285 lines
28 KiB
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285 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.981.53583" ID-GBIF-Dataset="8e14a5b5-aa20-4050-b2b4-f393764fc6c7" ID-Pensoft-Pub="1313-2970-981-1" ID-Pensoft-UUID="7C44C797C4125A71BAE032A55E6FA5DC" ID-ZooBank="5194AAC86B8A473F8A41470A60182A0B" ModsDocID="1313-2970-981-1" checkinTime="1604011718171" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Sajan, Sheikh, Tripathy, Basudev, Meng, Kaibaryer, Asami, Takahiro & Ablett, Jonathan D." docDate="2020" docId="584E5BE39FB15B909DD27F27B311041E" docLanguage="en" docName="ZooKeys 981: 1-220" docOrigin="ZooKeys 981" docSource="http://dx.doi.org/10.3897/zookeys.981.53583" docTitle="Metalycaeus Pilsbry 1900" docType="treatment" docVersion="5" id="7C44C797C4125A71BAE032A55E6FA5DC" lastPageNumber="1" masterDocId="7C44C797C4125A71BAE032A55E6FA5DC" masterDocTitle="Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue" masterLastPageNumber="220" masterPageNumber="1" pageNumber="1" updateTime="1634714035540" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sajan, Sheikh</mods:namePart>
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<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India & Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
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<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Meng, Kaibaryer</mods:namePart>
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<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Asami, Takahiro</mods:namePart>
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<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5706-0272</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
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<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>981</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>220</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.981.53583</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.981.53583</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-981-1</mods:identifier>
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<mods:identifier type="ZooBank">5194AAC86B8A473F8A41470A60182A0B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">7C44C797C4125A71BAE032A55E6FA5DC</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168715083" LSID="urn:lsid:plazi:treatment:584E5BE39FB15B909DD27F27B311041E" httpUri="http://treatment.plazi.org/id/584E5BE39FB15B909DD27F27B311041E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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Genus
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<taxonomicName LSID="584E5BE3-9FB1-5B90-9DD2-7F27B311041E" authority="Pilsbry, 1900" authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Metalycaeus Pilsbry, 1900</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Metalycaeus)" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Metalycaeus">Alycaeus (Metalycaeus)</taxonomicName>
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Pilsbry, 1900: 382. (section of
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<taxonomicName authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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)
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Raptomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Raptomphalus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Raptomphalus</taxonomicName>
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Godwin-Austen, 1914: 366. syn. nov.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Metalycaeus</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.82528" author="Bollinger, G" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="1" pagination="309 - 340" refId="B30" refString="Bollinger, G, 1918. Land-Mollusken von Celebes. Ausbeute der in den Jahren 1902 und 1903 ausgefuehrten zweiten Celebes-Reise der Herren Dr. P. und Dr. F. Sarasin. Revue Suisse de Zoologie 26: 309 - 340, DOI: https://doi.org/10.5962/bhl.part.82528" title="Land-Mollusken von Celebes. Ausbeute der in den Jahren 1902 und 1903 ausgefuehrten zweiten Celebes-Reise der Herren Dr. P. und Dr. F. Sarasin." url="https://doi.org/10.5962/bhl.part.82528" volume="26" year="1918">Bollinger 1918</bibRefCitation>
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: 316;
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Venus" pageId="0" pageNumber="1" pagination="1 - 16" refId="B157" refString="Pall-Gergely, B, Asami, T, 2017. A new species of Metalycaeus hitherto undistinguished from M. vinctus (Pilsbry, 1902). Venus 75 (1-4): 1 - 16" title="A new species of Metalycaeus hitherto undistinguished from M. vinctus (Pilsbry, 1902)." volume="75" year="2017">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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and Asami 2017
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</bibRefCitation>
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: 4;
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2017
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</bibRefCitation>
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: 73.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus (Metalycaeus)" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Metalycaeus">Chamalycaeus (Metalycaeus)</taxonomicName>
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-
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<bibRefCitation author="Thiele, J" journalOrPublisher="Verlag von Gustav Fischer, Jena" pageId="0" pageNumber="1" refId="B213" refString="Thiele, J, 1929. Handbuch der systematischen Weichtierkunde, Teil 1. Verlag von Gustav Fischer, Jena" title="Handbuch der systematischen Weichtierkunde, Teil 1." year="1929">Thiele 1929</bibRefCitation>
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: 108; Wenz 1938: 478;
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<bibRefCitation author="Egorov, RV" journalOrPublisher="Egorov, Moscow" pageId="0" pageNumber="1" refId="B45" refString="Egorov, RV, 2013. Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R. Egorov, Moscow" title="Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R." year="2013">Egorov 2013</bibRefCitation>
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: 37.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus (Raptomphalus)" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Raptomphalus">Chamalycaeus (Raptomphalus)</taxonomicName>
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-
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<bibRefCitation author="Thiele, J" journalOrPublisher="Verlag von Gustav Fischer, Jena" pageId="0" pageNumber="1" refId="B213" refString="Thiele, J, 1929. Handbuch der systematischen Weichtierkunde, Teil 1. Verlag von Gustav Fischer, Jena" title="Handbuch der systematischen Weichtierkunde, Teil 1." year="1929">Thiele 1929</bibRefCitation>
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: 108; Wenz 1938: 478;
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<bibRefCitation author="Egorov, RV" journalOrPublisher="Egorov, Moscow" pageId="0" pageNumber="1" refId="B45" refString="Egorov, RV, 2013. Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R. Egorov, Moscow" title="Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R." year="2013">Egorov 2013</bibRefCitation>
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: 37-38.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type species">
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<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Metalycaeus) melanopoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Melanopoma" subGenus="Metalycaeus">Alycaeus (Metalycaeus) melanopoma</taxonomicName>
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Pilsbry, 1900 (Fig.
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<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Type species of alycaeid genus-group taxa A Alycaeus melanopoma Pilsbry, 1900 (synonym of Chamalycaeus nipponensis (Reinhardt, 1877 )), lectotype (ANSP 78815; type species of Metalycaeus); B Metalycaeus magnificus (Godwin-Austen, 1914), syntype (NHMUK 1903.7.1.3115; type species of Raptomphalus). Photographs: downloaded from the website of ANSP (A), Harold Taylor (B)." figureDoi="10.3897/zookeys.981.53583.figure31" httpUri="https://binary.pensoft.net/fig/468676" pageId="0" pageNumber="1">31A</figureCitation>
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) (junior synonym of
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<taxonomicName baseAuthorityName="Reinhardt" baseAuthorityYear="1877" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus nipponensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Nipponensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus nipponensis</emphasis>
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</taxonomicName>
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[Reinhardt, 1877], see
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<bibRefCitation author="Minato, H" journalOrPublisher="Minato, Shirahama X" pageId="0" pageNumber="1" refId="B124" refString="Minato, H, 1988. A systematic and bibliographic list of the Japanese land snails. H. Minato, Shirahama X" title="A systematic and bibliographic list of the Japanese land snails. H." year="1988">Minato 1988</bibRefCitation>
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), SD
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<bibRefCitation author="Thiele, J" journalOrPublisher="Verlag von Gustav Fischer, Jena" pageId="0" pageNumber="1" refId="B213" refString="Thiele, J, 1929. Handbuch der systematischen Weichtierkunde, Teil 1. Verlag von Gustav Fischer, Jena" title="Handbuch der systematischen Weichtierkunde, Teil 1." year="1929">Thiele (1929</bibRefCitation>
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: 108).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell small to very large (D: 3-10 mm), spire usually low, protoconch usually elevated, spirally striated (key character); spiral striation of protoconch very rarely absent (see under
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<taxonomicName lsidName="M. laevis" pageId="0" pageNumber="1" rank="species" species="Laevis">
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<emphasis italics="true" pageId="0" pageNumber="1">M. laevis</emphasis>
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</taxonomicName>
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); R1 usually reticulated with sometimes prominent radial ribs; spiral lines always present on the teleoconch; R2 rarely short, usually long to very long, sometimes entirely smooth, but typically with widely spaced, straight, sharp ribs; R3 usually well-developed, although can be reduced. Operculum thin or relatively thickened, sometimes with funnel-shaped outer surface caused by the modification of the multispiral outer laminae. Radula usual for the family (central tooth with five cusps, broad, central cusp pointed).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
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</taxonomicName>
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is recognised based on the spirally striated protoconch, which distinguishes
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<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
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</taxonomicName>
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from species assigned to all other alycaeid genera.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
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</taxonomicName>
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is widely distributed from the southeastern Himalaya to the south of Honshu Island (Japan) and the northern islands of the Philippines, but it does not extend any further south than northern Laos and Vietnam (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of Chamalycaeus Moellendorff, 1897 (dark shaded area) and Metalycaeus Pilsbry, 1900 (light shaded area)." figureDoi="10.3897/zookeys.981.53583.figure9" httpUri="https://binary.pensoft.net/fig/468654" pageId="0" pageNumber="1">9</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The
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<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
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</taxonomicName>
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records in
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<bibRefCitation DOI="https://doi.org/10.1080/13235818.2019.1638541" author="Pall-Gergely, B" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="377 - 389" refId="B159" refString="Pall-Gergely, B, Auffenberg, K, 2019. A review of the Alycaeidae of the Philippines with descriptions of new species and subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea). Molluscan Research 39 (4): 377 - 389, DOI: https://doi.org/10.1080/13235818.2019.1638541" title="A review of the Alycaeidae of the Philippines with descriptions of new species and subspecies (Gastropoda: Caenogastropoda: Cyclophoroidea)." url="https://doi.org/10.1080/13235818.2019.1638541" volume="39" year="2019">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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and Auffenberg (2019)
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</bibRefCitation>
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from Borneo (
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<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus everetti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Everetti">
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<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus everetti</emphasis>
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</taxonomicName>
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) and Sumatra (
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<taxonomicName baseAuthorityName="E. von Martens" baseAuthorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus sumatranus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Sumatranus">
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<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus sumatranus</emphasis>
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</taxonomicName>
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) later proved to be
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<taxonomicName authorityName="Moellendorff" authorityYear="1897" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus</emphasis>
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</taxonomicName>
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species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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|
The type species of
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Raptomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Raptomphalus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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|
<emphasis italics="true" pageId="0" pageNumber="1">Raptomphalus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus magnificus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Magnificus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus magnificus</emphasis>
|
|
</taxonomicName>
|
|
) (Fig.
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|
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Type species of alycaeid genus-group taxa A Alycaeus melanopoma Pilsbry, 1900 (synonym of Chamalycaeus nipponensis (Reinhardt, 1877 )), lectotype (ANSP 78815; type species of Metalycaeus); B Metalycaeus magnificus (Godwin-Austen, 1914), syntype (NHMUK 1903.7.1.3115; type species of Raptomphalus). Photographs: downloaded from the website of ANSP (A), Harold Taylor (B)." figureDoi="10.3897/zookeys.981.53583.figure31" httpUri="https://binary.pensoft.net/fig/468676" pageId="0" pageNumber="1">31B</figureCitation>
|
|
) and
|
|
<taxonomicName lsidName="M. oakesi" pageId="0" pageNumber="1" rank="species" species="Oakesi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">M. oakesi</emphasis>
|
|
</taxonomicName>
|
|
have a prominently keeled umbilicus which serve as the distinctive character for the genus
|
|
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Raptomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Raptomphalus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Raptomphalus</emphasis>
|
|
</taxonomicName>
|
|
. A less prominently keeled umbilicus is observable in other species, such as
|
|
<taxonomicName baseAuthorityName="W. T. Blanford" baseAuthorityYear="1863" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus vulcani" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Vulcani">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus vulcani</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1886" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus brahma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Brahma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus brahma</emphasis>
|
|
</taxonomicName>
|
|
, which can be interpreted as intermediate forms between the non-keeled umbilicus of alycaeids and the keeled umbilicus of
|
|
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Raptomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Raptomphalus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Raptomphalus</emphasis>
|
|
</taxonomicName>
|
|
. Moreover, a variety of
|
|
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1886" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus brahma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Brahma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus brahma</emphasis>
|
|
</taxonomicName>
|
|
has much less prominent umbilical keel than typical species in the genus. Therefore, the keeled umbilicus cannot serve as a distinctive character between genera. Consequently,
|
|
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Raptomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Raptomphalus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Raptomphalus</emphasis>
|
|
</taxonomicName>
|
|
is a synonym of
|
|
<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">For sake of simplicity, this genus is divided here into typical and atypical (or questionable) species. The first includes those having sharp, widely spaced R2 ribs, whereas the second includes species with R2 sculpture other than typical, including those which could not be examined.</paragraph>
|
|
<caption doi="10.3897/zookeys.981.53583.figure31" httpUri="https://binary.pensoft.net/fig/468676" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
|
|
Type species of alycaeid genus-group taxa
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus melanopoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Melanopoma">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus melanopoma</emphasis>
|
|
</taxonomicName>
|
|
Pilsbry, 1900 (synonym of
|
|
<taxonomicName baseAuthorityName="Reinhardt" baseAuthorityYear="1877" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus nipponensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Nipponensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus nipponensis</emphasis>
|
|
</taxonomicName>
|
|
(Reinhardt, 1877)), lectotype (ANSP 78815; type species of
|
|
<taxonomicName authorityName="Pilsbry" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus</emphasis>
|
|
</taxonomicName>
|
|
);
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus magnificus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Magnificus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Metalycaeus magnificus</emphasis>
|
|
</taxonomicName>
|
|
(Godwin-Austen, 1914), syntype (NHMUK 1903.7.1.3115; type species of
|
|
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Raptomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Raptomphalus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Raptomphalus</emphasis>
|
|
</taxonomicName>
|
|
). Photographs: downloaded from the website of ANSP (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), Harold Taylor (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |