224 lines
23 KiB
XML
224 lines
23 KiB
XML
<document id="CEF389554C464B05F9A201D064FFE137" ID-DOI="10.1163/18759866-BJA10013" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343289" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1694663361057" checkinUser="felipe" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Tongkerd, Piyoros, Jeratthitikul, Ekgachai & Panha, Somsak" docDate="2021" docId="426CF52FA4534B6DFF1B81586759BD98" docLanguage="en" docName="ContribZool.90.1.21-69.pdf" docOrigin="Contributions to Zoology 90 (1)" docSource="http://dx.doi.org/10.1163/18759866-bja10013" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Taphrenalla zemia Pholyotha & Sutcharit & Tongkerd & Jeratthitikul & Panha 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="67" masterDocId="BE558D57A47D4B43FF9A80386371BE43" masterDocTitle="Integrative systematics reveals the new land-snail genus Taphrenalla (Eupulmonata: Ariophantidae) with a description of nine new species from Thailand" masterLastPageNumber="69" masterPageNumber="21" pageNumber="67" updateTime="1695043390670" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="330C2CE5AC191367D66FE79A856B2445" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A06F187826DE71A513DB7BBD93630C90">
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<mods:title id="56BDD64F616701D85E7C16A664AA3110">Integrative systematics reveals the new land-snail genus Taphrenalla (Eupulmonata: Ariophantidae) with a description of nine new species from Thailand</mods:title>
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<mods:name id="5C270450B532E8ACC4694ADACFD73DB2" type="personal">
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<mods:roleTerm id="55A25937522966F26A773D232DC041B8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="956B4FF97152DA2972CD8FED0D3BCCAF">Pholyotha, Arthit</mods:namePart>
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<mods:affiliation id="6979594126E2C5B0EA5DA00744382972">Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn</mods:affiliation>
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</mods:name>
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<mods:name id="A8FE672788F9B46CE2A0E6EA79A39DC4" type="personal">
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<mods:roleTerm id="2C67A1828BC9699FB2844C598879B7F0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8827FBDDA470ACBFE73904B4FFA466B3">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="E4CF6E0389E8DB5CF085D1819DE54D2D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="1B2DCE9B97F299FE3A875D29F088489E" type="personal">
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<mods:roleTerm id="10214F9D1B04EB5454D6B3755463F472">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F03F1FB984D8C56A8E177DC1F6942B86">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="3A3BAF60E88E43C68B90AA86B99BD811">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="F398ADEA5182A01788AF5C624A1A3476" type="personal">
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<mods:role id="A369A47819093E0402B1EC6D61DB3559">
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<mods:roleTerm id="7FFC9D24B4F2969033628215E1D5AE8E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E9458603FF4E78C1063FADBBB9C379B7">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="1CD0383BF432233320F0F7FA732C5501">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10330, Bangkok, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="0A76189F065EA9C4F668F8A85DDCD3A6" type="personal">
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<mods:role id="3328BC8C9808B188C22111F14C35851B">
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<mods:roleTerm id="8570DB06D92FCCA2B7455D1B85D36B22">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3D9C1743492EA6BCA5B5AFE4176EF212">Panha, Somsak</mods:namePart>
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<mods:affiliation id="E998DF325D86C237D53FE277EEF4E217">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:nameIdentifier id="D2FCADA45C05BD50DD7CA769433D2B47" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource id="3BCD7D1A4AC9F5CECF1A584A9EE901B8">text</mods:typeOfResource>
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<mods:title id="0B6C227EED6037BB4ACB81CE3903DB96">Contributions to Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="74340F59AB07672598FFE016D638D71D">
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<mods:date id="96250E09E5806529DD1A4B577F701D0A">2021</mods:date>
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<mods:number id="1CC87D8E268F14F2DB58794978B95D24">2020-12-07</mods:number>
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<mods:number id="972A3D6743DEE9868CFF0F54F0FA79F3">90</mods:number>
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<mods:detail id="413134428BEFE853E78D76D0822FE863" type="issue">
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<mods:number id="065B849AA07AEC065F1FA76F4C442ECC">1</mods:number>
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<mods:start id="9794F6B3437EFA582973A1C1A471E7B7">21</mods:start>
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<mods:url id="2A84EF0D881B7DC06471DD981B4420F7">http://dx.doi.org/10.1163/18759866-bja10013</mods:url>
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<mods:classification id="38CE263C54F381F9065E6B6E5E63DC32">journal article</mods:classification>
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<mods:identifier id="2A47F5E816B98FDD45E4E6DD80851C90" type="DOI">10.1163/18759866-BJA10013</mods:identifier>
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<mods:identifier id="691C7357BBF6F8BE69641BE98847E297" type="ISSN">1875-9866</mods:identifier>
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<mods:identifier id="2641C328C4B924CBBC0232099737B16E" type="Zenodo-Dep">8343289</mods:identifier>
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<treatment id="426CF52FA4534B6DFF1B81586759BD98" LSID="urn:lsid:plazi:treatment:426CF52FA4534B6DFF1B81586759BD98" httpUri="http://treatment.plazi.org/id/426CF52FA4534B6DFF1B81586759BD98" lastPageNumber="67" pageId="46" pageNumber="67">
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<subSubSection id="82DF17B2A4534B6DFF1B81586112BF3B" box="[129,611,352,377]" pageId="46" pageNumber="67" type="nomenclature">
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<paragraph id="CA7A4439A4534B6DFF1B81586112BF3B" blockId="46.[129,611,352,377]" box="[129,611,352,377]" pageId="46" pageNumber="67">
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<heading id="9132F355A4534B6DFF1B81586112BF3B" box="[129,611,352,377]" fontSize="10" level="2" pageId="46" pageNumber="67" reason="6">
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<taxonomicName id="0DC53FBAA4534B6DFF1B8158617FBF3A" ID-CoL="8L579" ID-ENA="2729321" authority="Pholyotha & Panha" authorityName="Pholyotha & Sutcharit & Tongkerd & Jeratthitikul & Panha" authorityYear="2021" box="[129,526,352,377]" class="Gastropoda" family="Ariophantidae" genus="Taphrenalla" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="67" phylum="Mollusca" rank="species" species="zemia" status="sp. nov.">
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<emphasis id="F8B1982BA4534B6DFF1B81586232BF3A" bold="true" box="[129,323,352,377]" italics="true" pageId="46" pageNumber="67">Taphrenalla zemia</emphasis>
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Pholyotha & Panha
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</taxonomicName>
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,
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<taxonomicNameLabel id="E3822550A4534B6DFD8281596112BF3B" box="[536,611,353,376]" pageId="46" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="82DF17B2A4534B6DFF1B81BD62F0BFDE" box="[129,385,389,413]" pageId="46" pageNumber="67" type="description">
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<paragraph id="CA7A4439A4534B6DFF1B81BD62F0BFDE" blockId="46.[129,385,389,413]" box="[129,385,389,413]" pageId="46" pageNumber="67">
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(
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<figureCitation id="52FE58BCA4534B6DFF1D81BD63CBBFDE" box="[135,186,389,413]" captionStart="Figure 1" captionStartId="3.[98,166,1062,1084]" captionTargetBox="[102,1153,164,1043]" captionTargetId="figure-208@3.[102,1153,164,1043]" captionTargetPageId="3" captionText="Figure 1 Map showing the sampling sites of the 11 Taphrenalla species along Southern Thailand.Each colour of the distribution range follows the phylogenetic tree of all Taphrenalla species in fig.3." figureDoi="http://doi.org/10.5281/zenodo.8343291" httpUri="https://zenodo.org/record/8343291/files/figure.png" pageId="46" pageNumber="67">figs 1</figureCitation>
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,
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||
<figureCitation id="52FE58BCA4534B6DFF5E81BD63A1BFDE" box="[196,208,389,413]" captionStart="Figure 3" captionStartId="13.[98,166,1547,1569]" captionTargetBox="[103,1152,168,1525]" captionTargetId="figure-2@13.[99,1155,164,1528]" captionTargetPageId="13" captionText="Figure 3 A representative ML tree showing only the Taphrenalla clade.The synopsis of the radial groove on shell sculpture, genitalia showing location of dart apparatus, and live specimens of Taphrenalla spp. Results of species delimitation analyses based on the COI gene using ABGD and PTP.Black circles represent nodes with both ML (when Ē 70%) and BI (when Ē 0.95) support.White circles represent nodes with ML BS support but no BI PP support (when <0.95). Each clade colour refers to the distribution range in fig. 1." figureDoi="http://doi.org/10.5281/zenodo.8343295" httpUri="https://zenodo.org/record/8343295/files/figure.png" pageId="46" pageNumber="67">3</figureCitation>
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||
, 13B, 14C, table 6)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="82DF17B2A4534B6DFF1B81F5612FBDF4" pageId="46" pageNumber="67" type="materials_examined">
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<paragraph id="CA7A4439A4534B6DFF1B81F562ACBC13" blockId="46.[129,641,460,592]" pageId="46" pageNumber="67">
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<materialsCitation id="7AAD4E64A4534B6DFF1B81F563C0BC4A" collectionCode="CUMZ" country="India" pageId="46" pageNumber="67" typeStatus="Type">
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<emphasis id="F8B1982BA4534B6DFF1B81F56207BFA7" box="[129,374,460,484]" italics="true" pageId="46" pageNumber="67">Type material examined.</emphasis>
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<typeStatus id="157EFA9BA4534B6DFEE081F462AABFA7" box="[378,475,460,484]" pageId="46" pageNumber="67">Holotype</typeStatus>
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||
:
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<collectionCode id="ACD4DCFCA4534B6DFE7881F56157BFA6" box="[482,550,461,485]" collectionName="United Kingdom, Cambridge, University, Museum of Zoology" pageId="46" pageNumber="67">CUMZ</collectionCode>
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7213 (fig. 13B).
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</materialsCitation>
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<materialsCitation id="7AAD4E64A4534B6DFF2E81C96151BC6F" collectionCode="CUMZ" country="India" pageId="46" pageNumber="67" typeStatus="Paratypes">
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<typeStatus id="157EFA9BA4534B6DFF2E81C96268BC4A" box="[180,281,497,521]" pageId="46" pageNumber="67">Paratypes</typeStatus>
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||
. Same data as holotype:
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||
<collectionCode id="ACD4DCFCA4534B6DFD9781C96120BC4A" box="[525,593,497,521]" collectionName="United Kingdom, Cambridge, University, Museum of Zoology" pageId="46" pageNumber="67">CUMZ</collectionCode>
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||
7214 (25 shells and
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||
<specimenCount id="DCC38FB0A4534B6DFE8C822C62D0BC6F" box="[278,417,532,556]" pageId="46" pageNumber="67" type="generic">30 specimens</specimenCount>
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in ethanol),
|
||
</materialsCitation>
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||
<materialsCitation id="7AAD4E64A4534B6DFDBD822D638DBC13" collectionCode="NHMUK" country="India" pageId="46" pageNumber="67">
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||
<collectionCode id="ACD4DCFCA4534B6DFDBD822D610EBC6F" box="[551,639,533,556]" collectionName="NHMUK" pageId="46" pageNumber="67">NHMUK</collectionCode>
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||
(two shells),
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||
</materialsCitation>
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||
<materialsCitation id="7AAD4E64A4534B6DFE98820062A8BC13" box="[258,473,568,592]" collectionCode="SMF" country="India" pageId="46" pageNumber="67">
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||
and
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||
<collectionCode id="ACD4DCFCA4534B6DFEB48200622DBC13" box="[302,348,568,592]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="46" pageNumber="67">SMF</collectionCode>
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||
(two shells)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="CA7A4439A4534B6DFF1B82B862EABD08" blockId="46.[129,640,640,843]" pageId="46" pageNumber="67">
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<materialsCitation id="7AAD4E64A4534B6DFF1B82B863C8BD47" collectionCode="N" country="India" latitude="8.475249" longitude="98.53911" pageId="46" pageNumber="67" stateProvince="Tham">
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||
<emphasis id="F8B1982BA4534B6DFF1B82B862F1BCDB" box="[129,384,640,664]" italics="true" pageId="46" pageNumber="67">Other material examined.</emphasis>
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||
<collectingRegion id="08018ADBA4534B6DFE1E82B862B1BCDB" box="[388,448,640,664]" pageId="46" pageNumber="67">Tham</collectingRegion>
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Pha Sua Bureau of Monks,
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<collectingRegion id="08018ADBA4534B6DFF4E829C620EBCFF" box="[212,383,676,700]" pageId="46" pageNumber="67">Mueang District</collectingRegion>
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, Phang-nga (
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<geoCoordinate id="AFF122FEA4534B6DFD92829C61F1BCFF" box="[520,640,676,700]" degrees="8" direction="north" minutes="28" orientation="latitude" pageId="46" pageNumber="67" precision="1" seconds="30.9" value="8.475249">8°28’30.9”N</geoCoordinate>
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<geoCoordinate id="AFF122FEA4534B6DFF1B82F06275BCA3" box="[129,260,712,736]" degrees="98" direction="east" minutes="32" orientation="longitude" pageId="46" pageNumber="67" precision="1" seconds="20.8" value="98.53911">98°32’20.8”E</geoCoordinate>
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):
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||
<collectionCode id="ACD4DCFCA4534B6DFE8F82F06228BCA3" box="[277,345,712,736]" collectionName="United Kingdom, Cambridge, University, Museum of Zoology" pageId="46" pageNumber="67">CUMZ</collectionCode>
|
||
7215 (
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||
<specimenCount id="DCC38FB0A4534B6DFE0282F06142BCA3" box="[408,563,712,736]" pageId="46" pageNumber="67" type="generic">five specimens</specimenCount>
|
||
in ethanol)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="7AAD4E64A4534B6DFF5082D462E6BD08" collectionCode="N" country="India" latitude="8.463305" longitude="98.569885" pageId="46" pageNumber="67" stateProvince="Wat Tham">
|
||
<collectingRegion id="08018ADBA4534B6DFF5082D4624CBD47" box="[202,317,748,772]" pageId="46" pageNumber="67">Wat Tham</collectingRegion>
|
||
Bang Toei,
|
||
<collectingRegion id="08018ADBA4534B6DFE5682D4610DBD47" box="[460,636,748,772]" pageId="46" pageNumber="67">Mueang District</collectingRegion>
|
||
, Phang-nga (
|
||
<geoCoordinate id="AFF122FEA4534B6DFF6683286203BD6B" box="[252,370,784,808]" degrees="8" direction="north" minutes="27" orientation="latitude" pageId="46" pageNumber="67" precision="1" seconds="47.9" value="8.463305">8°27’47.9”N</geoCoordinate>
|
||
<geoCoordinate id="AFF122FEA4534B6DFEE183286287BD6B" box="[379,502,784,808]" degrees="98" direction="east" minutes="34" orientation="longitude" pageId="46" pageNumber="67" precision="1" seconds="11.6" value="98.569885">98°34’11.6”E</geoCoordinate>
|
||
):
|
||
<collectionCode id="ACD4DCFCA4534B6DFD938328613CBD6B" box="[521,589,784,808]" collectionName="United Kingdom, Cambridge, University, Museum of Zoology" pageId="46" pageNumber="67">CUMZ</collectionCode>
|
||
7216 (
|
||
<specimenCount id="DCC38FB0A4534B6DFF1D830B626EBD08" box="[135,287,819,843]" pageId="46" pageNumber="67" type="generic">five specimens</specimenCount>
|
||
in ethanol)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CA7A4439A4534B6DFF1B8344612FBDF4" blockId="46.[129,640,891,951]" pageId="46" pageNumber="67">
|
||
<materialsCitation id="7AAD4E64A4534B6DFF1B8344612BBDF4" collectionCode="N" country="Thailand" latitude="8.531445" longitude="98.57816" pageId="46" pageNumber="67" typeStatus="Type">
|
||
<emphasis id="F8B1982BA4534B6DFF1B83446276BDD0" box="[129,263,891,915]" italics="true" pageId="46" pageNumber="67">
|
||
<typeStatus id="157EFA9BA4534B6DFF1B834463C1BDD0" box="[129,176,892,915]" pageId="46" pageNumber="67">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Wat Khiriwong, Thap Put District, Phang-nga,
|
||
<collectingCountry id="B2D204A9A4534B6DFF6D83A76223BDF4" box="[247,338,927,951]" name="Thailand" pageId="46" pageNumber="67" value="Thailand">Thailand</collectingCountry>
|
||
(
|
||
<geoCoordinate id="AFF122FEA4534B6DFEC483A762BEBDF4" box="[350,463,927,951]" degrees="8" direction="north" minutes="31" orientation="latitude" pageId="46" pageNumber="67" precision="1" seconds="53.2" value="8.531445">8°31’53.2”N</geoCoordinate>
|
||
<geoCoordinate id="AFF122FEA4534B6DFE4E83A76122BDF4" box="[468,595,927,951]" degrees="98" direction="east" minutes="34" orientation="longitude" pageId="46" pageNumber="67" precision="1" seconds="41.4" value="98.57816">98°34’41.4”E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82DF17B2A4534B6DFF1B83DF63A1BA28" pageId="46" pageNumber="67" type="etymology">
|
||
<paragraph id="CA7A4439A4534B6DFF1B83DF63A1BA28" blockId="46.[129,640,999,1131]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFF1B83DF639FBDBC" box="[129,238,999,1023]" italics="true" pageId="46" pageNumber="67">Etymology.</emphasis>
|
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The specific epithet is from the Greek word ‘
|
||
<taxonomicName id="0DC53FBAA4534B6DFF598434638EBA60" authorityName="Pholyotha & Sutcharit & Tongkerd & Jeratthitikul & Panha" authorityYear="2021" box="[195,255,1036,1059]" class="Gastropoda" family="Ariophantidae" genus="Taphrenalla" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="67" phylum="Mollusca" rank="species" species="zemia">
|
||
<emphasis id="F8B1982BA4534B6DFF598434638EBA60" box="[195,255,1036,1059]" italics="true" pageId="46" pageNumber="67">zemia</emphasis>
|
||
</taxonomicName>
|
||
’ meaning ‘loss’, which referrs to the shell being without radial grooves on the upper surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82DF17B2A4534B6DFF1B84A36398BB5D" pageId="46" pageNumber="67" type="diagnosis">
|
||
<paragraph id="CA7A4439A4534B6DFF1B84A36398BB5D" blockId="46.[129,641,1179,1310]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFF1B84A36398BAF1" box="[129,233,1179,1202]" italics="true" pageId="46" pageNumber="67">Diagnosis.</emphasis>
|
||
Shell moderate to large-sized, shallowly impressed suture, and without radial groove. Epiphallic caecum straight. Dart apparatus located on vagina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82DF17B2A4534B6DFF1B85776722BCFF" pageId="46" pageNumber="67" type="description">
|
||
<paragraph id="CA7A4439A4534B6DFF1B85776382BB25" blockId="46.[129,640,1359,1705]" box="[129,243,1359,1382]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFF1B85776382BB25" box="[129,243,1359,1382]" italics="true" pageId="46" pageNumber="67">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="CA7A4439A4534B6DFF1B854A6245B8EA" blockId="46.[129,640,1359,1705]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFF1B854A63C0BBC9" box="[129,177,1394,1418]" italics="true" pageId="46" pageNumber="67">Shell</emphasis>
|
||
(fig. 13B). Shell globosely depressed, moderate to nearly large (width up to
|
||
<quantity id="0D3DE9DCA4534B6DFE4085AE6143BBEE" box="[474,562,1430,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="46" pageNumber="67" unit="mm" value="21.3">21.3 mm</quantity>
|
||
; height up to
|
||
<quantity id="0D3DE9DCA4534B6DFF2485826263BB91" box="[190,274,1466,1490]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="46" pageNumber="67" unit="mm" value="11.9">11.9 mm</quantity>
|
||
) and yellowish-brown. Whorls 5–6; suture shallowly impressed. Upper shell surface smooth without radial grooves. Spire moderately elevated; apex raised with a spire angle of about 137–151°. Last whorl slightly shouldered and almost rounded periphery. Aperture lip simple; umbilicus narrowly opened.
|
||
</paragraph>
|
||
<paragraph id="CA7A4439A4534B6DFD3E809560E2BC4B" blockId="46.[676,1187,173,520]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFD3E80956045BE86" box="[676,820,173,197]" italics="true" pageId="46" pageNumber="67">Genital organs</emphasis>
|
||
(fig. 14C). Atrium long. Penis slender tube and slightly longer than vagina. Epiphallus cylindrical tube, nearly one and a half times total penis length. Epiphallic caecum short, and attached with penial retractor muscle. Flagellum large and slender tube. Vas deferens thin tube connected between distal end of epiphallus and free oviduct. Vagina long and cylindrical-shaped. Dart apparatus located near proximal vagina away from penis joining to atrium.
|
||
</paragraph>
|
||
<paragraph id="CA7A4439A4534B6DFD3E82006722BCFF" blockId="46.[676,1187,568,700]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFD3E8200603FBC13" box="[676,846,568,592]" italics="true" pageId="46" pageNumber="67">External features</emphasis>
|
||
(
|
||
<figureCitation id="52FE58BCA4534B6DFCC0820060FCBC13" box="[858,909,568,592]" captionStart="Figure 3" captionStartId="13.[98,166,1547,1569]" captionTargetBox="[103,1152,168,1525]" captionTargetId="figure-2@13.[99,1155,164,1528]" captionTargetPageId="13" captionText="Figure 3 A representative ML tree showing only the Taphrenalla clade.The synopsis of the radial groove on shell sculpture, genitalia showing location of dart apparatus, and live specimens of Taphrenalla spp. Results of species delimitation analyses based on the COI gene using ABGD and PTP.Black circles represent nodes with both ML (when Ē 70%) and BI (when Ē 0.95) support.White circles represent nodes with ML BS support but no BI PP support (when <0.95). Each clade colour refers to the distribution range in fig. 1." figureDoi="http://doi.org/10.5281/zenodo.8343295" httpUri="https://zenodo.org/record/8343295/files/figure.png" pageId="46" pageNumber="67">fig. 3</figureCitation>
|
||
). Animal dark gray in body colour with three pale milky yellow stripes running from head to caudal horn. One stripe at middle of the body and two stripes close to foot sole.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82DF17B2A4534B6DFD3E82D36759BD98" pageId="46" pageNumber="67" type="discussion">
|
||
<paragraph id="CA7A4439A4534B6DFD3E82D36759BD98" blockId="46.[676,1188,747,987]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFD3E82D360C6BD40" box="[676,951,747,771]" italics="true" pageId="46" pageNumber="67">
|
||
Remarks.
|
||
<taxonomicName id="0DC53FBAA4534B6DFC9B82D360C6BD40" authority="Pholyotha & Sutcharit & Tongkerd & Jeratthitikul & Panha, 2021" authorityName="Pholyotha & Sutcharit & Tongkerd & Jeratthitikul & Panha" authorityYear="2021" box="[769,951,747,771]" class="Gastropoda" family="Ariophantidae" genus="Taphrenalla" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="67" phylum="Mollusca" rank="species" species="zemia" status="sp. nov.">Taphrenalla zemia</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="E3822550A4534B6DFC2182D46775BD40" box="[955,1028,748,771]" pageId="46" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is distinguished from all other
|
||
<taxonomicName id="0DC53FBAA4534B6DFCB18337609DBD64" box="[811,1004,783,807]" pageId="46" pageNumber="67">
|
||
<emphasis id="F8B1982BA4534B6DFCB1833760D0BD64" box="[811,929,783,807]" italics="true" pageId="46" pageNumber="67">Taphrenalla</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the shell having a shallowly impressed suture without a subsutural band and no radial grooves on shell sculpture. Although we surveyed three localities during the wet season, only juvenile specimens were found for the examination and illustrated herein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |