317 lines
39 KiB
XML
317 lines
39 KiB
XML
<document id="A42D845A2FBFBCE75A4A4AAAAD01A14F" ID-DOI="10.26107/RBZ-2021-0016" ID-ISSN="2345-7600" ID-Zenodo-Dep="5352100" ID-ZooBank="563ADA2B-90D3-465C-9E2A-6595189F447E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630207664921" checkinUser="diego" docAuthor="Ahyong, Shane T. & Ng, Peter K. L." docDate="2021" docId="5C2F2B730A5B6836FF95F8EDFF7EFB1E" docLanguage="en" docName="RBZ-2021-0016.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Arcotheres lanensis Ahyong & Ng 2021, new combination" docType="treatment" docVersion="4" lastPageNumber="195" masterDocId="A016530B0A5F6831FFE2FFF7FF90FFAB" masterDocTitle="The pinnotherid crabs from the Gulf of Siam described by Rathbun (1909) (Decapoda: Brachyura): revisited and revised" masterLastPageNumber="211" masterPageNumber="188" pageNumber="192" updateTime="1699185341332" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3CD32D21EF5A5BF154560BDA9B90FA0E">The pinnotherid crabs from the Gulf of Siam described by Rathbun (1909) (Decapoda: Brachyura): revisited and revised</mods:title>
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<mods:namePart id="08E63BEA5AD176F5DAA3E0547C7326E2">Ahyong, Shane T.</mods:namePart>
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<mods:namePart id="F7D82BE07EAA2EE9AA761416BDD37099">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B245A718ACB426DD5EF61B8321E30CFB">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="79E98665BFBCBDA5AEC3546D2F2ACB5F">2021</mods:date>
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<paragraph id="D4399A650A5B6835FF95F8EDFD7AF899" blockId="4.[119,746,1818,1874]" box="[119,746,1818,1842]" pageId="4" pageNumber="192">
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<taxonomicName id="1386E1E60A5B6835FF95F8EDFD88F899" ID-CoL="8KPS3" authority="(Rathbun, 1909)" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[119,536,1818,1842]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="lanensis" status="comb. nov.">
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<emphasis id="E6F246770A5B6835FF95F8EDFEC2F899" bold="true" box="[119,338,1818,1842]" italics="true" pageId="4" pageNumber="192">Arcotheres lanensis</emphasis>
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(
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<bibRefCitation id="B017E7940A5B6835FE80F8EDFD80F899" author="Rathbun MJ" box="[354,528,1818,1842]" pageId="4" pageNumber="192" pagination="107 - 114" refId="ref15662" refString="Rathbun MJ (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22: 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="FDC1FB0C0A5B6835FDC7F8EDFD7AF899" box="[549,746,1818,1842]" pageId="4" pageNumber="192" rank="species">new combination</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="9C9CC9EE0A5B6835FE96F8CDFE7EF8F9" box="[372,494,1850,1874]" pageId="4" pageNumber="192" type="description">
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<paragraph id="D4399A650A5B6835FE96F8CDFE7EF8F9" blockId="4.[119,746,1818,1874]" box="[372,494,1850,1874]" pageId="4" pageNumber="192">
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(
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<figureCitation id="4CBD86E00A5B6835FE9EF8CDFE59F8F9" box="[380,457,1850,1874]" captionStart="Fig" captionStartId="5.[136,171,1686,1708]" captionTargetBox="[136,1475,189,1671]" captionTargetId="figure-7@5.[136,1475,189,1671]" captionTargetPageId="5" captionText="Fig. 4. Arcotheres lanensis (Rathbun, 1909), ovigerous female holotype, cl 2.3 mm, cw 2.6 mm, Koh Lan, Thailand, ZMUC CRU-7022. A, overall habitus; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.5352108" httpUri="https://zenodo.org/record/5352108/files/figure.png" pageId="4" pageNumber="192">Figs. 4</figureCitation>
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,
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<figureCitation id="4CBD86E00A5B6835FE35F8CDFE75F8F9" box="[471,485,1850,1874]" captionStart="Fig" captionStartId="6.[113,148,1617,1639]" captionTargetBox="[113,1451,190,1602]" captionTargetId="figure-10@6.[109,1457,168,1624]" captionTargetPageId="6" captionText="Fig. 5. Arcotheres lanensis (Rathbun, 1909), ovigerous female holotype, cl 2.3 mm, cw 2.6 mm, Koh Lan, Thailand, ZMUC CRU-7022. A, carapace; B, right maxilliped 3; C, lower buccal margin; D, anterior thoracic sternum; E, right chela; F–I, left P2–5, respectively; J, right P2; K–L, right P4, P5, respectively. Scale: A = 1.0 mm; B = 0.25 mm; C–L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5352110" httpUri="https://zenodo.org/record/5352110/files/figure.png" pageId="4" pageNumber="192">5</figureCitation>
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)
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</paragraph>
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<treatmentCitation id="5527BC740A5B6835FF93F88EFDB2F825" author="Rathbun MJ" box="[113,546,1913,1935]" page="109" pageId="4" pageNumber="192" year="1909">
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<taxonomicName id="1386E1E60A5B6835FF93F88EFDB2F825" ID-CoL="8JC5B" authority="Rathbun, 1909: 109" authorityName="Rathbun" authorityPageNumber="109" authorityYear="1909" box="[113,546,1913,1935]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="192" phylum="Arthropoda" rank="species" species="lanensis">
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<emphasis id="E6F246770A5B6835FF93F88EFED8F825" box="[113,328,1913,1934]" italics="true" pageId="4" pageNumber="192">Pinnotheres lanensis</emphasis>
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<bibRefCitation id="B017E7940A5B6835FEB0F88EFDB2F825" author="Rathbun MJ" box="[338,546,1913,1935]" pageId="4" pageNumber="192" pagination="107 - 114" refId="ref15662" refString="Rathbun MJ (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22: 107 - 114." type="journal article" year="1909">Rathbun, 1909: 109</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. —
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<treatmentCitation id="5527BC740A5B6835FDB0F88EFF2FF800" author="Rathbun MJ" page="332" pageId="4" pageNumber="192" year="1910">
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<bibRefCitation id="B017E7940A5B6835FDB0F88EFF2FF800" author="Rathbun MJ" pageId="4" pageNumber="192" pagination="301 - 367" refId="ref15689" refString="Rathbun MJ (1910) The Danish expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelige Matematiske Afhandlinger, 5: 301 - 367, pls. 1, 2, map." type="book chapter" year="1910">Rathbun, 1910: 332</bibRefCitation>
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</treatmentCitation>
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, fig. 14. —
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<treatmentCitation id="5527BC740A5B6835FEA3F861FE67F800" author="Tesch JJ" box="[321,503,1942,1964]" page="249" pageId="4" pageNumber="192" year="1918">
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<bibRefCitation id="B017E7940A5B6835FEA3F861FE67F800" author="Tesch JJ" box="[321,503,1942,1964]" pageId="4" pageNumber="192" pagination="149 - 295" refId="ref16469" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918: 249</bibRefCitation>
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</treatmentCitation>
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, 255. —
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||
<treatmentCitation id="5527BC740A5B6835FDBEF861FF2FF863" author="Gordon I" page="179" pageId="4" pageNumber="192" year="1936">
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||
<bibRefCitation id="B017E7940A5B6835FDBEF861FF2FF863" author="Gordon I" pageId="4" pageNumber="192" pagination="163 - 180" refId="ref13891" refString="Gordon I (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnean Society of London, Zoology, 40: 163 - 180." type="journal article" year="1936">Gordon, 1936: 179</bibRefCitation>
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||
</treatmentCitation>
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||
. —
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<treatmentCitation id="5527BC740A5B6835FF08F845FE07F863" author="Suvatti C" box="[234,407,1970,1992]" page="69" pageId="4" pageNumber="192" year="1938">
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<bibRefCitation id="B017E7940A5B6835FF08F845FE07F863" author="Suvatti C" box="[234,407,1970,1992]" pageId="4" pageNumber="192" refId="ref16286" refString="Suvatti C (1938) A check-list of aquatic fauna in Siam (excluding fishes). Bureau of Fisheries, Bangkok, 116 pp." type="book" year="1938">Suvatti, 1938: 69</bibRefCitation>
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||
</treatmentCitation>
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||
. —
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||
<treatmentCitation id="5527BC740A5B6835FE21F845FDEBF863" author="Suvatti C" box="[451,635,1970,1992]" page="159" pageId="4" pageNumber="192" year="1950">
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<bibRefCitation id="B017E7940A5B6835FE21F845FDEBF863" author="Suvatti C" box="[451,635,1970,1992]" pageId="4" pageNumber="192" refId="ref16312" refString="Suvatti C (1950) Fauna of Thailand. Department of Fisheries, Bangkok, ii + 1100 pp., 1 pl." type="book" year="1950">Suvatti, 1950: 159</bibRefCitation>
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||
</treatmentCitation>
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||
. —
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||
<treatmentCitation id="5527BC740A5B6835FD45F845FE06F84F" author="Silas EG & Alagarswami K" page="1201" pageId="4" pageNumber="192" year="1967">
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||
<bibRefCitation id="B017E7940A5B6835FD45F845FE06F84F" author="Silas EG & Alagarswami K" pageId="4" pageNumber="192" pagination="1161 - 1227" refId="ref16074" refString="Silas EG & Alagarswami K (1967) On an instance of parasitisation by the pea-crab (Pinnotheres sp.) on the backwater clam [Meretrix casta (Chemnitz)] from India, with a review of the work on systematics, ecology, biology and etiology of pea crabs of the genus Pinnotheres, Latreille, 1802. Proceedings of the Symposium on Crustacea held at Ernakulam, 1965, Symposium Series 2. Marine Biological Association of India, 3: 1161 - 1227." type="journal article" year="1967">Silas & Alagarswami, 1967: 1201</bibRefCitation>
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</treatmentCitation>
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||
, 1223. —
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<treatmentCitation id="5527BC740A5B6835FDE3F838FD7DF84E" author="Schmitt WL & McCain JC & Davidson E" box="[513,749,1999,2021]" page="86" pageId="4" pageNumber="192" year="1973">
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<bibRefCitation id="B017E7940A5B6835FDE3F838FD7DF84E" author="Schmitt WL & McCain JC & Davidson E" box="[513,749,1999,2021]" pageId="4" pageNumber="192" refId="ref16022" refString="Schmitt WL, McCain JC & Davidson E (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. In: Gruner H-E & Holthuis LB (eds.) Crustaceorum Catalogus. Pars 3. W. Junk B. V., Den Haag, 160 pp." type="book" year="1973">Schmitt et al., 1973: 86</bibRefCitation>
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</treatmentCitation>
|
||
. —
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<treatmentCitation id="5527BC740A5B6835FF56F81BFEE5F7AA" author="Naiyanetr P" box="[180,373,2028,2050]" page="42" pageId="4" pageNumber="192" year="1980">
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<bibRefCitation id="B017E7940A5B6835FF56F81BFEE5F7AA" author="Naiyanetr P" box="[180,373,2028,2050]" pageId="4" pageNumber="192" refId="ref14760" refString="Naiyanetr P (1980) Crustacean Fauna of Thailand (Decapoda and Stomatopoda). Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 73 pp." type="book" year="1980">Naiyanetr, 1980: 42</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="5527BC740A5B6835FE78F81BFDF6F7AA" author="Naiyanetr P" box="[410,614,2028,2050]" page="104" pageId="4" pageNumber="192" year="1998">
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<bibRefCitation id="B017E7940A5B6835FE78F81BFDF6F7AA" author="Naiyanetr P" box="[410,614,2028,2050]" pageId="4" pageNumber="192" refId="ref14791" refString="Naiyanetr P (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr, 1998: 104</bibRefCitation>
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</treatmentCitation>
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. —
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||
<treatmentCitation id="5527BC740A5B6835FD6EF81BFF6CF7B5" author="Naiyanetr P" page="118" pageId="4" pageNumber="192" year="2007">
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<bibRefCitation id="B017E7940A5B6835FD6EF81BFF6CF7B5" author="Naiyanetr P" pageId="4" pageNumber="192" refId="ref14826" refString="Naiyanetr P (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr, 2007: 118</bibRefCitation>
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</treatmentCitation>
|
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. —
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||
<treatmentCitation id="5527BC740A5B6835FEC4F7FEFE7FF7B4" author="Ng PKL & Guinot D & Davie PJF" box="[294,495,2057,2079]" page="250" pageId="4" pageNumber="192" year="2008">
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<bibRefCitation id="B017E7940A5B6835FEC4F7FEFE7FF7B4" author="Ng PKL & Guinot D & Davie PJF" box="[294,495,2057,2079]" pageId="4" pageNumber="192" pagination="1 - 286" refId="ref15203" refString="Ng PKL, Guinot D & Davie PJF (2008) Systema Brachyurorum; Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 250</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="9C9CC9EE0A5B6835FCCFF92DFB63F899" pageId="4" pageNumber="192" type="materials_examined">
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<paragraph id="D4399A650A5B6835FCCFF92DFB63F899" blockId="4.[813,1451,1754,1842]" pageId="4" pageNumber="192">
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<emphasis id="E6F246770A5B6835FCCFF92DFC4CF959" bold="true" box="[813,988,1754,1778]" pageId="4" pageNumber="192">Type material.</emphasis>
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<materialsCitation id="64EE90380A5B6835FC04F92DFB63F899" collectingDate="1900-03-02" collectionCode="ZMUC" collectorName="Koh Lan & Th. Mortensen" elevation="55" pageId="4" pageNumber="192" specimenCode="CRU-7022" specimenCount="1" typeStatus="holotype">
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<typeStatus id="0B3D24C70A5B6835FC04F92DFBC0F959" box="[998,1104,1754,1778]" pageId="4" pageNumber="192" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="B29702A00A5B6835FBBCF92DFB3CF959" box="[1118,1196,1754,1778]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="192">ZMUC</collectionCode>
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<specimenCode id="8420321E0A5B6835FB55F92DFAA1F959" box="[1207,1329,1754,1778]" collectionCode="CRU-" pageId="4" pageNumber="192">CRU-7022</specimenCode>
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, ovigerous female (cl
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, cw
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),
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<collectorName id="7973FFB30A5B6835FB73F90DFB62F8B9" box="[1169,1266,1786,1810]" pageId="4" pageNumber="192">Koh Lan</collectorName>
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, 30 fm (=
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</quantity>
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), mud, coll.
|
||
<collectorName id="7973FFB30A5B6835FC46F8EDFBD6F899" box="[932,1094,1818,1842]" pageId="4" pageNumber="192">Th. Mortensen</collectorName>
|
||
,
|
||
<date id="A038BCA50A5B6835FBB0F8EDFB7FF899" box="[1106,1263,1818,1842]" pageId="4" pageNumber="192" value="1900-03-02">
|
||
<collectingDate id="B07C454D0A5B6835FBB0F8EDFB7FF899" box="[1106,1263,1818,1842]" pageId="4" pageNumber="192" value="1900-03-02">2 March 1900</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C9CC9EE0A5B6836FCCFF8ADFE60FF5E" lastPageId="7" lastPageNumber="195" pageId="4" pageNumber="192" type="description">
|
||
<paragraph id="D4399A650A5B6834FCCFF8ADFE4CF8F3" blockId="4.[813,1453,1882,2098]" lastBlockId="5.[136,775,1792,1880]" lastPageId="5" lastPageNumber="193" pageId="4" pageNumber="192">
|
||
<emphasis id="E6F246770A5B6835FCCFF8ADFBC3F8D9" bold="true" box="[813,1107,1882,1906]" pageId="4" pageNumber="192">
|
||
Description of
|
||
<typeStatus id="0B3D24C70A5B6835FC07F8ADFBDEF8D9" box="[997,1102,1882,1906]" pageId="4" pageNumber="192" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Carapace slightly wider than long, soft, in poor condition, irregularly subcircular; front not produced, anterior margin convex in dorsal view; anterolateral margins poorly defined; dorsum smooth, glabrous; dorsal surface convex in profile. Epistome with narrow interantennular septum; median buccal margin with low, obtuse median point. Antennular sinus of similar size to orbit; antennules folded slightly obliquely. Antennal articles 1 and 2 fused to epistome. Eyes not visible in dorsal view, filling orbit, cornea pigmented.
|
||
</paragraph>
|
||
<caption id="80F9CAED0A5A6834FF6AF961FD00F962" ID-DOI="http://doi.org/10.5281/zenodo.5352108" ID-Zenodo-Dep="5352108" httpUri="https://zenodo.org/record/5352108/files/figure.png" pageId="5" pageNumber="193" startId="5.[136,171,1686,1708]" targetBox="[136,1475,189,1671]" targetPageId="5">
|
||
<paragraph id="D4399A650A5A6834FF6AF961FD00F962" blockId="5.[136,1475,1686,1737]" pageId="5" pageNumber="193">
|
||
Fig. 4.
|
||
<taxonomicName id="1386E1E60A5A6834FF32F961FDA7F906" authority="(Rathbun, 1909)" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[208,567,1686,1709]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="193" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A5A6834FF32F961FE1FF900" box="[208,399,1686,1707]" italics="true" pageId="5" pageNumber="193">Arcotheres lanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B017E7940A5A6834FE7EF961FDA0F906" author="Rathbun MJ" box="[412,560,1686,1709]" pageId="5" pageNumber="193" pagination="107 - 114" refId="ref15662" refString="Rathbun MJ (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22: 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ovigerous female holotype, cl 2.3 mm, cw 2.6 mm, Koh Lan, Thailand, ZMUC CRU-7022. A, overall habitus; B, ventral view of cephalothorax.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D4399A650A5A6834FF6AF876FAC6F893" blockId="5.[136,775,1920,2104]" lastBlockId="5.[834,1474,1792,1849]" pageId="5" pageNumber="193">Mxp3 ischiomerus length about 1.6× width; outer margin strongly convex; inner proximal margin convex; inner distal margin obtusely rounded. Carpus shorter than propodus. Propodus spatulate, apex rounded, dorsally and distally setose. Dactylus digitiform, distally setose, inserted slightly proximal to propodal midlength, apex not reaching end of propodus. Exopod inner margin almost straight, outer margin convex; flagellum with 2 articles, distally setose.</paragraph>
|
||
<paragraph id="D4399A650A5A6837FCA1F897FD10F8FB" blockId="5.[835,1475,1888,2104]" lastBlockId="6.[113,752,1752,1872]" lastPageId="6" lastPageNumber="194" pageId="5" pageNumber="193">Chelipeds symmetrical from left to right, surfaces glabrous except inner, lower distal margin. Dactylus and pollex relatively straight, crossing distally, without gape; apices simple, not expanded mesially. Dactylus occlusal margin with blunt triangular tooth proximal to midlength, margin straight in distal half, finely denticulate, sparsely setose. Pollex occlusal margin irregularly convex proximally, almost straight in distal half, finely denticulate, sparsely setose; with fringe of short setae on inner ventral margin. Propodus palm dorsal margin 1.5× height, 1.1× length of dactylus; palm ventral margin almost straight. Carpus unarmed.</paragraph>
|
||
<caption id="80F9CAED0A596837FF93F9A6FB95F90B" ID-DOI="http://doi.org/10.5281/zenodo.5352110" ID-Zenodo-Dep="5352110" httpUri="https://zenodo.org/record/5352110/files/figure.png" pageId="6" pageNumber="194" startId="6.[113,148,1617,1639]" targetBox="[113,1451,190,1602]" targetPageId="6">
|
||
<paragraph id="D4399A650A596837FF93F9A6FB95F90B" blockId="6.[113,1452,1617,1697]" pageId="6" pageNumber="194">
|
||
Fig. 5.
|
||
<taxonomicName id="1386E1E60A596837FF58F9A6FDB0F9CC" authority="(Rathbun, 1909)" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[186,544,1617,1639]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="194" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A596837FF58F9A6FEEAF9CD" box="[186,378,1617,1638]" italics="true" pageId="6" pageNumber="194">Arcotheres lanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B017E7940A596837FE64F9A6FD89F9CC" author="Rathbun MJ" box="[390,537,1617,1639]" pageId="6" pageNumber="194" pagination="107 - 114" refId="ref15662" refString="Rathbun MJ (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22: 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ovigerous female holotype, cl 2.3 mm, cw 2.6 mm, Koh Lan, Thailand, ZMUC CRU-7022. A, carapace; B, right maxilliped 3; C, lower buccal margin; D, anterior thoracic sternum; E, right chela; F–I, left P2–5, respectively; J, right P2; K–L, right P4, P5, respectively. Scale: A = 1.0 mm; B = 0.25 mm; C–L = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D4399A650A596837FF93F88FFCFBF87B" blockId="6.[112,754,1912,2096]" lastBlockId="6.[813,1453,1752,2000]" pageId="6" pageNumber="194">
|
||
P2–5 unarmed, largely glabrous, setae densest on P5 dactylus. P2, 3, and 5 similar, symmetrical from left to right; right P4 length 1.3× longer than left; relative lengths in decreasing order: P4 (longer)
|
||
<emphasis id="E6F246770A596837FEDBF82EFEDBF85B" box="[313,331,2009,2032]" italics="true" pageId="6" pageNumber="194">></emphasis>
|
||
P5
|
||
<emphasis id="E6F246770A596837FE97F82EFE17F85B" box="[373,391,2009,2032]" italics="true" pageId="6" pageNumber="194">></emphasis>
|
||
P3
|
||
<emphasis id="E6F246770A596837FE50F82EFE54F85B" box="[434,452,2009,2032]" italics="true" pageId="6" pageNumber="194">></emphasis>
|
||
P2. Longer P4 merus about 0.4× pcl. P2–3 dactylus subequal, 0.6× propodus length, longer than half length of P4 and 5 dactyli, evenly arcuate, distally acute, flexor margin unarmed, glabrous. P4 dactylus slender, weakly curved, sparsely setose, unarmed; left dactylus as long as propodus; right dactylus 0.7× propodus length. P5 dactylus slightly longer than propodus, slender, weakly curved, subequal to P4 dactylus; extensor margin with few short setae, flexor margin lined with long setae, unarmed. Relative dactylus lengths: P4 (longer) ~ P5> P3
|
||
<emphasis id="E6F246770A596837FCCCF84EFCD0F87B" box="[814,832,1977,2000]" italics="true" pageId="6" pageNumber="194">></emphasis>
|
||
P2.
|
||
</paragraph>
|
||
<paragraph id="D4399A650A596837FCCFF80FFB50F79B" blockId="6.[812,1451,2040,2096]" pageId="6" pageNumber="194">Thoracic sternum anterior margin weakly concave medially; sternites 1–3 indistinguishably fused.</paragraph>
|
||
<paragraph id="D4399A650A586836FF6AFF4AFE60FF5E" blockId="7.[136,774,189,245]" pageId="7" pageNumber="195">Pleon of 6 free somites and telson, extending to buccal region, covering bases of P2–P5.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C9CC9EE0A586836FF6AFEEAFDC9FBFE" pageId="7" pageNumber="195" type="discussion">
|
||
<paragraph id="D4399A650A586836FF6AFEEAFDC9FBFE" blockId="7.[135,776,285,1109]" pageId="7" pageNumber="195">
|
||
<emphasis id="E6F246770A586836FF6AFEEAFF67FE9E" bold="true" box="[136,247,285,309]" pageId="7" pageNumber="195">Remarks.</emphasis>
|
||
<taxonomicName id="1386E1E60A586836FF1FFEEAFE4BFE9E" authorityName="Rathbun" authorityYear="1909" box="[253,475,285,309]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FF1FFEEAFE4BFE9E" box="[253,475,285,309]" italics="true" pageId="7" pageNumber="195">Pinnotheres lanensis</emphasis>
|
||
</taxonomicName>
|
||
is referred to
|
||
<taxonomicName id="1386E1E60A586836FD92FEEAFD75FE9E" authorityName="Manning" authorityYear="1993" box="[624,741,285,309]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6F246770A586836FD92FEEAFD75FE9E" box="[624,741,285,309]" italics="true" pageId="7" pageNumber="195">Arcotheres</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the asymmetrical P4, the elongated and slender P4–5 dactyli that are longer than those of P2 and P3, and the perpendicular articulation between the ischium and merus of P5.
|
||
<taxonomicName id="1386E1E60A586836FF53FE6AFE14FE1E" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[177,388,413,437]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FF53FE6AFE14FE1E" box="[177,388,413,437]" italics="true" pageId="7" pageNumber="195">Arcotheres lanensis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="1386E1E60A586836FDABFE6AFD28FE1F" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[585,696,413,436]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis id="E6F246770A586836FDABFE6AFD28FE1F" box="[585,696,413,436]" italics="true" pageId="7" pageNumber="195">A. exiguus</emphasis>
|
||
</taxonomicName>
|
||
, which it resembles in most respects. The two species, however, differ in the proportional length of the P2 and P3 dactyli (distinctly exceeding half the length of the dactyli of P5 and the longer P
|
||
<quantity id="137E37800A586836FEEAFDEAFEA0FD9E" box="[264,304,541,565]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="195" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="1386E1E60A586836FED4FDEAFE3AFD9E" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[310,426,541,565]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FED4FDEAFE3AFD9E" box="[310,426,541,565]" italics="true" pageId="7" pageNumber="195">A. lanensis</emphasis>
|
||
</taxonomicName>
|
||
, distinctly shorter than half in
|
||
<taxonomicName id="1386E1E60A586836FD0DFDEAFF49FDFF" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis id="E6F246770A586836FD0DFDEAFF49FDFF" italics="true" pageId="7" pageNumber="195">A. exiguus</emphasis>
|
||
</taxonomicName>
|
||
), as well as the size at maturity.
|
||
<taxonomicName id="1386E1E60A586836FDD6FDCAFC96FDFE" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[564,774,573,597]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FDD6FDCAFC96FDFE" box="[564,774,573,597]" italics="true" pageId="7" pageNumber="195">Arcotheres lanensis</emphasis>
|
||
</taxonomicName>
|
||
is already ovigerous by cl
|
||
<quantity id="137E37800A586836FE53FDAAFD99FDDF" box="[433,521,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="195" unit="mm" value="2.3">2.3 mm</quantity>
|
||
, whereas
|
||
<taxonomicName id="1386E1E60A586836FD9BFDAAFD7AFDDF" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[633,746,605,628]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis id="E6F246770A586836FD9BFDAAFD7AFDDF" box="[633,746,605,628]" italics="true" pageId="7" pageNumber="195">A. exiguus</emphasis>
|
||
</taxonomicName>
|
||
is not mature until about twice that size (Ng & Ahyong, in preparation). In addition, even in
|
||
<taxonomicName id="1386E1E60A586836FDE4FD6AFDEAFD1F" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[518,634,669,692]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis id="E6F246770A586836FDE4FD6AFDEAFD1F" box="[518,634,669,692]" italics="true" pageId="7" pageNumber="195">A. exiguus</emphasis>
|
||
</taxonomicName>
|
||
of a similar size to the
|
||
<typeStatus id="0B3D24C70A586836FEE1FD4AFEF0FD7E" box="[259,352,701,725]" pageId="7" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="1386E1E60A586836FE68FD4AFD91FD7E" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[394,513,701,725]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FE68FD4AFD91FD7E" box="[394,513,701,725]" italics="true" pageId="7" pageNumber="195">A. lanensis</emphasis>
|
||
</taxonomicName>
|
||
, the P5 dactylus is still distinctly more elongated, resembling that of the largest specimens of
|
||
<taxonomicName id="1386E1E60A586836FEC1FD0AFE07FCBF" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[291,407,765,788]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis id="E6F246770A586836FEC1FD0AFE07FCBF" box="[291,407,765,788]" italics="true" pageId="7" pageNumber="195">A. exiguus</emphasis>
|
||
</taxonomicName>
|
||
(Ng & Ahyong, in preparation).
|
||
<taxonomicName id="1386E1E60A586836FF6AFCEAFECBFC9E" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[136,347,797,821]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FF6AFCEAFECBFC9E" box="[136,347,797,821]" italics="true" pageId="7" pageNumber="195">Arcotheres lanensis</emphasis>
|
||
</taxonomicName>
|
||
also resembles
|
||
<taxonomicName id="1386E1E60A586836FDE7FCEAFD1FFC9E" authorityName="Rathbun" authorityYear="1909" box="[517,655,797,821]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="E6F246770A586836FDE7FCEAFD1FFC9E" box="[517,655,797,821]" italics="true" pageId="7" pageNumber="195">A. quadratus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1386E1E60A586836FD7FFCEAFC90FC9E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[669,768,797,821]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="E6F246770A586836FD7FFCEAFC90FC9E" box="[669,768,797,821]" italics="true" pageId="7" pageNumber="195">A. similis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="1386E1E60A586836FF57FCCAFE3DFCFE" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[181,429,829,853]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="latus">
|
||
<emphasis id="E6F246770A586836FF57FCCAFE97FCFE" box="[181,263,829,853]" italics="true" pageId="7" pageNumber="195">A. latus</emphasis>
|
||
(
|
||
<bibRefCitation id="B017E7940A586836FEF7FCCAFE35FCFE" author="Burger O" box="[277,421,829,853]" pageId="7" pageNumber="195" pagination="361 - 390" refId="ref13553" refString="Burger O (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8: 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in the presence of the elongated P2 and P3 dactyli that exceed half the length of the dactyli of P5 and the longer P4, but is readily separated by the unarmed flexor margin of the P5 dactylus (versus one or two rows of spinules on the distal half of the flexor margin in
|
||
<taxonomicName id="1386E1E60A586836FD12FC4AFF64FC5E" authorityName="Rathbun" authorityYear="1909" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="E6F246770A586836FD12FC4AFF64FC5E" italics="true" pageId="7" pageNumber="195">A. quadratus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1386E1E60A586836FEE1FC2AFEF9FC5E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[259,361,989,1013]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="E6F246770A586836FEE1FC2AFEF9FC5E" box="[259,361,989,1013]" italics="true" pageId="7" pageNumber="195">A. similis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="1386E1E60A586836FE4AFC2AFE6AFC5E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[424,506,989,1013]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="latus">
|
||
<emphasis id="E6F246770A586836FE4AFC2AFE6AFC5E" box="[424,506,989,1013]" italics="true" pageId="7" pageNumber="195">A. latus</emphasis>
|
||
</taxonomicName>
|
||
). As with
|
||
<taxonomicName id="1386E1E60A586836FD90FC2AFD71FC5F" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[626,737,989,1012]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="exiguus">
|
||
<emphasis id="E6F246770A586836FD90FC2AFD71FC5F" box="[626,737,989,1012]" italics="true" pageId="7" pageNumber="195">A. exiguus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1386E1E60A586836FD12FC2AFF5EFBBE" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="E6F246770A586836FD12FC2AFF5EFBBE" italics="true" pageId="7" pageNumber="195">A. similis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1386E1E60A586836FEE6FC0AFEC7FBBE" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[260,343,1021,1045]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="latus">
|
||
<emphasis id="E6F246770A586836FEE6FC0AFEC7FBBE" box="[260,343,1021,1045]" italics="true" pageId="7" pageNumber="195">A. latus</emphasis>
|
||
</taxonomicName>
|
||
mature at at least least twice the size of
|
||
<taxonomicName id="1386E1E60A586836FF6BFBEAFF6FFB9E" authorityName="Ahyong & Ng" authorityYear="2021" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[137,255,1053,1077]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="lanensis">
|
||
<emphasis id="E6F246770A586836FF6BFBEAFF6FFB9E" box="[137,255,1053,1077]" italics="true" pageId="7" pageNumber="195">A. lanensis</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<taxonomicName id="1386E1E60A586836FE93FBEAFE6DFB9E" authorityName="Rathbun" authorityYear="1909" box="[369,509,1053,1077]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="195" phylum="Arthropoda" rank="species" species="quadratus">
|
||
<emphasis id="E6F246770A586836FE93FBEAFE6DFB9E" box="[369,509,1053,1077]" italics="true" pageId="7" pageNumber="195">A. quadratus</emphasis>
|
||
</taxonomicName>
|
||
is mature at a similarly small size (Ng & Ahyong, in preparation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C9CC9EE0A586836FF6AFB8BFF7EFB1E" pageId="7" pageNumber="195" type="distribution">
|
||
<paragraph id="D4399A650A586836FF6AFB8BFF7EFB1E" blockId="7.[136,774,1148,1205]" pageId="7" pageNumber="195">
|
||
<emphasis id="E6F246770A586836FF6AFB8BFEB6FB3F" bold="true" box="[136,294,1148,1172]" pageId="7" pageNumber="195">Distribution.</emphasis>
|
||
Presently known only from the Gulf of
|
||
<collectingCountry id="AC91DAF50A586836FF6AFB6AFF7AFB1E" box="[136,234,1181,1205]" name="Thailand" pageId="7" pageNumber="195">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |