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<document id="F3B9E0F43C1E977AE63A91EDCF39B17D" ID-DOI="10.5281/zenodo.5393502" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393502" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630318937109" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques &amp; Mauriès, Jean-Paul" docDate="2006" docId="03FEF24BA97A6A5DFCB4FC3A1DD3C4A7" docLanguage="en" docName="Zoosystema.2006.28.1.75-92.pdf" docOrigin="Zoosystema 28 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Lipseuma bernardi Golovatch &amp; Geoffroy &amp; Mauriès 2006, n. sp." docType="treatment" docVersion="4" lastPageNumber="91" masterDocId="FFC78A33A9756A4DFF9CFFAB1F53C162" masterDocTitle="Four new Chordeumatida (Diplopoda) from caves in China" masterLastPageNumber="92" masterPageNumber="75" pageNumber="90" updateTime="1699225294555" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1DC064A840F9BBCBD5A0FCD1A6358373" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0906F8B9F78F46C39906EB0128ECC5C2">
<mods:title id="B89CC3F47F4F53DCECA2FC88963A53AB">Four new Chordeumatida (Diplopoda) from caves in China</mods:title>
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<mods:name id="E977F43460E5A0A5D26239B7880C931D" type="personal">
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<mods:namePart id="581894D95B4B27F5CD83D9E7E231F369">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="DB69D8FD79DEC2F59B510DA3857F3C83">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:namePart id="83493102D3DF9DB1ECEDC91A8CEFE104">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="B6678A8EFB11DEDA38F7FDEFEC5C9005">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du Sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="EA8185810A41ED746AA12913CF9F42AB" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="4D7F5ECB5CAD27644913629DC2D60E01">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="4EFD0CC01E89D9D61ACF5C98A1B1501E">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602 Taxonomie et Collections, case postale 53, 61 rue Buffon, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:typeOfResource id="0CDF1177F9AEF91FE7EE5D60B15F06A8">text</mods:typeOfResource>
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<mods:title id="54AE06ADBD4C03599A6C45B5BA414039">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="A0E379DB86CC66F0A8AF15DC7865F439">
<mods:date id="A93DBD191D62D2790F8CA138A6A69623">2006</mods:date>
<mods:detail id="FF8499A75B4D193E4248C365A47EE636" type="volume">
<mods:number id="D127608994457FFC1F9B9EC17FF348BC">28</mods:number>
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<mods:identifier id="F4E5F689D217300FAE3EED5E88D9BD71" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03FEF24BA97A6A5DFCB4FC3A1DD3C4A7" ID-DOI="http://doi.org/10.5281/zenodo.5469453" ID-Zenodo-Dep="5469453" LSID="urn:lsid:plazi:treatment:03FEF24BA97A6A5DFCB4FC3A1DD3C4A7" httpUri="http://treatment.plazi.org/id/03FEF24BA97A6A5DFCB4FC3A1DD3C4A7" lastPageId="16" lastPageNumber="91" pageId="15" pageNumber="90">
<subSubSection id="C34D10D6A97A6A42FCB4FC3A1B64C2C9" box="[808,1079,912,939]" pageId="15" pageNumber="90" type="nomenclature">
<paragraph id="8BE8435DA97A6A42FCB4FC3A1B64C2C9" blockId="15.[808,1079,912,971]" box="[808,1079,912,939]" pageId="15" pageNumber="90">
<heading id="D0A0F431A97A6A42FCB4FC3A1B64C2C9" box="[808,1079,912,939]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="90" reason="2">
<taxonomicName id="4C5738DEA97A6A42FCB4FC3A1CA6C2C9" ID-CoL="3VC68" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[808,1013,912,939]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="90" phylum="Arthropoda" rank="species" species="bernardi" status="sp. nov.">
<emphasis id="B9239F4FA97A6A42FCB4FC3A1CA6C2C9" bold="true" box="[808,1013,912,939]" italics="true" pageId="15" pageNumber="90">Lipseuma bernardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A97A6A42FC60FC3A1B64C2C9" box="[1020,1079,913,939]" pageId="15" pageNumber="90" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A97A6A42FCE4FC1B1CB5C2A9" box="[888,998,944,971]" pageId="15" pageNumber="90" type="description">
<paragraph id="8BE8435DA97A6A42FCE4FC1B1CB5C2A9" blockId="15.[808,1079,912,971]" box="[888,998,944,971]" pageId="15" pageNumber="90">
(
<figureCitation id="136C5FD8A97A6A42FCE3FC1B1C90C2A9" box="[895,963,944,971]" captionStart="FIG" captionStartId="14.[113,124,1635,1652]" captionTargetBox="[220,1146,220,1556]" captionTargetId="figure-86@14.[163,1164,162,1592]" captionTargetPageId="14" captionText="FIG. 7. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, antenna; B, left half of body segment 7, front view; C, D, leg 6, front and caudal views, respectively; E, leg 7, front view; F-H, anterior gonopods (leg-pair 8), front and caudal views, respectively. Scale bars: A-E, 0.2 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393518" httpUri="https://zenodo.org/record/5393518/files/figure.png" pageId="15" pageNumber="90">Figs 7</figureCitation>
;
<figureCitation id="136C5FD8A97A6A42FC53FC1A1C8EC2A9" box="[975,989,945,971]" captionStart="FIG" captionStartId="15.[113,124,818,835]" captionTargetBox="[130,1186,220,772]" captionTargetId="figure-54@15.[56,1265,145,822]" captionTargetPageId="15" captionText="FIG. 8. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, leg 10, front view; B, posterior gonopods, caudal view. Scale bar: A, 0.2 mm; B, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393520" httpUri="https://zenodo.org/record/5393520/files/figure.png" pageId="15" pageNumber="90">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A97A6A42FD39FC5A1CCBC510" pageId="15" pageNumber="90" type="materials_examined">
<paragraph id="8BE8435DA97A6A42FD39FC5A1CCBC510" blockId="15.[677,1210,1008,1138]" pageId="15" pageNumber="90">
<materialsCitation id="3B3F4900A97A6A42FD39FC5A1B01C535" collectingDate="1999-08-07" collectionCode="MNHN" collectorName="Three Eyes Cave, No. &amp; B. &amp; J. Lips" country="China" county="Xin Long County" location="Three Eyes Cave" pageId="15" pageNumber="90" specimenCode="DA156" specimenCount="2" specimenCount-male="2" stateProvince="Sichuan" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus id="54ECFDFFA97A6A42FCC2FC5B1CEFC56A" box="[862,956,1008,1032]" pageId="15" pageNumber="90" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34003CDA97A6A42FC5FFC5B1B52C56A" box="[963,1025,1008,1032]" name="China" pageId="15" pageNumber="90">China</collectingCountry>
,
<collectingRegion id="49938DBFA97A6A42FB96FC5B1BDCC56A" box="[1034,1167,1008,1032]" country="China" name="Sichuan" pageId="15" pageNumber="90">Sichuan Prov.</collectingRegion>
,
<collectingCounty id="62893BD1A97A6A42FB09FC5B1C79C541" pageId="15" pageNumber="90">Xin Long County</collectingCounty>
,
<collectorName id="26A2268BA97A6A42FCAFFBA01B57C540" box="[819,1028,1035,1059]" pageId="15" pageNumber="90">
<location id="8E881586A97A6A42FCAFFBA01C87C541" LSID="urn:lsid:plazi:treatment:03FEF24BA97A6A5DFCB4FC3A1DD3C4A7:8E881586A97A6A42FCAFFBA01C87C541" box="[819,980,1035,1059]" country="China" county="Xin Long County" name="Three Eyes Cave" pageId="15" pageNumber="90" stateProvince="Sichuan">Three Eyes Cave</location>
, No.
</collectorName>
438,
<date id="FFE9659DA97A6A42FBA0FBA01BE6C541" box="[1084,1205,1035,1059]" pageId="15" pageNumber="90" value="1999-08-07">
<collectingDate id="EFAD9C75A97A6A42FBA0FBA01BE6C541" box="[1084,1205,1035,1059]" pageId="15" pageNumber="90" value="1999-08-07">7.VIII.1999</collectingDate>
</date>
, leg.
<collectorName id="26A2268BA97A6A42FD50FB8D1C12C55F" box="[716,833,1061,1085]" pageId="15" pageNumber="90">B. &amp; J. Lips</collectorName>
,
<specimenCount id="9D5188D4A97A6A42FCD6FB831C0DC55E" box="[842,862,1064,1084]" pageId="15" pageNumber="90" type="male"></specimenCount>
(
<collectionCode id="ED46DB98A97A6A42FCF7FB8E1CECC55F" box="[875,959,1061,1085]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="90">MNHN</collectionCode>
<specimenCode id="DBF1EB26A97A6A42FC58FB8E1B41C55F" box="[964,1042,1061,1085]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="90">DA156</specimenCode>
).
<typeStatus id="54ECFDFFA97A6A42FBBEFB8D1BD2C55C" box="[1058,1153,1062,1086]" pageId="15" pageNumber="90" type="paratype">Paratypes</typeStatus>
: same locality,
<specimenCount id="9D5188D4A97A6A42FD6AFBEB1C4FC535" box="[758,796,1088,1112]" pageId="15" pageNumber="90" type="male">1 ♂</specimenCount>
, incomplete (
<collectionCode id="ED46DB98A97A6A42FC3FFBEB1CA4C53A" box="[931,1015,1088,1112]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="90">MNHN</collectionCode>
<specimenCode id="DBF1EB26A97A6A42FC62FB941B18C535" box="[1022,1099,1087,1111]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="90">DA156</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F4900A97A6A42FBC0FBEB1CC0C510" collectionCode="MNHN" country="China" pageId="15" pageNumber="90" specimenCode="DA156" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<specimenCount id="9D5188D4A97A6A42FBC0FBEB1BD0C535" box="[1116,1155,1088,1112]" pageId="15" pageNumber="90" type="male">1 ♂</specimenCount>
, subadult (
<collectionCode id="ED46DB98A97A6A42FD78FBF11C6BC510" box="[740,824,1114,1138]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="90">MNHN</collectionCode>
<specimenCode id="DBF1EB26A97A6A42FCA3FBF11CDFC510" box="[831,908,1114,1138]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="90">DA156</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A97A6A42FD39FB3A1B6DC5CA" box="[677,1086,1168,1192]" pageId="15" pageNumber="90" type="etymology">
<paragraph id="8BE8435DA97A6A42FD39FB3A1B6DC5CA" blockId="15.[677,1086,1168,1192]" box="[677,1086,1168,1192]" pageId="15" pageNumber="90">ETYMOLOGY. — To honour Bernard Lips.</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A97A6A42FD39FB6C1B34C471" pageId="15" pageNumber="90" type="diagnosis">
<paragraph id="8BE8435DA97A6A42FD39FB6C1B34C471" blockId="15.[677,1210,1223,1300]" pageId="15" pageNumber="90">
DIAGNOSIS. — Differs from the only congener (
<taxonomicName id="4C5738DEA97A6A42FBE4FB631D82C59B" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="90" phylum="Arthropoda" rank="species" species="josianae" status="n. gen., n. sp.">
<emphasis id="B9239F4FA97A6A42FBE4FB631D82C59B" italics="true" pageId="15" pageNumber="90">L. josianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A97A6A42FD4BFB491C0DC59B" box="[727,862,1249,1274]" pageId="15" pageNumber="90" rank="species">n. gen., n. sp.</taxonomicNameLabel>
) by the slightly smaller size, as well as minor details of leg and gonopod structure.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A97A6A5DFD39FAE61E7AC32C" lastPageId="16" lastPageNumber="91" pageId="15" pageNumber="90" type="description">
<paragraph id="8BE8435DA97A6A42FD39FAE61C66C407" blockId="15.[677,1210,1357,1700]" box="[677,821,1357,1383]" pageId="15" pageNumber="90">DESCRIPTION</paragraph>
<paragraph id="8BE8435DA97A6A42FD39FAC71C29C484" blockId="15.[677,1210,1357,1700]" pageId="15" pageNumber="90">
Length 7.2 (
<typeStatus id="54ECFDFFA97A6A42FCB3FAC71CC0C4E5" box="[815,915,1388,1415]" pageId="15" pageNumber="90" type="holotype">holotype</typeStatus>
) or
<quantity id="4CAFEEB8A97A6A42FC58FAC71B4EC4E5" box="[964,1053,1388,1415]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="15" pageNumber="90" unit="mm" value="6.8">6.8 mm</quantity>
(subadult
<specimenCount id="9D5188D4A97A6A42FB0AFAC41BFAC4E7" box="[1174,1193,1391,1413]" pageId="15" pageNumber="90" type="female"></specimenCount>
); width of body segment 6,
<quantity id="4CAFEEB8A97A6A42FC5AFA271B7DC4C4" box="[966,1070,1420,1446]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="15" pageNumber="90" unit="mm" value="0.72">0.72 mm</quantity>
, of midbody segments,
<quantity id="4CAFEEB8A97A6A42FC8BFA001CD3C4A4" box="[791,896,1451,1478]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="90" unit="mm" value="0.78">0.78 mm</quantity>
, maximum breadth of head,
<quantity id="4CAFEEB8A97A6A42FD39FA601DA8C484" box="[677,763,1483,1510]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="90" unit="mm" value="0.9">0.9 mm</quantity>
(
<typeStatus id="54ECFDFFA97A6A42FC95FA601C3DC484" box="[777,878,1483,1510]" pageId="15" pageNumber="90" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BE8435DA97A6A42FD27FA401B07C746" blockId="15.[677,1210,1357,1700]" pageId="15" pageNumber="90">
Antennae (
<figureCitation id="136C5FD8A97A6A42FCB5FA401C27C767" box="[809,884,1515,1541]" captionStart="FIG" captionStartId="14.[113,124,1635,1652]" captionTargetBox="[220,1146,220,1556]" captionTargetId="figure-86@14.[163,1164,162,1592]" captionTargetPageId="14" captionText="FIG. 7. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, antenna; B, left half of body segment 7, front view; C, D, leg 6, front and caudal views, respectively; E, leg 7, front view; F-H, anterior gonopods (leg-pair 8), front and caudal views, respectively. Scale bars: A-E, 0.2 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393518" httpUri="https://zenodo.org/record/5393518/files/figure.png" pageId="15" pageNumber="90">Fig. 7A</figureCitation>
) with even more exposed, highly elongate, pointed apical cones on stalks.
</paragraph>
<paragraph id="8BE8435DA97A6A42FD27F9811CAFC7C6" blockId="15.[677,1210,1357,1700]" pageId="15" pageNumber="90">
All peripheral characters like in
<taxonomicName id="4C5738DEA97A6A42FBB9F9801BC9C726" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[1061,1178,1579,1604]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="90" phylum="Arthropoda" rank="species" species="josianae" status="n. gen., n. sp.">
<emphasis id="B9239F4FA97A6A42FBB9F9801BC9C726" box="[1061,1178,1579,1604]" italics="true" pageId="15" pageNumber="90">L. josianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A97A6A42FB3FF9801C44C706" pageId="15" pageNumber="90" rank="species">n. gen., n. sp.</taxonomicNameLabel>
except as follows: exterior macrochaetae on midbody segments
<quantity id="4CAFEEB8A97A6A42FC06F9C21CA3C7E6" box="[922,1008,1641,1668]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="90" unit="mm" value="0.6">0.6 mm</quantity>
long, i.e. nearly as wide as segment itself (
<figureCitation id="136C5FD8A97A6A42FC00F9221CBCC7C6" box="[924,1007,1673,1700]" captionStart="FIG" captionStartId="14.[113,124,1635,1652]" captionTargetBox="[220,1146,220,1556]" captionTargetId="figure-86@14.[163,1164,162,1592]" captionTargetPageId="14" captionText="FIG. 7. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, antenna; B, left half of body segment 7, front view; C, D, leg 6, front and caudal views, respectively; E, leg 7, front view; F-H, anterior gonopods (leg-pair 8), front and caudal views, respectively. Scale bars: A-E, 0.2 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393518" httpUri="https://zenodo.org/record/5393518/files/figure.png" pageId="15" pageNumber="90">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8BE8435DA9656A5DFF14FF731D3AC033" blockId="16.[113,647,216,590]" pageId="16" pageNumber="91">
Male legs 6 (
<figureCitation id="136C5FD8A9656A5DFE8AFF731EC1C190" box="[278,402,216,242]" captionStart="FIG" captionStartId="14.[113,124,1635,1652]" captionTargetBox="[220,1146,220,1556]" captionTargetId="figure-86@14.[163,1164,162,1592]" captionTargetPageId="14" captionText="FIG. 7. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, antenna; B, left half of body segment 7, front view; C, D, leg 6, front and caudal views, respectively; E, leg 7, front view; F-H, anterior gonopods (leg-pair 8), front and caudal views, respectively. Scale bars: A-E, 0.2 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393518" httpUri="https://zenodo.org/record/5393518/files/figure.png" pageId="16" pageNumber="91">Fig. 7C, D</figureCitation>
), 10 (
<figureCitation id="136C5FD8A9656A5DFE49FF731D7AC190" box="[469,553,216,242]" captionStart="FIG" captionStartId="15.[113,124,818,835]" captionTargetBox="[130,1186,220,772]" captionTargetId="figure-54@15.[56,1265,145,822]" captionTargetPageId="15" captionText="FIG. 8. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, leg 10, front view; B, posterior gonopods, caudal view. Scale bar: A, 0.2 mm; B, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393520" httpUri="https://zenodo.org/record/5393520/files/figure.png" pageId="16" pageNumber="91">Fig. 8A</figureCitation>
) and 11 virtually the same as in
<taxonomicName id="4C5738DEA9656A5DFEEFFF531EB0C073" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[371,483,248,273]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="species" species="josianae" status="n. gen., n. sp.">
<emphasis id="B9239F4FA9656A5DFEEFFF531EB0C073" box="[371,483,248,273]" italics="true" pageId="16" pageNumber="91">L. josianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFE77FF531DD1C070" box="[491,642,248,274]" pageId="16" pageNumber="91" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, but male legs 7 (
<figureCitation id="136C5FD8A9656A5DFE83FEBC1E3CC050" box="[287,367,279,306]" captionStart="FIG" captionStartId="14.[113,124,1635,1652]" captionTargetBox="[220,1146,220,1556]" captionTargetId="figure-86@14.[163,1164,162,1592]" captionTargetPageId="14" captionText="FIG. 7. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, antenna; B, left half of body segment 7, front view; C, D, leg 6, front and caudal views, respectively; E, leg 7, front view; F-H, anterior gonopods (leg-pair 8), front and caudal views, respectively. Scale bars: A-E, 0.2 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393518" httpUri="https://zenodo.org/record/5393518/files/figure.png" pageId="16" pageNumber="91">Fig. 7E</figureCitation>
) with a characteristic protuberance on tarsus ventrally at about midway.
</paragraph>
<paragraph id="8BE8435DA9656A5DFF14FEFD1EB9C0AD" blockId="16.[113,647,216,590]" pageId="16" pageNumber="91">
Anterior gonopods (
<figureCitation id="136C5FD8A9656A5DFEC3FEFD1E99C013" box="[351,458,342,369]" captionStart="FIG" captionStartId="14.[113,124,1635,1652]" captionTargetBox="[220,1146,220,1556]" captionTargetId="figure-86@14.[163,1164,162,1592]" captionTargetPageId="14" captionText="FIG. 7. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, antenna; B, left half of body segment 7, front view; C, D, leg 6, front and caudal views, respectively; E, leg 7, front view; F-H, anterior gonopods (leg-pair 8), front and caudal views, respectively. Scale bars: A-E, 0.2 mm; F, G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393518" httpUri="https://zenodo.org/record/5393518/files/figure.png" pageId="16" pageNumber="91">Fig. 7F, G</figureCitation>
) also virtually the same as in
<taxonomicName id="4C5738DEA9656A5DFF79FEDC1E07C0F2" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[229,340,375,400]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="species" species="josianae" status="n. gen., n. sp.">
<emphasis id="B9239F4FA9656A5DFF79FEDC1E07C0F2" box="[229,340,375,400]" italics="true" pageId="16" pageNumber="91">L. josianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFEC7FEDC1EBCC0F3" box="[347,495,375,401]" pageId="16" pageNumber="91" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, but posterior gonopods (
<figureCitation id="136C5FD8A9656A5DFF70FE3D1E6CC0D2" box="[236,319,406,432]" captionStart="FIG" captionStartId="15.[113,124,818,835]" captionTargetBox="[130,1186,220,772]" captionTargetId="figure-54@15.[56,1265,145,822]" captionTargetPageId="15" captionText="FIG. 8. — Lipseuma bernardi n. gen., n. sp., paratype ♂: A, leg 10, front view; B, posterior gonopods, caudal view. Scale bar: A, 0.2 mm; B, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393520" httpUri="https://zenodo.org/record/5393520/files/figure.png" pageId="16" pageNumber="91">Fig. 8B</figureCitation>
) with traces of a second telopoditomere distally on caudal face.
</paragraph>
<paragraph id="8BE8435DA9656A5DFF14FE7E1E7AC32C" blockId="16.[113,647,216,590]" pageId="16" pageNumber="91">Subadult male (26 body segments) with small but evident gonopod promordia, legs 7 almost as strongly incrassate as in adult male, legs 7 also slightly enlarged.</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9656A5DFFEDFDDF1DD3C4A7" pageId="16" pageNumber="91" type="discussion">
<paragraph id="8BE8435DA9656A5DFFEDFDDF1F86C3EE" blockId="16.[113,648,628,1478]" box="[113,213,628,654]" pageId="16" pageNumber="91">REMARKS</paragraph>
<paragraph id="8BE8435DA9656A5DFFEDFD381E70C22E" blockId="16.[113,648,628,1478]" pageId="16" pageNumber="91">
Both
<taxonomicName id="4C5738DEA9656A5DFF2DFD3F1E46C3CF" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[177,277,660,685]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9239F4FA9656A5DFF2DFD3F1E46C3CF" box="[177,277,660,685]" italics="true" pageId="16" pageNumber="91">Lipseuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFE82FD3F1E39C3CC" box="[286,362,660,686]" pageId="16" pageNumber="91" rank="genus">n. gen.</taxonomicNameLabel>
species are remarkable in being completely unpigmented and eyeless, as well as in bearing strongly elongated antennae, legs (especially tarsi) and exterior macrochaetae. All these features leave no doubt that we are dealing with troglobites.
</paragraph>
<paragraph id="8BE8435DA9656A5DFF14FCF91DD3C4A7" blockId="16.[113,648,628,1478]" pageId="16" pageNumber="91">
It is noteworthy that interspecific variation in
<taxonomicName id="4C5738DEA9656A5DFFEDFCD91F87C2E9" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[113,212,882,907]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9239F4FA9656A5DFFEDFCD91F87C2E9" box="[113,212,882,907]" italics="true" pageId="16" pageNumber="91">Lipseuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFF46FCD91E71C2EE" box="[218,290,882,908]" pageId="16" pageNumber="91" rank="genus">n. gen.</taxonomicNameLabel>
is very modest, in contrast to that observed in the other two kashmireumatid genera. In fact,
<taxonomicName id="4C5738DEA9656A5DFF54FC1A1E6BC2A8" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[200,312,945,970]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="species" species="josianae" status="n. gen., n. sp.">
<emphasis id="B9239F4FA9656A5DFF54FC1A1E6BC2A8" box="[200,312,945,970]" italics="true" pageId="16" pageNumber="91">L. josianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFEDDFC1A1E85C2A9" box="[321,470,945,971]" pageId="16" pageNumber="91" rank="species">n. gen., n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C5738DEA9656A5DFD93FC1A1DD5C2A8" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[527,646,944,970]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="species" species="bernardi" status="n. gen., n. sp.">
<emphasis id="B9239F4FA9656A5DFD93FC1A1DD5C2A8" box="[527,646,944,970]" italics="true" pageId="16" pageNumber="91">L. bernardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFFEDFC7A1E57C289" box="[113,260,977,1003]" pageId="16" pageNumber="91" rank="species">n. gen., n. sp.</taxonomicNameLabel>
are barely distinguishable. The very small differences noted between them suggest either a quite recent colonization of caves by their common ancestor or a fairly recent vicariance event with speciation within their respective cave systems. The latter scenario seems more plausible, given the profound troglomorphism of both species, which would require a geological time-scale to develop, along with the geographical proximity of Chuan Dong Zi and Three Eyes caves (both situated in the Xin Long geographical zone just at the border between
<collectingRegion id="49938DBFA9656A5DFF52FA861E41C425" box="[206,274,1325,1351]" country="China" name="Hubei" pageId="16" pageNumber="91">Hubei</collectingRegion>
and
<collectingRegion id="49938DBFA9656A5DFED8FA861ECAC425" box="[324,409,1325,1351]" country="China" name="Sichuan" pageId="16" pageNumber="91">Sichuan</collectingRegion>
provinces [
<bibRefCitation id="EFC63EACA9656A5DFD92FA861F9DC405" author="DEGOUVE P. &amp; LIPS B. &amp; LIPS J." pageId="16" pageNumber="91" refId="ref7818" refString="DEGOUVE P., LIPS B. &amp; LIPS J. 1997. - Speleologie au pays de l'homme sauvage no 3. Rapport de la 3 e expedition speleologique en Chine, A. K. L. 1997, province du Sichuan. Federation francaise de Speleologie, Paris, 130 p." type="book" year="1997">
Degouve
<emphasis id="B9239F4FA9656A5DFDE9FA861FDFC404" italics="true" pageId="16" pageNumber="91">et al.</emphasis>
1997
</bibRefCitation>
]). In other words, the old geological age of the genus
<taxonomicName id="4C5738DEA9656A5DFF41FAC61E13C4E4" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[221,320,1389,1414]" class="Diplopoda" family="Kashmireumatidae" genus="Lipseuma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="91" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9239F4FA9656A5DFF41FAC61E13C4E4" box="[221,320,1389,1414]" italics="true" pageId="16" pageNumber="91">Lipseuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9656A5DFEDBFAC61EC3C4E5" box="[327,400,1389,1415]" pageId="16" pageNumber="91" rank="genus">n. gen.</taxonomicNameLabel>
seems unquestionable, whereas the isolation of both hitherto known congeners probably occurred only relatively recently.
</paragraph>
</subSubSection>
</treatment>
</document>