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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e102683" ID-Pensoft-Pub="1864-8312-81-581" ID-Pensoft-UUID="E6F5D6150A225AFBB372E719EBBEDB68" ID-ZooBank="CA56E9F0B4264014B20BD7C455FD65A7" ModsDocID="1864-8312-81-581" checkinTime="1686864336769" checkinUser="pensoft" docAuthor="Wang, Ya-Lian, Peng, Zi-Qi, Jin, Feng-Liang, Chen, Hong-Wei &amp; Huang, Jia" docDate="2023" docId="5855C064A7BD5447B34914717F4E7DEA" docLanguage="en" docName="ArthropodSystPhyl 81: 581-610" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-06-15" docSource="http://dx.doi.org/10.3897/asp.81.e102683" docTitle="Stegana (Orthostegana) fuscofemorata Peng &amp; Chen 2023, sp. nov." docType="treatment" docUuid="D4FD7C40-8F9A-427A-8A08-4A8DDDCE2F3F" docUuidSource="ZooBank" docVersion="1" id="E6F5D6150A225AFBB372E719EBBEDB68" lastPageNumber="581" masterDocId="E6F5D6150A225AFBB372E719EBBEDB68" masterDocTitle="Integrative species delimitation in the morphologically conservative Stegana subgen. Orthostegana (Diptera, Drosophilidae)" masterLastPageNumber="610" masterPageNumber="581" pageNumber="581" updateTime="1686864336769" updateUser="pensoft">
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<mods:title id="46E15CAEC1F5DEFF703B798148CD19FF">Integrative species delimitation in the morphologically conservative Stegana subgen. Orthostegana (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="198260AF835BBEAF7F3E8D3DAB359559">Wang, Ya-Lian</mods:namePart>
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<mods:affiliation id="9366777FB158ECED6A67285CD37D6AAB">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="1F6D86F9EC1E0D3B248BA5025FC2A972">Peng, Zi-Qi</mods:namePart>
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<mods:affiliation id="144D109DA4F0D45006166244F20BBE2E">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="E84D3B173FE04185CF8A523436199DA7">Jin, Feng-Liang</mods:namePart>
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<mods:affiliation id="B1E42D11EDC8E6B956E0BFDD06D895A1">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="B9142A6D6E89B6DCD7B06BDA44DC17A9">Chen, Hong-Wei</mods:namePart>
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<mods:affiliation id="621092C217510C66CA11FEE309F879B1">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="84884973F6EC1D290A02D9A89A41EAD0">Huang, Jia</mods:namePart>
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<mods:affiliation id="ACA79F3B77416ED0C520F6A22B37AB91">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:date id="B54ACD90FADE46C715050E49FD4A36F3">2023</mods:date>
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<treatment id="5855C064A7BD5447B34914717F4E7DEA" LSID="urn:lsid:zoobank.org:act:D4FD7C40-8F9A-427A-8A08-4A8DDDCE2F3F" httpUri="http://treatment.plazi.org/id/5855C064A7BD5447B34914717F4E7DEA" lastPageNumber="581" pageId="0" pageNumber="581">
<subSubSection id="06E589014568C9DB2023B8319D975033" pageId="0" pageNumber="581" type="nomenclature">
<paragraph id="12FC6EFE7EA3C2AE77071FB25A7AD1D9" pageId="0" pageNumber="581">
3.2.1.4.
<taxonomicName id="604C7C0BFBC8622D17F3D30934A6FCC7" LSID="https://zoobank.org/D4FD7C40-8F9A-427A-8A08-4A8DDDCE2F3F" authority="Peng &amp; Chen" authorityName="Peng &amp; Chen" authorityYear="2023" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegana (Orthostegana) fuscofemorata" order="Diptera" pageId="0" pageNumber="581" phylum="Arthropoda" rank="species" species="fuscofemorata" status="sp. nov." subGenus="Orthostegana">Stegana (Orthostegana) fuscofemorata Peng &amp; Chen</taxonomicName>
<taxonomicNameLabel id="51C57CC56901F06C13B80EB2E8678857" pageId="0" pageNumber="581">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="5F91505782A06AAA484E900E53A71931" pageId="0" pageNumber="581" type="description">
<paragraph id="5BCDBDB44744A025DC19ED332BD9E1E8" pageId="0" pageNumber="581">
<figureCitation id="0DBC74E8BCDB0A77543084D178BC4BB3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) cuodi Peng &amp; Chen sp. nov. (male); C, D S. (O.) cuodi Peng &amp; Chen sp. nov. (female); E, F S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (male); G, H S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (female)." figureDoi="10.3897/asp.81.e102683.figure6" httpUri="https://binary.pensoft.net/fig/864473" pageId="0" pageNumber="581">Figs 6E-H</figureCitation>
<figureCitation id="18EA6EDC9221201F7C2025234F2B65DA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Mesotarsus of males in ventral view. A Stegana (Orthostegana) curvinervis (Hendle, 1914); B S. (O.) singularis (Sidorenko, 1990); C S. (O.) flavicauda Zhang and Chen, 2012; D S. (O.) hirsutina Zhang and Chen, 2012; E S. (O.) hylecoeta Zhang and Chen, 2012; F S. (O.) multicardua Zhang and Chen, 2012; G S. (O.) aini Peng &amp; Chen sp. nov.; H S. (O.) brevivittata Peng &amp; Chen sp. nov.; I S. (O.) cuodi Peng &amp; Chen sp. nov.; J S. (O.) fuscofemorata Peng &amp; Chen sp. nov. Scale bar = 0.2 mm." figureDoi="10.3897/asp.81.e102683.figure9" httpUri="https://binary.pensoft.net/fig/864476" pageId="0" pageNumber="581">, 9J</figureCitation>
<figureCitation id="6AF9DE2360009AD067D90C4F51738637" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="49ED2AEBC22C6D76F6B98C0C27E9CBA7" pageId="0" pageNumber="581" type="diagnosis">
<paragraph id="3D52B4A7F87A4A3A0B8B1392D12314E3" pageId="0" pageNumber="581">Diagnosis.</paragraph>
<paragraph id="E5743F0C2D93044A9004548A77CCC70D" pageId="0" pageNumber="581">
This species closely resembles
<taxonomicName id="3C2C7FC0AA5080B1B0B72B60800BF7C6" lsidName="S. (O.) brevivittata" pageId="0" pageNumber="581" rank="species" species="brevivittata" subGenus="O.">S. (O.) brevivittata</taxonomicName>
<emphasis id="CC609C019E8A5243C2DD140A142AFB38" bold="true" pageId="0" pageNumber="581">sp. nov.</emphasis>
in the shape of male terminalia (Figs
<figureCitation id="2CDB774C377DEB311ECCAC28DF354CB6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Stegana (Orthostegana) brevivittata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure13" httpUri="https://binary.pensoft.net/fig/864480" pageId="0" pageNumber="581">13</figureCitation>
,
<figureCitation id="DF921072812DC43D0F503C0EFBAC3756" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">15</figureCitation>
), but can be distinguished from the latter by having the mesotarsus ventrally with 2 rows of black, apically blunt, stout spines; metatarsus with 12/3+1 spines; 2nd to 4th segments with 3/1, 2/1, and 2/1 spines, respectively (Fig.
<figureCitation id="D636352B841581262E99C9A021AC49BA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Mesotarsus of males in ventral view. A Stegana (Orthostegana) curvinervis (Hendle, 1914); B S. (O.) singularis (Sidorenko, 1990); C S. (O.) flavicauda Zhang and Chen, 2012; D S. (O.) hirsutina Zhang and Chen, 2012; E S. (O.) hylecoeta Zhang and Chen, 2012; F S. (O.) multicardua Zhang and Chen, 2012; G S. (O.) aini Peng &amp; Chen sp. nov.; H S. (O.) brevivittata Peng &amp; Chen sp. nov.; I S. (O.) cuodi Peng &amp; Chen sp. nov.; J S. (O.) fuscofemorata Peng &amp; Chen sp. nov. Scale bar = 0.2 mm." figureDoi="10.3897/asp.81.e102683.figure9" httpUri="https://binary.pensoft.net/fig/864476" pageId="0" pageNumber="581">9J</figureCitation>
). Surstylus with some setae near ventral margin on inner surface (Fig.
<figureCitation id="C09153889B2DF106209F29D9549DFF49" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">15B</figureCitation>
).
</paragraph>
<caption id="24AF65C1634E9386A426028AD3FCCA03" doi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581" start="Figure 15" startId="F15">
<paragraph id="1AEA2FD0055553B391CDE2E7CFD9AD75" pageId="0" pageNumber="581">
<emphasis id="5DD797A992F7C21C9E6D29C4CFBCE89A" bold="true" pageId="0" pageNumber="581">Figure 15.</emphasis>
<taxonomicName id="C5A83538EEDD9FE4D12C4A65195F9AFC" authorityName="Peng &amp; Chen" authorityYear="2023" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegana (Orthostegana) fuscofemorata" order="Diptera" pageId="0" pageNumber="581" phylum="Arthropoda" rank="species" species="fuscofemorata" subGenus="Orthostegana">Stegana (Orthostegana) fuscofemorata</taxonomicName>
Peng &amp; Chen
<emphasis id="DB67934209A9484728A029048EFFCA51" bold="true" pageId="0" pageNumber="581">sp. nov.</emphasis>
, male terminalia.
<emphasis id="ADE3E72BC37AFAB5EFBB7BE9245AE7A0" bold="true" pageId="0" pageNumber="581">A</emphasis>
Epandrium, surstylus, and cercus in lateral view;
<emphasis id="8BDF3B3D17077E502F2D311DFF087680" bold="true" pageId="0" pageNumber="581">B</emphasis>
Surstylus in ventral view;
<emphasis id="45565DE86C61DEEE4924DDEA20565AC3" bold="true" pageId="0" pageNumber="581">C</emphasis>
,
<emphasis id="00632532774954320119C76A53021001" bold="true" pageId="0" pageNumber="581">D</emphasis>
Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="833F276D081B0A17A2A64716D4E08902" pageId="0" pageNumber="581" type="description">
<paragraph id="FC551C40AEE12FA0B21DD80C88261D70" pageId="0" pageNumber="581">Description.</paragraph>
<paragraph id="C9E76F8BBC42192C0D852526F26752F0" pageId="0" pageNumber="581">
<emphasis id="BEF276F322639D60E4C48B5B6945FC36" bold="true" pageId="0" pageNumber="581">
Male and female.
<emphasis id="A7464A1F35E614AF380EC70AEF403DC2" italics="true" pageId="0" pageNumber="581">Head</emphasis>
</emphasis>
: Ocellar triangle brown to dark brown, with 2 small setae posterior to ocellar setae. Postocellar seta absent. Frons brownish yellow to brown, with numerous minute interfrontal setulae. Pedicel yellow; first flagellomere dark brown distally, with black pubescence. Face and gena brown to dark brown. Clypeus dark brown medially and brownish yellow laterally. Palpus brown.
<emphasis id="590274F0559B8CE99BE8EE206D55853E" bold="true" italics="true" pageId="0" pageNumber="581">Thorax</emphasis>
: Mesonotum brownish yellow to brown (Fig.
<figureCitation id="1B1AA4DD60CBF159A8F471538A5B9848" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) cuodi Peng &amp; Chen sp. nov. (male); C, D S. (O.) cuodi Peng &amp; Chen sp. nov. (female); E, F S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (male); G, H S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (female)." figureDoi="10.3897/asp.81.e102683.figure6" httpUri="https://binary.pensoft.net/fig/864473" pageId="0" pageNumber="581">6E, G</figureCitation>
). Pleura with a distinct brown to dark brown longitudinal stripe above (Fig.
<figureCitation id="22A8540B822298851437262E9E1B159E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) cuodi Peng &amp; Chen sp. nov. (male); C, D S. (O.) cuodi Peng &amp; Chen sp. nov. (female); E, F S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (male); G, H S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (female)." figureDoi="10.3897/asp.81.e102683.figure6" httpUri="https://binary.pensoft.net/fig/864473" pageId="0" pageNumber="581">6F, H</figureCitation>
). Scutellum mostly brownish yellow to brown (Fig.
<figureCitation id="4553C76F6D0AC4B66FE90640DDBD5ACD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) cuodi Peng &amp; Chen sp. nov. (male); C, D S. (O.) cuodi Peng &amp; Chen sp. nov. (female); E, F S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (male); G, H S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (female)." figureDoi="10.3897/asp.81.e102683.figure6" httpUri="https://binary.pensoft.net/fig/864473" pageId="0" pageNumber="581">6E, G</figureCitation>
). Halter mostly brownish, with a yellow patch at knob (Fig.
<figureCitation id="60C82E507AC68B9895ABF05E51127041" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) cuodi Peng &amp; Chen sp. nov. (male); C, D S. (O.) cuodi Peng &amp; Chen sp. nov. (female); E, F S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (male); G, H S. (O.) fuscofemorata Peng &amp; Chen sp. nov. (female)." figureDoi="10.3897/asp.81.e102683.figure6" httpUri="https://binary.pensoft.net/fig/864473" pageId="0" pageNumber="581">6E-H</figureCitation>
). Legs yellowish (Fig.
<figureCitation id="618BE4BD48C8AA8FFA2EDC2D4AF53775" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Mesotarsus of males in ventral view. A Stegana (Orthostegana) curvinervis (Hendle, 1914); B S. (O.) singularis (Sidorenko, 1990); C S. (O.) flavicauda Zhang and Chen, 2012; D S. (O.) hirsutina Zhang and Chen, 2012; E S. (O.) hylecoeta Zhang and Chen, 2012; F S. (O.) multicardua Zhang and Chen, 2012; G S. (O.) aini Peng &amp; Chen sp. nov.; H S. (O.) brevivittata Peng &amp; Chen sp. nov.; I S. (O.) cuodi Peng &amp; Chen sp. nov.; J S. (O.) fuscofemorata Peng &amp; Chen sp. nov. Scale bar = 0.2 mm." figureDoi="10.3897/asp.81.e102683.figure9" httpUri="https://binary.pensoft.net/fig/864476" pageId="0" pageNumber="581">9J</figureCitation>
).
<emphasis id="D4A018BB0243E515AA45A4BF7AEDD92F" bold="true" italics="true" pageId="0" pageNumber="581">Abdominal tergites</emphasis>
: brownish to brown.
<emphasis id="FE9B0BD9AB6DEE9B810EB0CE294C0959" bold="true" italics="true" pageId="0" pageNumber="581">Male terminalia</emphasis>
: Epandrium mostly pubescent except for anterior margin, with
<emphasis id="EF2407C4906632B14993B0B8F8B0FCE7" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
16 setae on dorsal to posterior portions per side (Fig.
<figureCitation id="297A8D2EF057B54CA79F88D3878CBBA8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">15A</figureCitation>
). Surstylus separated from epandrium, narrowed dorsally, with
<emphasis id="0C915680DD837969FF405571F16C270D" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
14 long peg-shaped prensisetae near anterodorsal margin on inner surface (Fig.
<figureCitation id="4BE2C2222F54A0A60957F4D92548E4F1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">15B</figureCitation>
). Pregonites fused basally, with
<emphasis id="7419671218D56ED3FBFE3ABCC77D9907" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
3 sensilla distally per side (Fig.
<figureCitation id="896A8220BC1F11403768382AB9A5C752" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">15C, D</figureCitation>
). Aedeagus pubescent distally, with some hairs ventrally and apically (Fig.
<figureCitation id="BEC9D645974B1046AB119226FE460916" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Stegana (Orthostegana) fuscofemorata Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure15" httpUri="https://binary.pensoft.net/fig/864482" pageId="0" pageNumber="581">15C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="88DF51EA19F5DAE16F024B44F55BE7C5" pageId="0" pageNumber="581" type="description">
<paragraph id="33837A2BD08A3D7CF53EC7FDB12FE69D" pageId="0" pageNumber="581">Measurements and indices.</paragraph>
<paragraph id="4D12F45838F7DE40E7B460A7ED1DE0EF" pageId="0" pageNumber="581">
<emphasis id="AEFA854A82580A863F3487300224E852" bold="true" pageId="0" pageNumber="581">BL</emphasis>
= 3.20 mm in holotype (range in 4♂ and 3♀ paratypes: 3.00-3.13 in ♂, 3.00-3.10 in ♀),
<emphasis id="A73785442FD13A44605362140F693B31" bold="true" pageId="0" pageNumber="581">ThL</emphasis>
= 1.39 mm (1.27-1.30 in ♂, 1.20-1.27 in ♀),
<emphasis id="CAB81680D4B2AF8D1336C12C6F04C17A" bold="true" pageId="0" pageNumber="581">WL</emphasis>
= 2.20 mm (2.17-2.30 in ♂, 2.30-2.53 in ♀),
<emphasis id="44CC63A3E680818AFBA97C4379C9BA58" bold="true" pageId="0" pageNumber="581">WW</emphasis>
= 1.03 mm (1.03-1.07 in ♂, 1.03-1.23 in ♀),
<emphasis id="0C24CAA0EF8D336DCF0E9EA58EE4EB44" bold="true" pageId="0" pageNumber="581">arb</emphasis>
= 7/4 (6-7/4-5),
<emphasis id="42011FE0433FB153E5EB3E034E68ADD9" bold="true" pageId="0" pageNumber="581">avd</emphasis>
= 0.79 (0.80-0.85),
<emphasis id="249024B6B32C2E21EF417E5EF842AF2E" bold="true" pageId="0" pageNumber="581">adf</emphasis>
= 2.00 (1.71-2.17),
<emphasis id="69A7E35AAA99E6B27BE80FDFEB1810A9" bold="true" pageId="0" pageNumber="581">flw</emphasis>
= 2.14 (1.17-2.33),
<emphasis id="8B46E20F84254003D08866D55118DF89" bold="true" pageId="0" pageNumber="581">FW/HW</emphasis>
= 0.40 (0.35-0.40),
<emphasis id="8BA20623E3960F1BED0B092622B585A2" bold="true" pageId="0" pageNumber="581">ch/o</emphasis>
= 0.13 (0.13),
<emphasis id="FA9C8EEA5C0B1AE4008B8FF8C9D3EB03" bold="true" pageId="0" pageNumber="581">prorb</emphasis>
= 0.95 (0.81-1.05),
<emphasis id="92DEC7EDB11D113F91186770D9EF7D80" bold="true" pageId="0" pageNumber="581">rcorb</emphasis>
= 0.71 (0.68-0.75),
<emphasis id="990DF0E18FB039F176E4E9B5842E3855" bold="true" pageId="0" pageNumber="581">vb</emphasis>
= 0.67 (0.40-0.67),
<emphasis id="43C7612CB1682B436AAB2D98A876136E" bold="true" pageId="0" pageNumber="581">dcl</emphasis>
= 0.48 (0.40-0.50),
<emphasis id="D7531C03FC1BA5724F60C499A6EEF6F5" bold="true" pageId="0" pageNumber="581">presctl</emphasis>
= 0.48 (0.44-0.53),
<emphasis id="CC48E16D54A983438F81AF255F8C6D31" bold="true" pageId="0" pageNumber="581">sctl</emphasis>
= 1.19 (1.31-1.48),
<emphasis id="4C8747AA8B155F5BDE8C01481F8CD1EB" bold="true" pageId="0" pageNumber="581">sterno</emphasis>
= 0.78 (0.62-0.81),
<emphasis id="8E4D8FE0FBCB272F6F840D2896FBABD6" bold="true" pageId="0" pageNumber="581">orbito</emphasis>
= 2.00 (2.00-2.25),
<emphasis id="1945B7F8406255AF397D6C0352B431AD" bold="true" pageId="0" pageNumber="581">dcp</emphasis>
= 0.24 (0.23-0.26),
<emphasis id="CDCAC316905BF8F83397E994A16CBA06" bold="true" pageId="0" pageNumber="581">sctlp</emphasis>
= 0.92 (0.91-1.10),
<emphasis id="9AD451164098F7B989E5290B3CB6FA18" bold="true" pageId="0" pageNumber="581">C</emphasis>
= 2.22 (2.15-2.41),
<emphasis id="2B3E64F426954414AD2AD5FF892484E2" bold="true" pageId="0" pageNumber="581">4c</emphasis>
= 0.95 (0.81-0.95),
<emphasis id="774FA91A82A0CB1DC7645707C2D3F17A" bold="true" pageId="0" pageNumber="581">4v</emphasis>
= 1.68 (1.63-1.65),
<emphasis id="13C997DA7A94F9568F8F42E3F6276C76" bold="true" pageId="0" pageNumber="581">5x</emphasis>
= 1.14 (1.13-1.14),
<emphasis id="4BD9ADAE1CD79DB3AF214C5B36703DAD" bold="true" pageId="0" pageNumber="581">ac</emphasis>
= 12.00 (10.00-12.00),
<emphasis id="8FB831F959A3CE15EC18F4491EBDF8C4" bold="true" pageId="0" pageNumber="581">M</emphasis>
= 0.42 (0.38-0.43),
<emphasis id="85CC325E5ED3313E12AA146D4B2BA44C" bold="true" pageId="0" pageNumber="581">C3F</emphasis>
= 0.72 (0.72-0.83).
</paragraph>
</subSubSection>
<subSubSection id="189C03E848BBBBAF29F140F22358205E" pageId="0" pageNumber="581" type="distribution">
<paragraph id="2FF3FCA528D07DF7D71111E400DB7AF9" pageId="0" pageNumber="581">Distribution.</paragraph>
<paragraph id="5FE71F61F9199C5BF2C6A0E676E5AD95" pageId="0" pageNumber="581">China (Yunnan).</paragraph>
</subSubSection>
<subSubSection id="BCC522A731B9E8E73D51EADD426B74B1" pageId="0" pageNumber="581" type="type material">
<paragraph id="2F01933A5C01E24DCD295086347A8F3E" pageId="0" pageNumber="581">Type material examined.</paragraph>
<paragraph id="16A6D1A32CE00B194CF8240A3A9EB941" pageId="0" pageNumber="581">
<emphasis id="C9F4518E66A2A6325DBB0D0A836E220A" bold="true" pageId="0" pageNumber="581">Holotype</emphasis>
: ♂ (SCAU, No. 112117), China: Muyiji Park, Ximeng, Yunnan,
<geoCoordinate id="C04006A2691E5B8E5CD5AD3D4062F2A8" degrees="22" direction="north" minutes="37" orientation="latitude" precision="15" seconds="15" value="22.620834">22°3715″N</geoCoordinate>
,
<geoCoordinate id="DAA693D8DE97DD2450619CFAC183BB54" degrees="99" direction="east" minutes="35" orientation="longitude" precision="15" seconds="42" value="99.595">99°3542″E</geoCoordinate>
, altitude 1203 m, 29.iv.2016, swept from tree trunk, J. Huang.
<emphasis id="7D89351CD71F8A02881A2F736ACEBF70" bold="true" pageId="0" pageNumber="581">Paratypes</emphasis>
: 1♂ (SCAU, No. 112118), Y.L. Wang, other data same as the holotype; 3♂, 1♀ (SCAU, Nos. 112119-22), Menglun, Mengla, Yunnan,
<geoCoordinate id="7E4AF4797D11C2BDB0D26DF7B00E519F" degrees="21" direction="north" minutes="55" orientation="latitude" precision="15" seconds="11" value="21.919722">21°5511″N</geoCoordinate>
, 101°1640″ E, altitudes 800-900 m, 13.ix.2002, swept from tree trunks, H.W. Chen; 1♀ (SCAU, No. 112123), Baihualing, Baoshan, Yunnan,
<geoCoordinate id="A1546FF0E4CF88440D9C6157EE6A9687" degrees="25" direction="north" minutes="17" orientation="latitude" precision="15" seconds="53" value="25.298056">25°1753″N</geoCoordinate>
,
<geoCoordinate id="8469A6A8B2C9ADD64F4A45ED8540E6A2" degrees="98" direction="east" minutes="48" orientation="longitude" precision="15" seconds="09" value="98.802505">98°4809″E</geoCoordinate>
, altitude 1370 m, 15.vi.2011, swept from tree trunk, J.J. Gao; 1♀ (SCAU, No. 112124), Wangtianshu, Mengla, Yunnan,
<geoCoordinate id="41F685C044762E562045FDA5F5E5FE30" degrees="21" direction="north" minutes="37" orientation="latitude" precision="15" seconds="10" value="21.619444">21°3710″N</geoCoordinate>
,
<geoCoordinate id="D380F066B022384206AA2761504C3CA3" degrees="101" direction="east" minutes="35" orientation="longitude" precision="15" seconds="17" value="101.58806">101°3517″E</geoCoordinate>
, altitude 570 m, 11.ix.2002, swept from tree trunk, H.W. Chen.
</paragraph>
</subSubSection>
<subSubSection id="9B0E44B63D192ECD5FB0B3C2C63A5863" pageId="0" pageNumber="581" type="etymology">
<paragraph id="34E22049ECACBDC571D287BE411B9298" pageId="0" pageNumber="581">Etymology.</paragraph>
<paragraph id="1EEA7A9CACD9FBFEA838137F81B58F28" pageId="0" pageNumber="581">
A combination of the Latin words &quot;
<emphasis id="FA68529839DF91D1F153B97D0C10624B" italics="true" pageId="0" pageNumber="581">fuscus</emphasis>
&quot; (= brown) and &quot;
<emphasis id="4C269E989904E56D316F6FE1A72B9BD8" italics="true" pageId="0" pageNumber="581">femoralis</emphasis>
&quot; (= femora), referring to all femora distally brown in females.
</paragraph>
</subSubSection>
</treatment>
</document>