridge. Clypeolabrum transverse, with rounded corners. Mandibles symmetrical, strong, sharply pointed, with 3 teeth, the basal one smallest. Molar area smooth. Terminal segment of maxillary palpus long and slender, slightly concave, about twice as long as segment 3. Labial palpus straight, terminal segment long, base slightly inflated, the slender apex is blunt. Antennae missing beyond pedicel (
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<figureCitation id="4CC948BFFF93BC792F81FA4D97240DB3" box="[931,1040,1444,1469]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1943,1966]" captionTargetBox="[241,1349,193,1902]" captionTargetId="figure@3.[235,1349,193,1920]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cygnocyphon ibex, n. gen., n. sp., male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2 – 4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium." httpUri="https://zenodo.org/record/235847/files/figure.png" pageId="2" pageNumber="453">Figs. 2–4</figureCitation>
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).
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</paragraph>
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<paragraph id="D44D543AFF93BC792CE5FA2190230E42" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
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Pronotum strongly transverse, rather flat, the explanate sides only a little lower than the middle of the disc.
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<collectingCountry id="ACE514AAFF93BC792CB5FA0593FB0E0A" box="[151,207,1516,1541]" name="India" pageId="2" pageNumber="453">Hind</collectingCountry>
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margin projecting towards the triangular scutellum with convex sides. Rear corners of pronotum rounded, sides converging forward, lowered towards the rounded front corners. Fore margin medially very little projecting. Legs unmodified, not saltatorial, tibiae with paired keels.
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</paragraph>
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<paragraph id="D44D543AFF93BC792CE5F9BE90BD0F47" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
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Underside. The subgenal ridge ends far from the eyes on a small raised knob and is at the level of the maxillary base separated from the raised edge of the oral cavity by a groove (sg in
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<figureCitation id="4CC948BFFF93BC792FD1F995970F0E9B" box="[1011,1083,1660,1685]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1943,1966]" captionTargetBox="[241,1349,193,1902]" captionTargetId="figure@3.[235,1349,193,1920]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cygnocyphon ibex, n. gen., n. sp., male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2 – 4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium." httpUri="https://zenodo.org/record/235847/files/figure.png" pageId="2" pageNumber="453">Fig. 2</figureCitation>
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||
). In side view, the lower head contour has a small but distinct opening (buttonhole configuration,
|
||
<bibRefCitation id="B06329CBFF93BC792FBAF97697070EB7" author="Zwick" box="[920,1075,1695,1720]" pageId="2" pageNumber="453" refString="Zwick, P. (2013 b) Australian Marsh Beetles (Coleoptera: Scirtidae). 3. A restricted concept of genus Cyphon, Australian species of Cyphon s. str., and the new Australasian genus Nanocyphon. Genus, 24 (2), 163 - 189." type="journal article" year="2013" yearSuffix="b">Zwick 2013b</bibRefCitation>
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||
). Prosternal process very small, drop-shaped. Receiving mesosternal groove minute, indistinctly rhomboid, without delimiting ridges. There are two vertical crests on the vertical wall in front of it. The mesoventral process separates the coxae completely but is narrow and caudally truncate. It meets the metaventral process which has the form of a slender triangle. Discrimen visible over most of metaventrite. Abdominal sternites unmodified.
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||
</paragraph>
|
||
<paragraph id="D44D543AFF93BC792CE5F8BD90380FD6" blockId="2.[151,1437,831,2009]" pageId="2" pageNumber="453">
|
||
<emphasis id="E6868828FF93BC792CE5F8BD92300F62" bold="true" box="[199,260,1876,1901]" pageId="2" pageNumber="453">Male</emphasis>
|
||
. T8 broken, a pair of straight strong apodemes and small fragments of the plate covered with microtrichia and small socketed setae remain (not shown). S8 not found. T9 with very strong straight apodemes curving caudally mediad and blending with the hind margin of the bare, short, transverse plate. S9 is a weakly sclerotized approximately rectangular caudally pilose plate (
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<figureCitation id="4CC948BFFF93BC792E98F82991CB0FD6" box="[698,767,1984,2009]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[175,1411,201,1234]" captionTargetId="figure@4.[156,1431,193,1246]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Cygnocyphon ibex, n. gen., n. sp., male. 6, T 9 and S 9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus." httpUri="https://zenodo.org/record/235848/files/figure.png" pageId="2" pageNumber="453">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="808D04B2FF92BC782CB5F87E92350FE4" httpUri="https://zenodo.org/record/235847/files/figure.png" pageId="3" pageNumber="454" targetBox="[241,1349,193,1902]" targetPageId="3">
|
||
<paragraph id="D44D543AFF92BC782CB5F87E92350FE4" blockId="3.[151,1436,1943,2027]" pageId="3" pageNumber="454">
|
||
<emphasis id="E6868828FF92BC782CB5F87E92090FA2" bold="true" box="[151,317,1943,1966]" pageId="3" pageNumber="454">FIGURES 1–5.</emphasis>
|
||
<taxonomicName id="13F22FB9FF92BC782D65F87191310FA1" box="[327,517,1944,1966]" class="Insecta" family="Scirtidae" genus="Cygnocyphon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="ibex" status="gen. nov.">
|
||
<emphasis id="E6868828FF92BC782D65F87191310FA1" box="[327,517,1944,1966]" italics="true" pageId="3" pageNumber="454">Cygnocyphon ibex</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDB53553FF92BC782E34F87191550FA2" box="[534,609,1944,1965]" pageId="3" pageNumber="454" rank="genus">
|
||
<emphasis id="E6868828FF92BC782E34F87191550FA2" bold="true" box="[534,609,1944,1965]" pageId="3" pageNumber="454">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="E6868828FF92BC782E52F87191990FA2" bold="true" box="[624,685,1944,1965]" pageId="3" pageNumber="454">n. sp.</emphasis>
|
||
, male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2–4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="808D04B2FF95BC7F2CB5FB1A97B00D26" httpUri="https://zenodo.org/record/235848/files/figure.png" pageId="4" pageNumber="455" targetBox="[175,1411,201,1234]" targetPageId="4">
|
||
<paragraph id="D44D543AFF95BC7F2CB5FB1A97B00D26" blockId="4.[151,1436,1267,1321]" pageId="4" pageNumber="455">
|
||
<emphasis id="E6868828FF95BC7F2CB5FB1A920E0D06" bold="true" box="[151,314,1267,1290]" pageId="4" pageNumber="455">FIGURES 6–8.</emphasis>
|
||
<taxonomicName id="13F22FB9FF95BC7F2D62FB1D92CF0D05" box="[320,507,1268,1290]" class="Insecta" family="Scirtidae" genus="Cygnocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="ibex" status="gen. nov.">
|
||
<emphasis id="E6868828FF95BC7F2D62FB1D92CF0D05" box="[320,507,1268,1290]" italics="true" pageId="4" pageNumber="455">Cygnocyphon ibex</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDB53553FF95BC7F2E2AFB1D917B0D06" box="[520,591,1268,1289]" pageId="4" pageNumber="455" rank="genus">
|
||
<emphasis id="E6868828FF95BC7F2E2AFB1D917B0D06" bold="true" box="[520,591,1268,1289]" pageId="4" pageNumber="455">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="E6868828FF95BC7F2E79FB1D91A70D06" bold="true" box="[603,659,1268,1289]" pageId="4" pageNumber="455">n. sp.</emphasis>
|
||
, male. 6, T9 and S9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D44D543AFF95BC7F2CE5FABC964A0E65" blockId="4.[151,1437,1365,2038]" pageId="4" pageNumber="455">
|
||
Penis with very wide roundly excised pala (
|
||
<figureCitation id="4CC948BFFF95BC7F2E94FABC90310D61" box="[694,773,1365,1390]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1943,1966]" captionTargetBox="[241,1349,193,1902]" captionTargetId="figure@3.[235,1349,193,1920]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cygnocyphon ibex, n. gen., n. sp., male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2 – 4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium." httpUri="https://zenodo.org/record/235847/files/figure.png" pageId="4" pageNumber="455">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="4CC948BFFF95BC7F2F30FABF90160D61" box="[786,802,1366,1390]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[175,1411,201,1234]" captionTargetId="figure@4.[156,1431,193,1246]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Cygnocyphon ibex, n. gen., n. sp., male. 6, T 9 and S 9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus." httpUri="https://zenodo.org/record/235848/files/figure.png" pageId="4" pageNumber="455">8</figureCitation>
|
||
). The trigonium is a short hard cupule between the very large curved horn-like parameroids. Each horn bears between base and tip a regular series of triangular incisions demarcating an appressed tooth. The tegmen (
|
||
<figureCitation id="4CC948BFFF95BC7F2E88FA7491C40DB9" box="[682,752,1437,1462]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[175,1411,201,1234]" captionTargetId="figure@4.[156,1431,193,1246]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Cygnocyphon ibex, n. gen., n. sp., male. 6, T 9 and S 9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus." httpUri="https://zenodo.org/record/235848/files/figure.png" pageId="4" pageNumber="455">Fig. 7</figureCitation>
|
||
) is a narrow rectangular sclerotized frame turning caudally into two slender lateral struts reaching back to the level of the trigonium. The struts support elongate flat blade-like parameres ending in a transverse series of fine longitudinal plicae. Each paramere possesses basally, to the inside of the struts, an elongate invagination containing delicate spinules and little toothed ribs (enlarged inset in
|
||
<figureCitation id="4CC948BFFF95BC7F2965F9E096B90E2D" box="[1351,1421,1545,1570]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[175,1411,201,1234]" captionTargetId="figure@4.[156,1431,193,1246]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Cygnocyphon ibex, n. gen., n. sp., male. 6, T 9 and S 9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus." httpUri="https://zenodo.org/record/235848/files/figure.png" pageId="4" pageNumber="455">Fig. 7</figureCitation>
|
||
); details not recognized. On the outside the complex of tegmen and parameres is surrounded by a tough membrane. On each side is a partly sclerotized angular basal projection which I regard as a rudimentary stylus (st in
|
||
<figureCitation id="4CC948BFFF95BC7F290EF9B8965B0E65" box="[1324,1391,1617,1642]" captionStart="FIGURES 6 – 8" captionStartId="4.[151,264,1267,1290]" captionTargetBox="[175,1411,201,1234]" captionTargetId="figure@4.[156,1431,193,1246]" captionTargetPageId="4" captionText="FIGURES 6 – 8. Cygnocyphon ibex, n. gen., n. sp., male. 6, T 9 and S 9, superimposed; 7, tegmen, with detail of the invaginated pocket; 8, penis. Figs. 7 (excl. inset) and 8 to the same scale. st?, the presumed rudimentary stylus." httpUri="https://zenodo.org/record/235848/files/figure.png" pageId="4" pageNumber="455">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF95BC7F2CE5F99C92960E81" blockId="4.[151,1437,1365,2038]" box="[199,418,1653,1678]" pageId="4" pageNumber="455">
|
||
<emphasis id="E6868828FF95BC7F2CE5F99C92110E81" bold="true" box="[199,293,1653,1678]" pageId="4" pageNumber="455">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF95BC7F2CE5F97090C20FDD" blockId="4.[151,1437,1365,2038]" pageId="4" pageNumber="455">
|
||
<emphasis id="E6868828FF95BC7F2CE5F97092320EBD" bold="true" box="[199,262,1689,1714]" pageId="4" pageNumber="455">Note.</emphasis>
|
||
There is no similar Australian or Oriental species, and
|
||
<taxonomicName id="13F22FB9FF95BC7F2F58F97390FE0EBD" box="[890,970,1689,1714]" class="Insecta" family="Scirtidae" genus="Cygnocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="ibex">
|
||
<emphasis id="E6868828FF95BC7F2F58F97390FE0EBD" box="[890,970,1689,1714]" italics="true" pageId="4" pageNumber="455">C. ibex</emphasis>
|
||
</taxonomicName>
|
||
fits in no known genus. In the search for relatives, negative characters like the poor development of prosternal process and mesoventral groove do not help. The buttonhole configuration occurs also in
|
||
<taxonomicName id="13F22FB9FF95BC7F2EB9F90890040EF5" box="[667,816,1761,1786]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF95BC7F2EB9F90890040EF5" box="[667,816,1761,1786]" italics="true" pageId="4" pageNumber="455">Contacyphon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13F22FB9FF95BC7F2F60F90890E40EF5" box="[834,976,1761,1786]" class="Insecta" family="Scirtidae" genus="Petrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF95BC7F2F60F90890E40EF5" box="[834,976,1761,1786]" italics="true" pageId="4" pageNumber="455">Petrocyphon</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="E6868828FF95BC7F2836F90897950EF5" box="[1044,1185,1761,1786]" italics="true" pageId="4" pageNumber="455">Nanocyphon</emphasis>
|
||
whose males share a poorly developed or missing S8.
|
||
<emphasis id="E6868828FF95BC7F2E2FF8EC91930F11" box="[525,679,1797,1822]" italics="true" pageId="4" pageNumber="455">Cygnocyphon</emphasis>
|
||
may be related to this group.
|
||
<emphasis id="E6868828FF95BC7F2FD9F8EC97BC0F11" box="[1019,1160,1797,1822]" italics="true" pageId="4" pageNumber="455">Nanocyphon</emphasis>
|
||
differs in habitus, by its modified genitalia and the short conical end segment of the labial palpus. Habitus and structure are very diverse in
|
||
<taxonomicName id="13F22FB9FF95BC7F2CB5F8A492180F69" box="[151,300,1869,1894]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF95BC7F2CB5F8A492180F69" box="[151,300,1869,1894]" italics="true" pageId="4" pageNumber="455">Contacyphon</emphasis>
|
||
</taxonomicName>
|
||
but none of the few Australian species is at all similar. Even slender
|
||
<taxonomicName id="13F22FB9FF95BC7F2849F8A496340F69" box="[1131,1280,1869,1894]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF95BC7F2849F8A496340F69" box="[1131,1280,1869,1894]" italics="true" pageId="4" pageNumber="455">Contacyphon</emphasis>
|
||
</taxonomicName>
|
||
species have projecting or at least marked front angles of pronotum. Also,
|
||
<taxonomicName id="13F22FB9FF95BC7F2F52F89897310F85" box="[880,1029,1905,1930]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF95BC7F2F52F89897310F85" box="[880,1029,1905,1930]" italics="true" pageId="4" pageNumber="455">Contacyphon</emphasis>
|
||
</taxonomicName>
|
||
species lack a trigonium but have instead paired prosthemes. This is not the case in
|
||
<emphasis id="E6868828FF95BC7F2EECF87C905C0FA1" box="[718,872,1941,1966]" italics="true" pageId="4" pageNumber="455">Cygnocyphon</emphasis>
|
||
. The long last segment of the maxillary palpus is unique, and none of the compared genera has three mandibular teeth (
|
||
<figureCitation id="4CC948BFFF95BC7F2F86F85090DD0FDD" box="[932,1001,1977,2002]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1943,1966]" captionTargetBox="[241,1349,193,1902]" captionTargetId="figure@3.[235,1349,193,1920]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Cygnocyphon ibex, n. gen., n. sp., male. 1, habitus; 2, head, combined dorsal and ventral views; 3, right mandible; 4, labial palpus; 5, terminalia. Figs. 2 – 4 to the same, 1 and 5 not to scale. gs, gular suture; sg, subgenal ridge; tt, tentorium." httpUri="https://zenodo.org/record/235847/files/figure.png" pageId="4" pageNumber="455">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CE807B1FF95BC7E2CE5F8349214096C" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455" type="etymology">
|
||
<paragraph id="D44D543AFF95BC7E2CE5F834963F08DA" blockId="4.[151,1437,1365,2038]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455">
|
||
<emphasis id="E6868828FF95BC7F2CE5F83492720FF9" bold="true" box="[199,326,2013,2038]" pageId="4" pageNumber="455">Etymology</emphasis>
|
||
. It must be close to 100 years since the only specimen was taken on the Swan River. The scientific name of swans (
|
||
<taxonomicName id="13F22FB9FF94BC7E2D6FFF71929508A0" box="[333,417,152,175]" class="Aves" family="Anatidae" genus="Cygnus" kingdom="Animalia" order="Anseriformes" pageId="5" pageNumber="456" phylum="Chordata" rank="genus">
|
||
<emphasis id="E6868828FF94BC7E2D6FFF71929508A0" box="[333,417,152,175]" italics="true" pageId="5" pageNumber="456">Cygnus</emphasis>
|
||
</taxonomicName>
|
||
) and the historical name
|
||
<taxonomicName id="13F22FB9FF94BC7E2E99FF7E902608BF" box="[699,786,151,176]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF94BC7E2E99FF7E902608BF" box="[699,786,151,176]" italics="true" pageId="5" pageNumber="456">Cyphon</emphasis>
|
||
</taxonomicName>
|
||
are connected in allusion to the
|
||
<typeStatus id="0B49EA98FF94BC7E285EFF71979808BF" box="[1148,1196,152,176]" pageId="5" pageNumber="456">type</typeStatus>
|
||
locality. The specific name, a noun in apposition, refers to the resemblance of the parameroids with the horns of
|
||
<taxonomicName id="13F22FB9FF94BC7E28B3FF5597F508DA" box="[1169,1217,188,213]" class="Insecta" family="Scirtidae" genus="Cygnocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="455" phylum="Arthropoda" rank="species" species="ibex">
|
||
<emphasis id="E6868828FF94BC7E28B3FF5597F508DA" box="[1169,1217,188,213]" italics="true" pageId="5" pageNumber="456">Ibex</emphasis>
|
||
</taxonomicName>
|
||
goats.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CB5FEC19214096C" blockId="5.[151,619,296,355]" pageId="5" pageNumber="456">
|
||
<emphasis id="E6868828FF94BC7E2CB5FEC1915F094E" bold="true" box="[151,619,296,322]" pageId="5" pageNumber="456">
|
||
<taxonomicName id="13F22FB9FF94BC7E2CB5FEC19292094D" box="[151,422,296,322]" class="Insecta" family="Scirtidae" genus="Eximiocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="excisus" status="gen. nov.">
|
||
<emphasis id="E6868828FF94BC7E2CB5FEC19292094D" bold="true" box="[151,422,296,322]" italics="true" pageId="5" pageNumber="456">Eximiocyphon excisus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDB53553FF94BC7E2D96FEC19132094D" box="[436,518,296,322]" pageId="5" pageNumber="456" rank="genus">n. gen.</taxonomicNameLabel>
|
||
et n. sp.
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="4CC948BFFF94BC7E2CBDFEA3922C096C" box="[159,280,330,355]" captionStart-0="FIGURES 9 – 15" captionStart-1="FIGURES 16 – 20" captionStartId-0="6.[151,264,1904,1927]" captionStartId-1="7.[151,264,1663,1686]" captionTargetBox-0="[218,1350,205,1860]" captionTargetBox-1="[170,1415,185,1622]" captionTargetId-0="figure@6.[209,1369,193,1875]" captionTargetId-1="figure@7.[156,1430,175,1642]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." captionText-1="FIGURES 16 – 20. Eximiocyphon excisus, n. gen., n. sp., male. 16, T 8; 17, T 9; 18, S 9; 19, penis and tegmen, superimposed; 20, same, apices enlarged. 16 – 18 and 19, 20 to the same scale, respectively. pa, pala; pe, paramere; te, tegmen." httpUri-0="https://zenodo.org/record/235849/files/figure.png" httpUri-1="https://zenodo.org/record/235850/files/figure.png" pageId="5" pageNumber="456">Figs. 9–20</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CE807B1FF94BC7E2CB5FE7B907D09FC" pageId="5" pageNumber="456" type="materials_examined">
|
||
<paragraph id="D44D543AFF94BC7E2CB5FE7B91D909A4" blockId="5.[151,1437,402,2011]" box="[151,749,402,427]" pageId="5" pageNumber="456">
|
||
<emphasis id="E6868828FF94BC7E2CB5FE7B92AE09A4" bold="true" box="[151,410,402,427]" pageId="5" pageNumber="456">
|
||
<typeStatus id="0B49EA98FF94BC7E2CB5FE7B93E609A4" box="[151,210,402,427]" pageId="5" pageNumber="456">Type</typeStatus>
|
||
and only species
|
||
</emphasis>
|
||
:
|
||
<taxonomicName id="13F22FB9FF94BC7E2D8AFE7B919409A4" box="[424,672,402,427]" class="Insecta" family="Scirtidae" genus="Eximiocyphon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="excisus" status="sp. nov.">
|
||
<emphasis id="E6868828FF94BC7E2D8AFE7B919409A4" box="[424,672,402,427]" italics="true" pageId="5" pageNumber="456">Eximiocyphon excisus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDB53553FF94BC7E2E8CFE7B91D909A4" box="[686,749,402,427]" pageId="5" pageNumber="456" rank="species">
|
||
<emphasis id="E6868828FF94BC7E2E8CFE7B91D909A4" bold="true" box="[686,749,402,427]" pageId="5" pageNumber="456">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CE5FE5F907D09FC" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
<emphasis id="E6868828FF94BC7E2CE5FE5F913009C0" bold="true" box="[199,516,438,463]" pageId="5" pageNumber="456">
|
||
<typeStatus id="0B49EA98FF94BC7E2CE5FE5F923609C0" box="[199,258,438,463]" pageId="5" pageNumber="456">Type</typeStatus>
|
||
material. ♂
|
||
<typeStatus id="0B49EA98FF94BC7E2DBFFE5F913009C0" box="[413,516,438,463]" pageId="5" pageNumber="456" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: Carbine Tableland
|
||
<emphasis id="E6868828FF94BC7E2ECEFE5F900109C0" bold="true" box="[748,821,438,463]" pageId="5" pageNumber="456">N.Qld</emphasis>
|
||
. Upper Doolins Ck.,
|
||
<date id="A04C72FAFF94BC7E280EFE5E97F109C0" box="[1068,1221,439,463]" pageId="5" pageNumber="456" value="1990-11-30">30 Nov 1990</date>
|
||
, 1150m, Monteith, Janetzki & Sheridan Pyrethrum Trees, logs & rocks (
|
||
<collectionCode id="B2E3CCFFFF94BC7E2ECAFE32900909FC" box="[744,829,475,499]" pageId="5" pageNumber="456">QMSB</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CE807B1FF94BC7E2CE5FE1796BC0B54" pageId="5" pageNumber="456" type="diagnosis">
|
||
<paragraph id="D44D543AFF94BC7E2CE5FE1796BC0B54" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
<emphasis id="E6868828FF94BC7E2CE5FE17920E0A18" bold="true" box="[199,314,510,535]" pageId="5" pageNumber="456">Diagnosis</emphasis>
|
||
. Stout, oval, moderately domed,
|
||
<emphasis id="E6868828FF94BC7E2EEAFE1691D60A19" box="[712,738,511,534]" italics="true" pageId="5" pageNumber="456">ca</emphasis>
|
||
<quantity id="130AF9DFFF94BC7E2ECCFE1690010A19" box="[750,821,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="456" unit="mm" value="2.0">2 mm</quantity>
|
||
long. Slender ambulatory legs. Long antennae with unmodified scape and pedicel and minute antennomere 3. Frontoclypeus forming lateral extensions in front of antennal insertions. Mandibles without teeth. Last segment of maxillary palpus bottle-shaped, terminal segment of labial palpus inserted at an angle near middle of second segment. Pronotum at base as wide as elytra, narrowing towards straight front margin, angles blunt, not projecting. Underside: subgenal ridge ending freely in front. Prosternal process barely developed, not reaching level of coxal tips. Mesoventrite without receiving groove, mesoventral process short, triangular, not reaching the triangular metaventral process, middle coxae partly contiguous. Metacoxae transverse, their mesal portion large, square. Elytral epipleura wide in front, excavate. Male: terminal abdominal sternites with long pilosity curling around a large caudal notch on sternite 7. Genitalia long, spindle-shaped, tegmen and penis closely appressed, both with handle-like bases, penis apex asymmetrical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CE807B1FF94BC732CE5FC8F976E0A8A" lastPageId="8" lastPageNumber="459" pageId="5" pageNumber="456" type="description">
|
||
<paragraph id="D44D543AFF94BC7E2CE5FC8F914B0C3C" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
<emphasis id="E6868828FF94BC7E2CE5FC8F92F10B70" bold="true" box="[199,453,870,895]" pageId="5" pageNumber="456">Description. Habitus.</emphasis>
|
||
BL
|
||
<quantity id="130AF9DFFF94BC7E2DD7FC8E91680B71" box="[501,604,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" pageId="5" pageNumber="456" unit="mm" value="2.04">2.04 mm</quantity>
|
||
. Body stout, oval, 1.4 times longer than wide (
|
||
<figureCitation id="4CC948BFFF94BC7E2843FC8F97910B70" box="[1121,1189,870,895]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 9</figureCitation>
|
||
), moderately convex. Head large,
|
||
<quantity id="130AF9DFFF94BC7E2D00FC6292BC0BAD" box="[290,392,907,931]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="456" unit="mm" value="0.74">0.74 mm</quantity>
|
||
wide (
|
||
<figureCitation id="4CC948BFFF94BC7E2DF5FC6391180BAC" box="[471,556,906,931]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 12</figureCitation>
|
||
). The frontoclypeus is laterally expanded forming broad lobes in front of the antennal insertions. The labrum is small, rectangular, transverse. Mandibles slender, acutely pointed, mesal edge smooth, no teeth, undulations or setulae (
|
||
<figureCitation id="4CC948BFFF94BC7E2E4CFC3B91F60BE4" box="[622,706,978,1003]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 13</figureCitation>
|
||
). The bottle-shaped terminal segment of the maxillary palpus is about as long as the penultimate one. Labial palpus with last segment standing at a right angle near midlength of the enlarged penultimate segment (
|
||
<figureCitation id="4CC948BFFF94BC7E2E3DFBF391460C3C" box="[543,626,1050,1075]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CE5FBD6909A0CCC" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
Pronotum with straight front margin, angles not projecting. Rear margin gently arched but straight in front of the scutellum which is an equilateral triangle. The irregularly punctate elytra are regularly arched, widest near midlength. There is a shallow longitudinal depression near the lateral margin in the anterior third of the elytra (
|
||
<figureCitation id="4CC948BFFF94BC7E294DFB6F93930CCC" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 9</figureCitation>
|
||
). The anterior portion of the epipleura is wide and concave (
|
||
<figureCitation id="4CC948BFFF94BC7E2F6FFB4390940CCC" box="[845,928,1194,1219]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CE5FB27923A0D20" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">Entire surface covered with simple punctures, those on the front half of body quite large, finer towards the elytral apex. The integument is shining, dark brown. The short brown setae curve forward on the head, backward elsewhere.</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CE5FAD291160DB0" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
Antennae and legs are brown, except scape and pedicel yellowish. Antennae half as long as body. pedicel smaller than scape, antennomere 3 minute, much shorter than pedicel. Antennomeres 4 and following are cylindrical, 3–4 times as long as wide, the last with blunt tip (
|
||
<figureCitation id="4CC948BFFF94BC7E2F6FFA6B90950D94" box="[845,929,1410,1435]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 14</figureCitation>
|
||
). Legs slender and long (hind legs now lost), keels on the tibiae with fine spines.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CE5FA2397620E98" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
Ventral face. The short gular sutures end at the tentorial invaginations from where only a faint line extends forward. Front end of the subgenal ridge unconnected, bent inward for a very short distance, parallel to the fold coming from the gular suture (
|
||
<figureCitation id="4CC948BFFF94BC7E2DD4F9FB917F0E24" box="[502,587,1554,1579]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 13</figureCitation>
|
||
). Prosternal process minute, a low crest not reaching the level of the front coxae, no distinct groove to receive it on mesoventrite. The middle coxae are medially contiguous (
|
||
<figureCitation id="4CC948BFFF94BC7E2923F9DF96600E40" box="[1281,1364,1590,1615]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 10</figureCitation>
|
||
), both the mesoventral and the metaventral process are short triangles partly separating the coxae. The ventral part of the hind coxae is large, rectangular, extending outward as an almost parallel narrow strip.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7E2CE5F94B90410F60" blockId="5.[151,1437,402,2011]" pageId="5" pageNumber="456">
|
||
<emphasis id="E6868828FF94BC7E2CE5F94B92300EB4" bold="true" box="[199,260,1698,1723]" pageId="5" pageNumber="456">Male</emphasis>
|
||
. Sternites 3–6 unmodified, S7 with deep U-shaped notch (
|
||
<figureCitation id="4CC948BFFF94BC7E2F8AF94B90C80EB4" box="[936,1020,1698,1723]" captionStart="FIGURES 9 – 15" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[218,1350,205,1860]" captionTargetId="figure@6.[209,1369,193,1875]" captionTargetPageId="6" captionText="FIGURES 9 – 15. Eximiocyphon excisus, n. gen., n. sp., male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12 – 14 to the same scale. c 1, c 2, c 3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process." httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="5" pageNumber="456">Fig. 11</figureCitation>
|
||
). Pilosity on S6 divergent, long and curving around the notch on S7. S8 not observed. T8 (
|
||
<figureCitation id="4CC948BFFF94BC7E2F1BF92F90A50ED0" box="[825,913,1734,1759]" captionStart="FIGURES 16 – 20" captionStartId="7.[151,264,1663,1686]" captionTargetBox="[170,1415,185,1622]" captionTargetId="figure@7.[156,1430,175,1642]" captionTargetPageId="7" captionText="FIGURES 16 – 20. Eximiocyphon excisus, n. gen., n. sp., male. 16, T 8; 17, T 9; 18, S 9; 19, penis and tegmen, superimposed; 20, same, apices enlarged. 16 – 18 and 19, 20 to the same scale, respectively. pa, pala; pe, paramere; te, tegmen." httpUri="https://zenodo.org/record/235850/files/figure.png" pageId="5" pageNumber="456">Fig. 16</figureCitation>
|
||
) with long curved apodemes, the antecosta connecting them has a small forward projection in the middle. Plate long, with transparent window in middle, bare except for a fringe of fine microtrichia along the rounded caudal margin. T9 not sclerotized, transparent, free part of apodemes longer than those of T8. S9 is a small transparent anteriorly unpaired sclerite with 2 tongue-shaped caudal lobes, each with 1 or 2 small socketed setae (
|
||
<figureCitation id="4CC948BFFF94BC7E2EC2F8BF905C0F60" box="[736,872,1878,1903]" captionStart="FIGURES 16 – 20" captionStartId="7.[151,264,1663,1686]" captionTargetBox="[170,1415,185,1622]" captionTargetId="figure@7.[156,1430,175,1642]" captionTargetPageId="7" captionText="FIGURES 16 – 20. Eximiocyphon excisus, n. gen., n. sp., male. 16, T 8; 17, T 9; 18, S 9; 19, penis and tegmen, superimposed; 20, same, apices enlarged. 16 – 18 and 19, 20 to the same scale, respectively. pa, pala; pe, paramere; te, tegmen." httpUri="https://zenodo.org/record/235850/files/figure.png" pageId="5" pageNumber="456">Figs. 17, 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF94BC7C2CE5F89397700F4D" blockId="5.[151,1437,402,2011]" lastBlockId="7.[151,1437,1761,2038]" lastPageId="7" lastPageNumber="458" pageId="5" pageNumber="456">
|
||
Penis and tegmen (
|
||
<figureCitation id="4CC948BFFF94BC7E2D86F89391040F9C" box="[420,560,1914,1939]" captionStart="FIGURES 16 – 20" captionStartId="7.[151,264,1663,1686]" captionTargetBox="[170,1415,185,1622]" captionTargetId="figure@7.[156,1430,175,1642]" captionTargetPageId="7" captionText="FIGURES 16 – 20. Eximiocyphon excisus, n. gen., n. sp., male. 16, T 8; 17, T 9; 18, S 9; 19, penis and tegmen, superimposed; 20, same, apices enlarged. 16 – 18 and 19, 20 to the same scale, respectively. pa, pala; pe, paramere; te, tegmen." httpUri="https://zenodo.org/record/235850/files/figure.png" pageId="5" pageNumber="456">Figs. 19, 20</figureCitation>
|
||
) form a spindle-shaped complex about
|
||
<quantity id="130AF9DFFF94BC7E2FDFF89297740F9D" box="[1021,1088,1915,1939]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="456" unit="mm" value="1.0">1 mm</quantity>
|
||
long but only
|
||
<quantity id="130AF9DFFF94BC7E28CDF89296610F9D" box="[1263,1365,1915,1939]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="456" unit="mm" value="0.12">0.12 mm</quantity>
|
||
wide. Attempts to separate parts were unsuccessful, the oblique position of the components is an artifact. The parameres are caudally narrowly divided. The long and narrow handle-like base seems to represent the tegmen, its front end is slightly widened and asymmetrical. The pale penis is a little shorter and much narrower, with long and narrow pala. The caudal third is asymmetrically divided into two long struts. The longer one has a wider base and caudolaterally a few teeth. The shorter strut has only a single minute outward tooth at the blunt tip.
|
||
</paragraph>
|
||
<caption id="808D04B2FF97BC7D2CB5F899925C0FEC" httpUri="https://zenodo.org/record/235849/files/figure.png" pageId="6" pageNumber="457" targetBox="[218,1350,205,1860]" targetPageId="6">
|
||
<paragraph id="D44D543AFF97BC7D2CB5F899925C0FEC" blockId="6.[151,1436,1904,2019]" pageId="6" pageNumber="457">
|
||
<emphasis id="E6868828FF97BC7D2CB5F89992730F89" bold="true" box="[151,327,1904,1927]" pageId="6" pageNumber="457">FIGURES 9–15.</emphasis>
|
||
<taxonomicName id="13F22FB9FF97BC7D2D6FF89891190F88" box="[333,557,1905,1927]" class="Insecta" family="Scirtidae" genus="Eximiocyphon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="excisus" status="gen. nov.">
|
||
<emphasis id="E6868828FF97BC7D2D6FF89891190F88" box="[333,557,1905,1927]" italics="true" pageId="6" pageNumber="457">Eximiocyphon excisus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDB53553FF97BC7D2E18F89891B60F89" box="[570,642,1905,1926]" pageId="6" pageNumber="457" rank="genus">
|
||
<emphasis id="E6868828FF97BC7D2E18F89891B60F89" bold="true" box="[570,642,1905,1926]" pageId="6" pageNumber="457">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="E6868828FF97BC7D2EACF89891FC0F89" bold="true" box="[654,712,1905,1926]" pageId="6" pageNumber="457">n. sp.</emphasis>
|
||
, male. 9, body contour; 10, ventral view, after removal of head, middle and hind legs, and abdomen, diagrammatic; 11, tip of abdomen, ventral; 12, 13, head, dorsal and ventral; 14, antenna; 15, labium. Figs. 9, 10 not to scale, 12–14 to the same scale. c1, c2, c3, pro-, meso-, and metacoxa; ms, mesoventral process; mt, metaventral process.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="808D04B2FF96BC7C2CB5F99697E50EBB" httpUri="https://zenodo.org/record/235850/files/figure.png" pageId="7" pageNumber="458" targetBox="[170,1415,185,1622]" targetPageId="7">
|
||
<paragraph id="D44D543AFF96BC7C2CB5F99697E50EBB" blockId="7.[151,1436,1663,1716]" pageId="7" pageNumber="458">
|
||
<emphasis id="E6868828FF96BC7C2CB5F99692670E9B" bold="true" box="[151,339,1663,1686]" pageId="7" pageNumber="458">FIGURES 16–20.</emphasis>
|
||
<taxonomicName id="13F22FB9FF96BC7C2D7AF996910C0E99" box="[344,568,1663,1686]" class="Insecta" family="Scirtidae" genus="Eximiocyphon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="excisus" status="gen. nov.">
|
||
<emphasis id="E6868828FF96BC7C2D7AF996910C0E99" box="[344,568,1663,1686]" italics="true" pageId="7" pageNumber="458">Eximiocyphon excisus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E6868828FF96BC7C2E66F96991BF0E9A" bold="true" box="[580,651,1664,1685]" pageId="7" pageNumber="458">
|
||
<taxonomicNameLabel id="FDB53553FF96BC7C2E66F96991BF0E9A" box="[580,651,1664,1685]" pageId="7" pageNumber="458" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="E6868828FF96BC7C2EB4F96991FB0E9A" bold="true" box="[662,719,1664,1685]" pageId="7" pageNumber="458">n. sp.</emphasis>
|
||
, male. 16, T8; 17, T9; 18, S9; 19, penis and tegmen, superimposed; 20, same, apices enlarged. 16–18 and 19, 20 to the same scale, respectively. pa, pala; pe, paramere; te, tegmen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D44D543AFF96BC7C2CE5F8A492840F69" blockId="7.[151,1437,1761,2038]" box="[199,432,1869,1894]" pageId="7" pageNumber="458">
|
||
<emphasis id="E6868828FF96BC7C2CE5F8A492110F69" bold="true" box="[199,293,1869,1894]" pageId="7" pageNumber="458">Female.</emphasis>
|
||
Not known.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF96BC7C2CE5F89893CB0FF9" blockId="7.[151,1437,1761,2038]" pageId="7" pageNumber="458">
|
||
<emphasis id="E6868828FF96BC7C2CE5F89892250F85" bold="true" box="[199,273,1905,1930]" pageId="7" pageNumber="458">Notes.</emphasis>
|
||
The closest relative of
|
||
<emphasis id="E6868828FF96BC7C2E3AF898918C0F85" box="[536,696,1905,1930]" italics="true" pageId="7" pageNumber="458">Eximiocyphon</emphasis>
|
||
is unknown. Each of the remarkable character expressions of this little beetle resembles some other genus. However, resemblances concern only single characters, others differ greatly. Therefore, similarities with other taxa seem to have arisen independently and provide no evidence of close relations.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF99BC732CB5FF71911308F7" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="459">
|
||
• A laterally widened frontoclypeus (not to be confused with the forward projecting lobes of some
|
||
<taxonomicName id="13F22FB9FF99BC7328DFFF7E96A808BF" box="[1277,1436,151,176]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC7328DFFF7E96A808BF" box="[1277,1436,151,176]" italics="true" pageId="8" pageNumber="459">Prionocyphon</emphasis>
|
||
</taxonomicName>
|
||
spp.!) occurs in some
|
||
<emphasis id="E6868828FF99BC732D9FFF54912408DB" box="[445,528,189,212]" italics="true" pageId="8" pageNumber="459">
|
||
<taxonomicName id="13F22FB9FF99BC732D9FFF54913808DB" box="[445,524,189,212]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">Scirtes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
e.g.,
|
||
<taxonomicName id="13F22FB9FF99BC732E6FFF54900708DA" authority="Watts. The" authorityName="Watts. The" box="[589,819,188,213]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="kaytae">
|
||
<emphasis id="E6868828FF99BC732E6FFF54918608DA" box="[589,690,188,213]" italics="true" pageId="8" pageNumber="459">S. kaytae</emphasis>
|
||
Watts. The
|
||
</taxonomicName>
|
||
square coxal plates also remind one of these saltatorial forms (e.g.,
|
||
<bibRefCitation id="B06329CBFF99BC732D6DFF0992E608F7" author="Watts" box="[335,466,224,248]" pageId="8" pageNumber="459" refString="Watts, C. H. S. (2004) Revision of Australian Scirtes Illiger and Ora Clark (Coleoptera: Scirtidae). Transactions of the Royal Society of South Australia, 128 (1), 131 - 168." type="journal article" year="2004">Watts 2004</bibRefCitation>
|
||
: fig. 1).
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF99BC732CB5FEEC97250913" blockId="8.[151,1437,151,645]" box="[151,1041,260,285]" pageId="8" pageNumber="459">
|
||
• The free, slightly recurved end of the subgenal ridge resembles
|
||
<taxonomicName id="13F22FB9FF99BC732FB2FEEC90E80913" box="[912,988,261,284]" class="Insecta" family="Scirtidae" genus="Scirtes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC732FB2FEEC90E80913" box="[912,988,261,284]" italics="true" pageId="8" pageNumber="459">Scirtes</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF99BC732CB5FEC19070094F" blockId="8.[151,1437,151,645]" box="[151,836,295,320]" pageId="8" pageNumber="459">• The extremely reduced prosternal process is exceptional.</paragraph>
|
||
<paragraph id="D44D543AFF99BC732CB5FEA4924709A2" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="459">
|
||
• The short mesoventral process and the partly contiguous middle coxae resemble the redescription of the African-Madagascan genus
|
||
<taxonomicName id="13F22FB9FF99BC732E23FE8691970987" box="[513,675,367,392]" class="Insecta" family="Scirtidae" genus="Brachycyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC732E23FE8691970987" box="[513,675,367,392]" italics="true" pageId="8" pageNumber="459">Brachycyphon</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="B06329CBFF99BC732EEEFE9990620987" author="Ruta" box="[716,854,368,392]" pageId="8" pageNumber="459" refString="Ruta, R. (2009) Brachycyphon Fairmaire, 1896, a neglected genus of Afrotropical Scirtidae (Coleoptera). Annales Zoologici (Warszawa), 59 (4), 621 - 628." type="journal article" year="2009">Ruta (2009)</bibRefCitation>
|
||
. However, other characters, especially the genitalia, do not fit at all.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF99BC732CB5FE5197EB09FA" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="459">
|
||
• The spindle shape of the genitalia superficially resembles some Oriental species, e.g., the Japanese
|
||
<taxonomicName id="13F22FB9FF99BC732CE5FE3591A409FA" authority="Yoshitomi, 2005" authorityName="Yoshitomi" authorityYear="2005" box="[199,656,476,501]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="amami">
|
||
<emphasis id="E6868828FF99BC732CE5FE35928809FB" box="[199,444,476,501]" italics="true" pageId="8" pageNumber="459">Ypsiloncyphon amami</emphasis>
|
||
(
|
||
<bibRefCitation id="B06329CBFF99BC732DE9FE3591BC09FA" author="Yoshitomi" box="[459,648,476,501]" pageId="8" pageNumber="459" refString="Yoshitomi, H. (2005) Systematic revision of the family Scirtidae of Japan, with phylogeny, morphology and bionomics (Insecta: Coleoptera, Scirtoidea). Japanese Journal of Systematic Entomology, Monographic Series, 3, 1 - 212." type="journal article" year="2005">Yoshitomi, 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
with a cigar-shaped penis but very different tegmen.
|
||
</paragraph>
|
||
<paragraph id="D44D543AFF99BC732CB5FDE9976E0A8A" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="459">
|
||
• The narrow handle shaped base of the penis resembles Species Group 3 of
|
||
<taxonomicName id="13F22FB9FF99BC732834FE16978C0A17" box="[1046,1208,511,536]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC732834FE16978C0A17" box="[1046,1208,511,536]" italics="true" pageId="8" pageNumber="459">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
. Species Group 3 is endemic in
|
||
<collectingCountry id="ACE514AAFF99BC732D6BFDCD92840A32" box="[329,432,548,573]" name="Australia" pageId="8" pageNumber="459">Australia</collectingCountry>
|
||
and New
|
||
<collectingCountry id="ACE514AAFF99BC732E00FDCD91470A32" box="[546,627,548,573]" name="Guinea-Bissau" pageId="8" pageNumber="459">Guinea</collectingCountry>
|
||
(
|
||
<bibRefCitation id="B06329CBFF99BC732EA3FDCD902F0A32" author="Zwick" box="[641,795,548,573]" pageId="8" pageNumber="459" refString="Zwick, P. (2014 b) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014" yearSuffix="b">Zwick 2014b</bibRefCitation>
|
||
), the other species groups of
|
||
<taxonomicName id="13F22FB9FF99BC732842FDCD96360A32" box="[1120,1282,548,573]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC732842FDCD96360A32" box="[1120,1282,548,573]" italics="true" pageId="8" pageNumber="459">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
are Oriental. However, there are differences in many other structures. For example, in
|
||
<taxonomicName id="13F22FB9FF99BC732814FDAE97EC0A6F" box="[1078,1240,583,608]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC732814FDAE97EC0A6F" box="[1078,1240,583,608]" italics="true" pageId="8" pageNumber="459">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
the mesoventral process is well developed and often caudally widened (
|
||
<bibRefCitation id="B06329CBFF99BC732F12FD85909E0A8B" author="Ruta" box="[816,938,620,644]" pageId="8" pageNumber="459" refString="Ruta, R. (2009) Brachycyphon Fairmaire, 1896, a neglected genus of Afrotropical Scirtidae (Coleoptera). Annales Zoologici (Warszawa), 59 (4), 621 - 628." type="journal article" year="2009">Ruta 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B06329CBFF99BC732F97FD85977B0A8A" author="Zwick" box="[949,1103,620,645]" pageId="8" pageNumber="459" refString="Zwick, P. (2014 b) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014" yearSuffix="b">Zwick 2014b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CE807B1FF99BC732CE5FD5D96B60B1B" pageId="8" pageNumber="459" type="etymology">
|
||
<paragraph id="D44D543AFF99BC732CE5FD5D96B60B1B" blockId="8.[151,1436,692,789]" pageId="8" pageNumber="459">
|
||
<emphasis id="E6868828FF99BC732CE5FD5D92720AC2" bold="true" box="[199,326,692,717]" pageId="8" pageNumber="459">Etymology</emphasis>
|
||
. The genus name is an agglutination of Latin
|
||
<emphasis id="E6868828FF99BC732F5DFD5C90EF0AC3" box="[895,987,693,716]" italics="true" pageId="8" pageNumber="459">eximius,</emphasis>
|
||
extraordinary, distinguished, with the classical name,
|
||
<taxonomicName id="13F22FB9FF99BC732D6AFD3E92AB0AFF" box="[328,415,727,752]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6868828FF99BC732D6AFD3E92AB0AFF" box="[328,415,727,752]" italics="true" pageId="8" pageNumber="459">Cyphon</emphasis>
|
||
</taxonomicName>
|
||
. It was chosen to underline that the beetle is distinguished by several unusual, presumably derived character expressions. The specific name (the Latin adjective
|
||
<taxonomicName id="13F22FB9FF99BC732F87FD1490C20B1B" box="[933,1014,765,788]" class="Insecta" family="Scirtidae" genus="Eximiocyphon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="excisus">
|
||
<emphasis id="E6868828FF99BC732F87FD1490C20B1B" box="[933,1014,765,788]" italics="true" pageId="8" pageNumber="459">excisus</emphasis>
|
||
</taxonomicName>
|
||
, excised) refers to the notch on S7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |