248 lines
40 KiB
XML
248 lines
40 KiB
XML
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<mods:title id="CDAAE918C491A5A7558CCD957938629F">Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)</mods:title>
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<mods:namePart id="F60426AE4563CC38EAF427DD532B8F35">Murányi, Dávid</mods:namePart>
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<mods:nameIdentifier id="0D0A56919414C3C16770491E1FF91613" type="email">muranyi@zool.nhmus.hu</mods:nameIdentifier>
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<mods:namePart id="C535473AE31690CF86C9BE01EBFA5627">Gamboa, Maribet</mods:namePart>
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<mods:nameIdentifier id="CAD94FDE2ADE6C4CD5D67147D4936AB6" type="email">maribetg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4DC0E4DD1EEF818C3805D3BEC67A3E29">Orci, Kirill Márk</mods:namePart>
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<mods:nameIdentifier id="5B128D497CCC5A72A7E433EFEBDD9270" type="email">muranyi@zool.nhmus.hu</mods:nameIdentifier>
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<mods:title id="462FA34A36B295D3C0743BEC05D9977F">Zootaxa</mods:title>
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<mods:date id="4A127869BC63DD5A1E110EF20DB2898E">2014</mods:date>
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<treatment id="03F99336FF3CFFDB1BE4580FD6A6F83F" ID-DOI="http://doi.org/10.5281/zenodo.4919265" ID-GBIF-Taxon="183709168" ID-Zenodo-Dep="4919265" LSID="urn:lsid:plazi:treatment:03F99336FF3CFFDB1BE4580FD6A6F83F" httpUri="http://treatment.plazi.org/id/03F99336FF3CFFDB1BE4580FD6A6F83F" lastPageNumber="53" pageId="52" pageNumber="53">
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<paragraph id="8BEF2220FF3CFFDB1BE4580FD6B2FCBB" blockId="52.[151,907,844,903]" box="[151,907,844,870]" pageId="52" pageNumber="53">
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<heading id="D0A7954CFF3CFFDB1BE4580FD6B2FCBB" bold="true" box="[151,907,844,870]" fontSize="11" level="2" pageId="52" pageNumber="53" reason="4">
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<emphasis id="B924FE32FF3CFFDB1BE4580FD6B2FCBB" bold="true" box="[151,907,844,870]" pageId="52" pageNumber="53">
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<taxonomicName id="4C5059A3FF3CFFDB1BE4580FD621FCBB" ID-CoL="7L82B" authority="turkestanica (Kimmins, 1950)" authorityName="turkestanica (Kimmins" authorityYear="1950" box="[151,792,844,870]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="turkestanica" status="comb. nov." subSpecies="turkestanica">
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<emphasis id="B924FE32FF3CFFDB1BE4580FD779FCBB" bold="true" box="[151,576,844,870]" italics="true" pageId="52" pageNumber="53">Zwicknia turkestanica turkestanica</emphasis>
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(Kimmins, 1950)
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</taxonomicName>
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<taxonomicNameLabel id="A2174349FF3CFFDB186C580FD6B2FCBB" box="[799,907,844,870]" pageId="52" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C34A71ABFF3CFFDB1BE4582DD758FC5A" box="[151,609,878,903]" pageId="52" pageNumber="53" type="description">
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<paragraph id="8BEF2220FF3CFFDB1BE4582DD758FC5A" blockId="52.[151,907,844,903]" box="[151,609,878,903]" pageId="52" pageNumber="53">
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(
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<figureCitation id="136B3EA5FF3CFFDB1BEC582DD47AFC5A" box="[159,323,878,903]" captionStart="FIGURES 102–105" captionStartId="53.[151,264,1955,1978]" captionTargetBox="[235,1330,185,1912]" captionTargetId="figure-52@53.[235,1352,171,1935]" captionTargetPageId="53" captionText="FIGURES 102–105. Male terminalia of Zwicknia turkestanica turkestanica (Kimmins, 1950); Kazakhstan, East Kazakhstan Province, Tarbagatai Mts., Ayagos—102: dorsal view; 103: ventral view; 104: lateral view; 105: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919134" httpUri="https://zenodo.org/record/4919134/files/figure.png" pageId="52" pageNumber="53">Figs. 102–105</figureCitation>
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,
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<figureCitation id="136B3EA5FF3CFFDB1A3D582DD440FC5A" box="[334,377,878,903]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi & Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="52" pageNumber="53">113</figureCitation>
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,
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<figureCitation id="136B3EA5FF3CFFDB1AF6582DD488FC5A" box="[389,433,878,903]" captionStart="FIGURES 124–126" captionStartId="58.[151,264,1502,1525]" captionTargetBox="[166,1399,232,1399]" captionTargetId="figure-159@58.[151,1436,223,1410]" captionTargetPageId="58" captionText="FIGURES 124–126. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—124: Z. sevanica (Zhiltzova, 1964); 125: Z. tuberculata (Zhiltzova, 1964); 126: Z. turkestanica turkestanica (Kimmins, 1950)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919144" httpUri="https://zenodo.org/record/4919144/files/figure.png" pageId="52" pageNumber="53">126</figureCitation>
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,
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<figureCitation id="136B3EA5FF3CFFDB1ACE582DD4D3FC5A" box="[445,490,878,903]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi & Orci, sp. n.; 131–132: Z. kovacsi Murányi & Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci & Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="52" pageNumber="53">147</figureCitation>
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,
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||
<figureCitation id="136B3EA5FF3CFFDB1A86582DD71BFC5A" box="[501,546,878,903]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi & Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci & Gamboa, sp. n.; 160: Z. kovacsi Murányi & Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="52" pageNumber="53">164</figureCitation>
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,
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<figureCitation id="136B3EA5FF3CFFDB195E582DD761FC5A" box="[557,600,878,903]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="52" pageNumber="53">196</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A71ABFF3CFFDB1BE458F5D056FBD7" pageId="52" pageNumber="53" type="reference_group">
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<paragraph id="8BEF2220FF3CFFDB1BE458F5D1ECFC11" blockId="52.[151,1437,950,1034]" box="[151,1237,950,972]" pageId="52" pageNumber="53">
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<treatmentCitationGroup id="AB40050EFF3CFFDB1BE458F5D1ECFC11" box="[151,1237,950,972]" pageId="52" pageNumber="53">
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<taxonomicName id="4C5059A3FF3CFFDB1BE458F5D492FC11" ID-CoL="5X9KQ" authority="(Pictet, 1833)" authorityName="sensu Morton" authorityYear="1896" baseAuthorityName="Pictet" baseAuthorityYear="1833" box="[151,427,950,972]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="nigra">
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<emphasis id="B924FE32FF3CFFDB1BE458F5D426FC11" box="[151,287,950,972]" italics="true" pageId="52" pageNumber="53">Capnia nigra</emphasis>
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(
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<bibRefCitation id="EFC15FD1FF3CFFDB1A5F58F5D49AFC11" author="Pictet, F. J." box="[300,419,950,972]" pageId="52" pageNumber="53" pagination="44 - 65" refId="ref44601" refString="Pictet, F. J. (1833) Memoire sur les Metamorphoses des Perles. Annales des Sciences Naturales, 28, 44 - 65." type="journal article" year="1833">Pictet, 1833</bibRefCitation>
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)
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||
</taxonomicName>
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||
—
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||
<treatmentCitation id="0AF10431FF3CFFDB1AB658F5D74FFC11" author="Morton, K. J." box="[453,630,950,972]" page="60" pageId="52" pageNumber="53" year="1896">
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<bibRefCitation id="EFC15FD1FF3CFFDB1AB658F5D74FFC11" author="Morton, K. J." box="[453,630,950,972]" pageId="52" pageNumber="53" pagination="55 - 63" refId="ref43808" refString="Morton, K. J. (1896) New and little-known Palaearctic Perlidae. The Transactions of the Entomological Society of London, 1896 (1), 55 - 63." type="journal article" year="1896">Morton 1896: 60</bibRefCitation>
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</treatmentCitation>
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. (
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<figureCitation id="136B3EA5FF3CFFDB19FB58F5D7FAFC11" box="[648,707,950,972]" captionStart="FIGURES 1–6" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[193,1399,221,1867]" captionTargetId="figure-86@8.[151,1436,195,1900]" captionTargetPageId="8" captionText="FIGURES 1–6. Male terminalia of West Palaearctic Capniidae—1–2: Zwicknia bifrons (Newman); 3–4: Capnia s.s. nigra (Pictet); 5–6: Capnia s.l. vidua Klapálek; 1, 3, 5: dorso-caudal view; 2, 4, 6: ventro-caudal view (B-scl: basal sclerite; Cs: caudal setae; Ec: eversible crest; Ep-scl: main epiproct sclerite (shaded in grey); I-scl: inner sclerite; Lb-scl: laterobasal sclerite; Ll: lower limb; Tg 10: tergite 10)—not to scale." figureDoi="http://doi.org/10.5281/zenodo.4919081" httpUri="https://zenodo.org/record/4919081/files/figure.png" pageId="52" pageNumber="53">Fig. 3</figureCitation>
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on Plate 2 refers to the
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<typeStatus id="54EB9C82FF3CFFDB18C658F5D137FC11" box="[949,1038,950,972]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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||
of
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||
<taxonomicName id="4C5059A3FF3CFFDB1F5C58F4D1FEFC11" ID-CoL="7GGKJ" authorityName="Murányi & Gamboa & Orci" authorityYear="2014" baseAuthorityName="Kimmins" baseAuthorityYear="1950" box="[1071,1223,950,972]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="turkestanica">
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||
<emphasis id="B924FE32FF3CFFDB1F5C58F4D1FEFC11" box="[1071,1223,950,972]" italics="true" pageId="52" pageNumber="53">Z. turkestanica</emphasis>
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||
</taxonomicName>
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||
).
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BEF2220FF3CFFDB1BE45896D056FBD7" blockId="52.[151,1437,950,1034]" pageId="52" pageNumber="53">
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<treatmentCitationGroup id="AB40050EFF3CFFDB1BE45896D056FBD7" pageId="52" pageNumber="53">
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<taxonomicName id="4C5059A3FF3CFFDB1BE45896D726FC36" ID-CoL="5WXK8" authority="Kimmins, 1950" authorityName="Kimmins" authorityYear="1950" box="[151,543,981,1003]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="turkestanica">
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<emphasis id="B924FE32FF3CFFDB1BE45896D454FC36" box="[151,365,981,1003]" italics="true" pageId="52" pageNumber="53">Capnia turkestanica</emphasis>
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||
Kimmins, 1950
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</taxonomicName>
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||
—
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<treatmentCitation id="0AF10431FF3CFFDB194A5896D61BFC36" author="Kimmins, D. E." box="[569,802,981,1003]" page="187" pageId="52" pageNumber="53" year="1950">
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<bibRefCitation id="EFC15FD1FF3CFFDB194A5896D61BFC36" author="Kimmins, D. E." box="[569,802,981,1003]" pageId="52" pageNumber="53" pagination="177 - 192" refId="ref43063" refString="Kimmins, D. E. (1950 a) Some new species of Asiatic Plecoptera. The Annals and Magazine of Natural History, 12 (3), 177 - 192." type="journal article" year="1950">Kimmins 1950a: 187</bibRefCitation>
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||
</treatmentCitation>
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||
. (original description of the adult);
|
||
<treatmentCitation id="0AF10431FF3CFFDB1FCA5896D0A0FC36" author="Zhiltzova, L. A." box="[1209,1433,981,1003]" page="603" pageId="52" pageNumber="53" year="1969">
|
||
<bibRefCitation id="EFC15FD1FF3CFFDB1FCA5896D0A0FC36" author="Zhiltzova, L. A." box="[1209,1433,981,1003]" pageId="52" pageNumber="53" pagination="593 - 611" refId="ref46461" refString="Zhiltzova, L. A. (1969) Novie I redkie vidi vesnianok sem. Capniidae (Plecoptera) iz Srednei Azii. Revue d'Entomologie de l'USSR, 48 (3), 593 - 611." type="journal article" year="1969">Zhiltzova 1969: 603</bibRefCitation>
|
||
</treatmentCitation>
|
||
. (complementary description and comparison with
|
||
<taxonomicName id="4C5059A3FF3CFFDB19B758B6D66AFBD7" ID-CoL="7GGKH" authorityName="Murányi & Gamboa & Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[708,851,1012,1034]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tuberculata">
|
||
<emphasis id="B924FE32FF3CFFDB19B758B6D66AFBD7" box="[708,851,1012,1034]" italics="true" pageId="52" pageNumber="53">Z. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<treatmentCitation id="0AF10431FF3CFFDB181A58B7D100FBD7" author="Zhiltzova, L. A." box="[873,1081,1012,1034]" page="349" pageId="52" pageNumber="53" year="2003">
|
||
<bibRefCitation id="EFC15FD1FF3CFFDB181A58B7D100FBD7" author="Zhiltzova, L. A." box="[873,1081,1012,1034]" pageId="52" pageNumber="53" pagination="1 - 538" refId="ref46559" refString="Zhiltzova, L. A. (2003) Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538." type="journal article" year="2003">Zhiltzova 2003: 349</bibRefCitation>
|
||
</treatmentCitation>
|
||
. (complementary description).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A71ABFF3CFFDB1BE45F75D7A8FB64" pageId="52" pageNumber="53" type="diagnosis">
|
||
<paragraph id="8BEF2220FF3CFFDB1BE45F75D7A8FB64" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
<emphasis id="B924FE32FF3CFFDB1BE45F75D428FB92" bold="true" box="[151,273,1078,1103]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
|
||
Male epiproct: Ep-scl narrow and acute in dorsal view, short, tip slightly downcurved in lateral view, membranous portion long; ventral membranous section reaching base in lateral view, apical spines long, distributed also on the apex of Ep-scl. Process of male Tg 9: high, perpendicularly elevated, <2× wider than Ep-scl, rounded and with continuously narrow sides caudally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A71ABFF3CFFDB1BB45F80D6C9FAFE" pageId="52" pageNumber="53" type="materials_examined">
|
||
<paragraph id="8BEF2220FF3CFFDB1BB45F80D6C9FAFE" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
<emphasis id="B924FE32FF3CFFDB1BB45F80D488FB01" bold="true" box="[199,433,1219,1244]" pageId="52" pageNumber="53">Material examined.</emphasis>
|
||
<materialsCitation id="3B38287DFF3CFFDB1ACA5F80D6D5FAFE" ID-GBIF-Occurrence="3325453312" collectionCode="V, PZC" collectorName="I. Devyatkov" country="Kazakhstan" elevation="6" location="River Ayagos" municipality="Tarbagatai Mts." pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Shyghys Qazaqstan Oblysy">
|
||
<collectingCountry id="F34762B0FF3CFFDB1ACA5F80D742FB01" box="[441,635,1219,1244]" name="Kazakhstan" pageId="52" pageNumber="53">KAZAKHSTAN</collectingCountry>
|
||
:
|
||
<collectingRegion id="4994ECC2FF3CFFDB19F55F87D689FB01" box="[646,944,1219,1244]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="52" pageNumber="53">East Kazakhstan Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B8BB85AFF3CFFDB18C85F80D157FB06" box="[955,1134,1219,1244]" pageId="52" pageNumber="53">Tarbagatai Mts.</collectingMunicipality>
|
||
,
|
||
<location id="8E8F74FBFF3CFFDB1F085F80D02DFB01" LSID="urn:lsid:plazi:treatment:03F99336FF3CFFDB1BE4580FD6A6F83F:8E8F74FBFF3CFFDB1F085F80D02DFB01" box="[1147,1300,1219,1244]" country="Kazakhstan" municipality="Tarbagatai Mts." name="River Ayagos" pageId="52" pageNumber="53" stateProvince="Shyghys Qazaqstan Oblysy">River Ayagos</location>
|
||
<locationDeviation id="9A5D3B02FF3CFFDB1E6F5F80D41AFB22" location="Oskemen" pageId="52" pageNumber="53">
|
||
<quantity id="4CA88FC5FF3CFFDB1E6F5F80D050FB01" box="[1308,1385,1219,1244]" metricMagnitude="5" metricUnit="m" metricValue="2.7" pageId="52" pageNumber="53" unit="km" value="270.0">270km</quantity>
|
||
SW of Öskemen
|
||
</locationDeviation>
|
||
(Ust-Kamenogorsk), 05.04.200 3, leg., det.
|
||
<collectionCode id="ED41BAE5FF3CFFDB186C5FA4D60AFB23" box="[799,819,1255,1278]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="52" pageNumber="53" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="26A547F6FF3CFFDB18325FA4D6E2FB22" box="[833,987,1254,1279]" pageId="52" pageNumber="53">I. Devyatkov</collectorName>
|
||
:
|
||
<quantity id="4CA88FC5FF3CFFDB18995FA5D137FB22" box="[1002,1038,1254,1279]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="m" value="6.0">
|
||
<elevation id="007DC513FF3CFFDB18995FA5D137FB22" box="[1002,1038,1254,1279]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="m" value="6.0">6m</elevation>
|
||
</quantity>
|
||
6f (
|
||
<collectionCode id="ED41BAE5FF3CFFDB1F315FA5D143FB22" box="[1090,1146,1254,1279]" pageId="52" pageNumber="53">PZC</collectionCode>
|
||
; used for drawings
|
||
<figureCitation id="136B3EA5FF3CFFDB1E165FA5D5C2FAFE" captionStart="FIGURES 102–105" captionStartId="53.[151,264,1955,1978]" captionTargetBox="[235,1330,185,1912]" captionTargetId="figure-52@53.[235,1352,171,1935]" captionTargetPageId="53" captionText="FIGURES 102–105. Male terminalia of Zwicknia turkestanica turkestanica (Kimmins, 1950); Kazakhstan, East Kazakhstan Province, Tarbagatai Mts., Ayagos—102: dorsal view; 103: ventral view; 104: lateral view; 105: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919134" httpUri="https://zenodo.org/record/4919134/files/figure.png" pageId="52" pageNumber="53">Figs. 102–105</figureCitation>
|
||
,
|
||
<figureCitation id="136B3EA5FF3CFFDB1A745E49D40AFAFE" box="[263,307,1290,1315]" captionStart="FIGURES 124–126" captionStartId="58.[151,264,1502,1525]" captionTargetBox="[166,1399,232,1399]" captionTargetId="figure-159@58.[151,1436,223,1410]" captionTargetPageId="58" captionText="FIGURES 124–126. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—124: Z. sevanica (Zhiltzova, 1964); 125: Z. tuberculata (Zhiltzova, 1964); 126: Z. turkestanica turkestanica (Kimmins, 1950)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919144" httpUri="https://zenodo.org/record/4919144/files/figure.png" pageId="52" pageNumber="53">126</figureCitation>
|
||
,
|
||
<figureCitation id="136B3EA5FF3CFFDB1A4C5E49D452FAFE" box="[319,363,1290,1315]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi & Orci, sp. n.; 131–132: Z. kovacsi Murányi & Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci & Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="52" pageNumber="53">147</figureCitation>
|
||
,
|
||
<figureCitation id="136B3EA5FF3CFFDB1A045E49D49DFAFE" box="[375,420,1290,1315]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi & Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci & Gamboa, sp. n.; 160: Z. kovacsi Murányi & Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="52" pageNumber="53">164</figureCitation>
|
||
;
|
||
<specimenCount id="9D56E9A9FF3CFFDB1AC35E49D72CFAFE" box="[432,533,1290,1315]" pageId="52" pageNumber="53" type="male">one male</specimenCount>
|
||
terminalia prepared for SEM to
|
||
<figureCitation id="136B3EA5FF3CFFDB18F05E49D6DAFAFE" box="[899,995,1290,1315]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi & Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="52" pageNumber="53">Fig. 113</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A71ABFF3CFFDB1BB45E6ED1B2F8A4" pageId="52" pageNumber="53" type="description">
|
||
<paragraph id="8BEF2220FF3CFFDB1BB45E6ED14FFA56" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
<emphasis id="B924FE32FF3CFFDB1BB45E6ED461FA9B" bold="true" box="[199,344,1325,1350]" pageId="52" pageNumber="53">Description.</emphasis>
|
||
Head, thorax, appendages and basal segments of the abdomen generotypic. Males micropterous or macropterous, females macropterous. Dimensions of the presently examined specimens: body length: males 5.5–6.5, females 6.0–7.5 mm; forewing length: males 1.0–6.0 mm, females 6.5–7.5 mm.
|
||
</paragraph>
|
||
<paragraph id="8BEF2220FF3CFFDB1BB45ED5D6D0F97B" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
Male terminalia (
|
||
<figureCitation id="136B3EA5FF3CFFDB1AFA5ED5D717FA72" box="[393,558,1430,1455]" captionStart="FIGURES 102–105" captionStartId="53.[151,264,1955,1978]" captionTargetBox="[235,1330,185,1912]" captionTargetId="figure-52@53.[235,1352,171,1935]" captionTargetPageId="53" captionText="FIGURES 102–105. Male terminalia of Zwicknia turkestanica turkestanica (Kimmins, 1950); Kazakhstan, East Kazakhstan Province, Tarbagatai Mts., Ayagos—102: dorsal view; 103: ventral view; 104: lateral view; 105: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919134" httpUri="https://zenodo.org/record/4919134/files/figure.png" pageId="52" pageNumber="53">Figs. 102–105</figureCitation>
|
||
,
|
||
<figureCitation id="136B3EA5FF3CFFDB19495ED5D751FA72" box="[570,616,1430,1455]" captionStart="FIGURES 124–126" captionStartId="58.[151,264,1502,1525]" captionTargetBox="[166,1399,232,1399]" captionTargetId="figure-159@58.[151,1436,223,1410]" captionTargetPageId="58" captionText="FIGURES 124–126. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—124: Z. sevanica (Zhiltzova, 1964); 125: Z. tuberculata (Zhiltzova, 1964); 126: Z. turkestanica turkestanica (Kimmins, 1950)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919144" httpUri="https://zenodo.org/record/4919144/files/figure.png" pageId="52" pageNumber="53">126</figureCitation>
|
||
): Process of Tg 9 high, perpendicularly elevated, apex is <2× wider than the medial section of Ep-scl; its shape is rounded, the apex smooth; sides continuously narrow in caudal view, the membranous portion narrowest apically (
|
||
<figureCitation id="136B3EA5FF3CFFDB191B5E9ED7F5FA2B" box="[616,716,1501,1526]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi & Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci & Gamboa, sp. n.; 160: Z. kovacsi Murányi & Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="52" pageNumber="53">Fig. 164</figureCitation>
|
||
). Tg 10, B-scl and Lb-scl generotypic. Ep-scl narrow and acute in dorsal view, short, medially not swollen, its medial width is ~½ of basal width; tip slightly downcurved in lateral view, divided section long. Ventral membranous portion between the division of Ep-scl reaching base in lateral view; apical spines moderatelly long, extending to the Ep-scl (
|
||
<figureCitation id="136B3EA5FF3CFFDB18045D05D6DCF982" box="[887,997,1606,1631]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi & Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="52" pageNumber="53">Figs. 113</figureCitation>
|
||
,
|
||
<figureCitation id="136B3EA5FF3CFFDB18865D05D11AF982" box="[1013,1059,1606,1631]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi & Orci, sp. n.; 131–132: Z. kovacsi Murányi & Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci & Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="52" pageNumber="53">147</figureCitation>
|
||
). I-scl generotypic, Ec long and contorted on the available specimens. St 9 not projecting medially, vesicle vestigial. Sg wide and rounded not as pronounced triangular shape, tip rounded. Pp, Fp, Rp and cerci generotypic.
|
||
</paragraph>
|
||
<paragraph id="8BEF2220FF3CFFDB1BB45DF3D487F8D2" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
Female subgenital plate: Rectangular, posterior margin rounded, equal to the segment`s posterior margin. Antero-lateral recess distinct, the plate brown, sometimes with paler lateral areas; lateral sclerites small (see
|
||
<bibRefCitation id="EFC15FD1FF3CFFDB1BE45DB5D47CF8D2" author="Zhiltzova, L. A." box="[151,325,1782,1807]" pageId="52" pageNumber="53" pagination="1 - 538" refId="ref46559" refString="Zhiltzova, L. A. (2003) Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538." type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
|
||
: Fig. 585).
|
||
</paragraph>
|
||
<paragraph id="8BEF2220FF3CFFDB1BB45C59D1B2F8A4" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
<emphasis id="B924FE32FF3CFFDB1BB45C59D405F8EC" bold="true" box="[199,316,1818,1843]" pageId="52" pageNumber="53">Affinities.</emphasis>
|
||
The relationship of
|
||
<taxonomicName id="4C5059A3FF3CFFDB19515C58D7F2F8EE" authorityName="Kimmins" authorityYear="1950" box="[546,715,1818,1843]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="turkestanica">
|
||
<emphasis id="B924FE32FF3CFFDB19515C58D7F2F8EE" box="[546,715,1818,1843]" italics="true" pageId="52" pageNumber="53">Z. turkestanica</emphasis>
|
||
</taxonomicName>
|
||
with closely related
|
||
<taxonomicName id="4C5059A3FF3CFFDB18CB5C58D16FF8EE" authorityName="Murányi & Gamboa & Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[952,1110,1818,1843]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tuberculata">
|
||
<emphasis id="B924FE32FF3CFFDB18CB5C58D16FF8EE" box="[952,1110,1818,1843]" italics="true" pageId="52" pageNumber="53">Z. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5059A3FF3CFFDB1FFD5C58D037F8EF" authorityName="Murányi & Gamboa & Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[1166,1294,1819,1842]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sevanica">
|
||
<emphasis id="B924FE32FF3CFFDB1FFD5C58D037F8EF" box="[1166,1294,1819,1842]" italics="true" pageId="52" pageNumber="53">Z. sevanica</emphasis>
|
||
</taxonomicName>
|
||
is discussed above. Males are easily distinguishable from all other
|
||
<taxonomicName id="4C5059A3FF3CFFDB18615C7ED6EDF88B" box="[786,980,1853,1878]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B924FE32FF3CFFDB18615C7ED641F88B" box="[786,888,1853,1878]" italics="true" pageId="52" pageNumber="53">Zwicknia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by high, but rounded process of Tg 9. Females are difficult to distinguish from other species of the genus. The larva is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A71ABFF3CFFDB1BB45CC0D6A6F83F" pageId="52" pageNumber="53" type="distribution">
|
||
<paragraph id="8BEF2220FF3CFFDB1BB45CC0D6A6F83F" blockId="52.[151,1437,1078,2019]" pageId="52" pageNumber="53">
|
||
<emphasis id="B924FE32FF3CFFDB1BB45CC0D4C3F841" bold="true" box="[199,506,1923,1948]" pageId="52" pageNumber="53">Distribution and ecology.</emphasis>
|
||
The species was described from a single male from`Turkestan` without exact locality. It is known from
|
||
<collectingCountry id="F34762B0FF3CFFDB1ABA5CE5D703F862" box="[457,570,1958,1983]" name="Tajikistan" pageId="52" pageNumber="53">Tajikistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34762B0FF3CFFDB193B5CE4D7F0F862" box="[584,713,1959,1983]" name="Kyrgyzstan" pageId="52" pageNumber="53">Kyrgyzstan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F34762B0FF3CFFDB18755CE5D6B0F862" box="[774,905,1958,1983]" name="Kazakhstan" pageId="52" pageNumber="53">Kazakhstan</collectingCountry>
|
||
(
|
||
<figureCitation id="136B3EA5FF3CFFDB18E85CE5D139F862" box="[923,1024,1958,1983]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="52" pageNumber="53">Fig. 196</figureCitation>
|
||
). Adults apparently are active from March to April associated with mountain streams as high as
|
||
<quantity id="4CA88FC5FF3CFFDB18485C89D6A3F83F" box="[827,922,1994,2019]" metricMagnitude="0" metricUnit="m" metricValue="2.4" pageId="52" pageNumber="53" unit="m" value="2.4">2,400 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |