220 lines
32 KiB
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220 lines
32 KiB
XML
<document id="5F2187B9CA82992C933A5473FEF846E8" ID-DOI="10.37520/aemnp.2021.009" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037330" ID-ZooBank="27EA8466-6B29-40F4-8236-DD49061C523B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624895377019" checkinUser="carolina" docAuthor="Liu, Tian-tian, Ono, Hiroki & Maruyama, Munetoshi" docDate="2021" docId="03CB023BAC1FFF9A4C9AFC66155FFC18" docLanguage="en" docName="ActaEntMusNatPra.61.1.163-201.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.009" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Bryothinusa aikoae Liu & Ono & Maruyama 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="187" masterDocId="FFF27A43AC0AFF824F48FFFB175FFFCD" masterDocTitle="Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="201" masterPageNumber="163" pageNumber="184" updateTime="1698977316832" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="60BB9E04DE1D0177280B21C627BFCD04">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="35A118CB691D9D4DD3354786763F4833">Liu, Tian-tian</mods:namePart>
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<mods:affiliation id="98FC8E90A59E2B8722EF500687E34A58">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395, Japan & The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:roleTerm id="0958D8DC1D94CBF38F7FC7854E63D1CE">Author</mods:roleTerm>
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<mods:namePart id="42B727BAE3B1E568A0811F73BC0C96BD">Ono, Hiroki</mods:namePart>
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<mods:affiliation id="92FA8BF2C13707BF53A26418AAB12565">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan</mods:affiliation>
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<mods:roleTerm id="A900AC3FFDF87EC939DBAFC6DB24A230">Author</mods:roleTerm>
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<mods:namePart id="E298F69F6D2D2710C22FC727DA9D6FF9">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="EF04DF089CC72549F1EAF845AB1F33E6">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:nameIdentifier id="C4C9DA2BAA8364F0B93ADE763A8F1093" type="email">dendrolasius@gmail.com</mods:nameIdentifier>
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<mods:title id="582FDD332C17D5D35A3AAEE8611961F9">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="E81073A6581D3E3926916F3740D50279">2021</mods:date>
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<mods:title id="81D5E99E399384C020CA915E95BED6A7">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="EA9AAAE0EC2E3E6E10534C8592EB49E5">2021-05-07</mods:number>
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<mods:number id="BE642A5E960C137C8419BE757C479A71">61</mods:number>
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<treatment id="03CB023BAC1FFF9A4C9AFC66155FFC18" LSID="urn:lsid:plazi:treatment:03CB023BAC1FFF9A4C9AFC66155FFC18" httpUri="http://treatment.plazi.org/id/03CB023BAC1FFF9A4C9AFC66155FFC18" lastPageId="24" lastPageNumber="187" pageId="21" pageNumber="184">
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<subSubSection id="C378E0A6AC1FFF974C9AFC661259FC78" box="[978,1286,925,949]" pageId="21" pageNumber="184" type="nomenclature">
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<paragraph id="8BDDB32DAC1FFF974C9AFC661259FC78" blockId="21.[978,1286,925,949]" box="[978,1286,925,949]" pageId="21" pageNumber="184">
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<heading id="D0950441AC1FFF974C9AFC661259FC78" box="[978,1286,925,949]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="184" reason="3">
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<emphasis id="B9166F3FAC1FFF974C9AFC661259FC78" bold="true" box="[978,1286,925,949]" pageId="21" pageNumber="184">
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<taxonomicName id="4C62C8AEAC1FFF974C9AFC6613F5FC78" ID-CoL="8XW4F" authority="Liu & Ono & Maruyama, 2021" authorityName="Liu & Ono & Maruyama" authorityYear="2021" box="[978,1194,925,949]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="184" phylum="Arthropoda" rank="species" species="aikoae" status="sp. nov.">
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<emphasis id="B9166F3FAC1FFF974C9AFC6613F5FC78" bold="true" box="[978,1194,925,949]" italics="true" pageId="21" pageNumber="184">Bryothinusa aikoae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A225D244AC1FFF974BF8FC651259FC78" box="[1200,1286,926,949]" pageId="21" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C378E0A6AC1FFF9A4CB2FC4715EDFE38" lastPageId="24" lastPageNumber="187" pageId="21" pageNumber="184" type="description">
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<paragraph id="8BDDB32DAC1FFF974CB2FC471382FC02" blockId="21.[828,1439,956,1124]" box="[1018,1245,956,975]" pageId="21" pageNumber="184">
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(
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<figureCitation id="1359AFA8AC1FFF974B48FC47131AFC02" box="[1024,1093,956,975]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="21" pageNumber="184">Figs 1L</figureCitation>
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;
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<figureCitation id="1359AFA8AC1FFF974B07FC471336FC02" box="[1103,1129,956,975]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="21" pageNumber="184">2L</figureCitation>
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;
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<figureCitation id="1359AFA8AC1FFF974B3AFC4713D6FC02" box="[1138,1161,956,975]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">14</figureCitation>
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;
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<figureCitation id="1359AFA8AC1FFF974BDAFC4713E7FC02" box="[1170,1208,956,975]" captionStart="Fig" captionStartId="34.[151,182,1955,1974]" captionTargetBox="[167,1408,193,1935]" captionTargetId="figure-13@34.[161,1426,189,1938]" captionTargetPageId="34" captionText="Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka." figureDoi="http://doi.org/10.5281/zenodo.5037374" httpUri="https://zenodo.org/record/5037374/files/figure.png" pageId="21" pageNumber="184">21B</figureCitation>
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;
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<figureCitation id="1359AFA8AC1FFF974B89FC471388FC02" box="[1217,1239,956,975]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="21" pageNumber="184">22</figureCitation>
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)
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</paragraph>
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<paragraph id="8BDDB32DAC1FFF974CF8FC2C13DCFBA9" blockId="21.[828,1439,956,1124]" pageId="21" pageNumber="184">
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(Japanese name: Hime-nagisa-hanekakushi)
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<emphasis id="B9166F3FAC1FFF974C74FBFA14E3FBD9" bold="true" box="[828,956,1025,1044]" pageId="21" pageNumber="184">Type material.</emphasis>
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Hඈඅඈඍඒඉൾ:
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<specimenCount id="9D6478A4AC1FFF974B68FC01136FFBDE" box="[1056,1072,1018,1043]" pageId="21" pageNumber="184" type="female">♀</specimenCount>
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,
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<emphasis id="B9166F3FAC1FFF974B71FBFA1385FBD9" bold="true" box="[1081,1242,1025,1044]" pageId="21" pageNumber="184">
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<collectingCountry id="F375F3BDAC1FFF974B71FBFA13DFFBD9" box="[1081,1152,1025,1044]" name="Japan" pageId="21" pageNumber="184">JAPAN</collectingCountry>
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: HඈඇඌΗǕ:
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</emphasis>
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Mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken,
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<date id="FFDC95EDAC1FFF974B1BFBE013EFFBE3" box="[1107,1200,1051,1070]" pageId="21" pageNumber="184" value="2009-03-12">12.iii.2009</date>
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, H. Ono (KUM).Pൺ*©ൺඍඒඉൾඌ:
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<emphasis id="B9166F3FAC1FFF974C74FBCD14CFFB84" bold="true" box="[828,912,1078,1097]" pageId="21" pageNumber="184">HඈඇඌΗǕ:</emphasis>
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<specimenCount id="9D6478A4AC1FFF974CDEFBCD1499FB84" box="[918,966,1072,1097]" pageId="21" pageNumber="184" type="female">3 ♀♀</specimenCount>
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, same data as
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<typeStatus id="54D90D8FAC1FFF974B0FFBCD13CDFB84" box="[1095,1170,1078,1097]" pageId="21" pageNumber="184" type="holotype">holotype</typeStatus>
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(KUM);
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<specimenCount id="9D6478A4AC1FFF974BADFBCD1259FB84" box="[1253,1286,1072,1097]" pageId="21" pageNumber="184" type="female">1 ♀</specimenCount>
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, same locality as
|
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<typeStatus id="54D90D8FAC1FFF974C74FBAA14D6FBA9" box="[828,905,1105,1124]" pageId="21" pageNumber="184" type="holotype">holotype</typeStatus>
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,
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<date id="FFDC95EDAC1FFF974CD9FBAA14B0FBA9" box="[913,1007,1105,1124]" pageId="21" pageNumber="184" value="2009-03-16">16.iii.2009</date>
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, H. Ono (KUM).
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</paragraph>
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<paragraph id="8BDDB32DAC1FFF974C74FB871270FA9E" blockId="21.[828,1437,1148,2067]" pageId="21" pageNumber="184">
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<emphasis id="B9166F3FAC1FFF974C74FB871496FB5E" bold="true" box="[828,969,1148,1171]" pageId="21" pageNumber="184">Description.</emphasis>
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Body (
|
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<figureCitation id="1359AFA8AC1FFF974B57FB87132CFB5E" box="[1055,1139,1148,1171]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="21" pageNumber="184">Fig. 1L</figureCitation>
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) small, about
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<quantity id="4C9A1EC8AC1FFF974A5FFB8712C4FB5E" box="[1303,1435,1148,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="21" pageNumber="184" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
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in length, covered with minute setae. Basically brown, head darker than pronotum and elytra, pronotum and elytra similarly coloured, from lower part of posteriorly abdominal tergite V to upper part of tergite VII infuscate; antenna, mouthparts and legs similarly coloured, lightly paler than head colour and paler toward apex.
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</paragraph>
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<paragraph id="8BDDB32DAC1FFF974C12FAA714F6F9BE" blockId="21.[828,1437,1148,2067]" pageId="21" pageNumber="184">
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Head oval, wider than long, shorter than elytra; eyes (
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<figureCitation id="1359AFA8AC1FFF974C0CFA8714CAFA5E" box="[836,917,1404,1427]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="21" pageNumber="184">Fig. 2L</figureCitation>
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) large, reniform, longer than 1/2 of head length, with 22–24 facets; antennal length little shorter than distance from anterior margin of head to posterior margin of elytra; segment I 1.4 times longer than II, almost 3 times longer than III, and longer than XI; antennal segments I–III and XI longer than wide, antennal segments IV–VI almost as long as wide, antennal segments VII–X wider than long.
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||
</paragraph>
|
||
<paragraph id="8BDDB32DAC1FFF974C12F98712D6F83E" blockId="21.[828,1437,1148,2067]" pageId="21" pageNumber="184">
|
||
Mouthparts (
|
||
<figureCitation id="1359AFA8AC1FFF974CAFF9871331F95E" box="[999,1134,1660,1683]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">Figs 14A–E</figureCitation>
|
||
) with distribution of labral setae as in
|
||
<figureCitation id="1359AFA8AC1FFF974CE4F9671379F97E" box="[940,1062,1692,1715]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">Figure 14A</figureCitation>
|
||
, with large and thick spiniform seta at antero-laterally; mandibles (
|
||
<figureCitation id="1359AFA8AC1FFF974BC3F94713B2F91E" box="[1163,1261,1724,1747]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">Fig. 14B</figureCitation>
|
||
) elongate triangle, asymmetrical, right with median tooth; median tooth shorter than 1/4 of mandibles; left without tooth, blunt at apex; maxilla with apical apex of lacinia (
|
||
<figureCitation id="1359AFA8AC1FFF974BB6F8E71201F8FE" box="[1278,1374,1820,1843]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">Fig. 14C</figureCitation>
|
||
) with second spines small, at tip with two spines not closely interdigitating; 3 setae on inner margin of lacinia; labial palpus (
|
||
<figureCitation id="1359AFA8AC1FFF974CC5F88714B4F85E" box="[909,1003,1916,1939]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">Fig. 14D</figureCitation>
|
||
) with most setae located basally; mentum (
|
||
<figureCitation id="1359AFA8AC1FFF974C0BF86714C1F87E" box="[835,926,1948,1971]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="21" pageNumber="184">Fig. 14E</figureCitation>
|
||
) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “v seta” of mentum slightly lower than level of anterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC1FFF974C12F8071214F7DE" blockId="21.[828,1437,1148,2067]" box="[858,1355,2044,2067]" pageId="21" pageNumber="184">Pronotum shorter than elytra, wider than head.</paragraph>
|
||
<caption id="DF1DE3A5AC1CFF944FDFF8161267F7D7" ID-DOI="http://doi.org/10.5281/zenodo.5037358" ID-Zenodo-Dep="5037358" httpUri="https://zenodo.org/record/5037358/files/figure.png" pageId="22" pageNumber="185" startId="22.[151,182,2029,2048]" targetBox="[151,1436,191,1993]" targetPageId="22">
|
||
<paragraph id="8BDDB32DAC1CFF944FDFF8161267F7D7" blockId="22.[151,1437,2028,2074]" pageId="22" pageNumber="185">
|
||
Fig. 13.
|
||
<taxonomicName id="4C62C8AEAC1CFF944F97F81716DCF832" authority="Liu & Ono & Maruyama, 2021" authorityName="Liu & Ono & Maruyama" authorityYear="2021" box="[223,387,2028,2047]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="185" phylum="Arthropoda" rank="species" species="itsuroi" status="sp. nov.">
|
||
<emphasis id="B9166F3FAC1CFF944F97F81716DCF832" box="[223,387,2028,2047]" italics="true" pageId="22" pageNumber="185">Bryothinusa itsuroi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225D244AC1CFF944EC1F8161693F7CD" box="[393,460,2029,2048]" pageId="22" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1DE3A5AC1DFF954FDFF80F1684F7EF" ID-DOI="http://doi.org/10.5281/zenodo.5037360" ID-Zenodo-Dep="5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="23" pageNumber="186" startId="23.[151,182,2036,2055]" targetBox="[160,1417,189,1979]" targetPageId="23">
|
||
<paragraph id="8BDDB32DAC1DFF954FDFF80F1684F7EF" blockId="23.[151,1437,2036,2082]" pageId="23" pageNumber="186">
|
||
Fig. 14.
|
||
<taxonomicName id="4C62C8AEAC1DFF954F95F80F16DCF7CA" authority="Liu & Ono & Maruyama, 2021" authorityName="Liu & Ono & Maruyama" authorityYear="2021" box="[221,387,2036,2055]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="186" phylum="Arthropoda" rank="species" species="aikoae" status="sp. nov.">
|
||
<emphasis id="B9166F3FAC1DFF954F95F80F16DCF7CA" box="[221,387,2036,2055]" italics="true" pageId="23" pageNumber="186">Bryothinusa aikoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225D244AC1DFF954EC0F80F1694F7CA" box="[392,459,2036,2055]" pageId="23" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDDB32DAC12FF9A4FFDFF4516D3FF38" blockId="24.[151,761,190,981]" pageId="24" pageNumber="187">Elytra subquadrate, almost as wide as pronotum; hind wings fully developed.</paragraph>
|
||
<paragraph id="8BDDB32DAC12FF9A4FFDFF051601FED8" blockId="24.[151,761,190,981]" box="[181,350,254,277]" pageId="24" pageNumber="187">Male unknown.</paragraph>
|
||
<paragraph id="8BDDB32DAC12FF9A4FFDFEE516F8FEB8" blockId="24.[151,761,190,981]" pageId="24" pageNumber="187">
|
||
Female abdominal sternite VIII in
|
||
<figureCitation id="1359AFA8AC12FF9A4D7FFEE515EEFEF8" box="[567,689,286,309]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="24" pageNumber="187">
|
||
Figure
|
||
<specimenCount id="9D6478A4AC12FF9A4DCEFEE515EEFEF8" box="[646,689,286,309]" pageId="24" pageNumber="187" type="generic">14F</specimenCount>
|
||
</figureCitation>
|
||
. Spermatheca (
|
||
<figureCitation id="1359AFA8AC12FF9A4FB5FEC51602FE98" box="[253,349,318,341]" captionStart="Fig" captionStartId="23.[151,182,2036,2055]" captionTargetBox="[160,1417,189,1979]" captionTargetId="figure-14@23.[159,1427,189,1980]" captionTargetPageId="23" captionText="Fig. 14. Bryothinusa aikoae sp. nov.A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – female sternite VIII; G – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037360" httpUri="https://zenodo.org/record/5037360/files/figure.png" pageId="24" pageNumber="187">Fig. 14G</figureCitation>
|
||
) very small, simplified in shape, about one time coiled near base.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC12FF9A4FDFFE8515EDFE38" blockId="24.[151,761,190,981]" pageId="24" pageNumber="187">
|
||
<emphasis id="B9166F3FAC12FF9A4FDFFE85161EFE58" bold="true" box="[151,321,382,405]" pageId="24" pageNumber="187">Measurements.</emphasis>
|
||
BL 1.50–1.80, FBL 0.83–0.85, HL 0.23– 0.28, HW 0.31–0.32, PL 0.29–0.30, PW 0.34–0.35, EL 0.35–0.36, EW 0.35–0.36, HTL 0.28–0.30, AL 0.58–0.63, AS-I 0.10–0.11, AS-II 0.07–0.08, AS-III 0.03–0.04.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC12FF9A4FDFFE05152FFDB8" pageId="24" pageNumber="187" type="diagnosis">
|
||
<paragraph id="8BDDB32DAC12FF9A4FDFFE05152FFDB8" blockId="24.[151,761,190,981]" pageId="24" pageNumber="187">
|
||
<emphasis id="B9166F3FAC12FF9A4FDFFE0516D7FDD8" bold="true" box="[151,392,510,533]" pageId="24" pageNumber="187">Differential diagnosis.</emphasis>
|
||
The species is distinct based on the body colour and shape. In body size this species is close to
|
||
<taxonomicName id="4C62C8AEAC12FF9A4FDFFDC517A3FD98" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[151,252,574,597]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="187" phylum="Arthropoda" rank="species" species="minuta">
|
||
<emphasis id="B9166F3FAC12FF9A4FDFFDC517A3FD98" box="[151,252,574,597]" italics="true" pageId="24" pageNumber="187">B. minuta</emphasis>
|
||
</taxonomicName>
|
||
but easily distinguished from it by the number of facets and by the fully developed hind wings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC12FF9A4FDFFD851682FD18" pageId="24" pageNumber="187" type="etymology">
|
||
<paragraph id="8BDDB32DAC12FF9A4FDFFD851682FD18" blockId="24.[151,761,190,981]" pageId="24" pageNumber="187">
|
||
<emphasis id="B9166F3FAC12FF9A4FDFFD85164BFD58" bold="true" box="[151,276,638,661]" pageId="24" pageNumber="187">Etymology.</emphasis>
|
||
This new species is named after Mrs.Aiko Ono for supporting the entomological life of her husband H. Ono, the second author of this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC12FF9A4FDFFD251505FD38" box="[151,602,734,757]" pageId="24" pageNumber="187" type="distribution">
|
||
<paragraph id="8BDDB32DAC12FF9A4FDFFD251505FD38" blockId="24.[151,761,190,981]" box="[151,602,734,757]" pageId="24" pageNumber="187">
|
||
<emphasis id="B9166F3FAC12FF9A4FDFFD25167AFD38" bold="true" box="[151,293,734,757]" pageId="24" pageNumber="187">Distribution.</emphasis>
|
||
<collectingCountry id="F375F3BDAC12FF9A4E64FD251637FD38" box="[300,360,734,757]" name="Japan" pageId="24" pageNumber="187">Japan</collectingCountry>
|
||
(
|
||
<emphasis id="B9166F3FAC12FF9A4E3EFD251688FD38" bold="true" box="[374,471,734,757]" pageId="24" pageNumber="187">Honshû:</emphasis>
|
||
Chiba-ken).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC12FF9A4FDFFD05155FFC18" pageId="24" pageNumber="187" type="discussion">
|
||
<paragraph id="8BDDB32DAC12FF9A4FDFFD05155FFC18" blockId="24.[151,761,190,981]" pageId="24" pageNumber="187">
|
||
<emphasis id="B9166F3FAC12FF9A4FDFFD05165BFCD8" bold="true" box="[151,260,766,789]" pageId="24" pageNumber="187">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54D90D8FAC12FF9A4E76FD051634FCD8" box="[318,363,766,789]" pageId="24" pageNumber="187">type</typeStatus>
|
||
series consists of only female specimens. The second author repeatedly searched for this species in the
|
||
<typeStatus id="54D90D8FAC12FF9A4F9CFCC5165EFC98" box="[212,257,830,853]" pageId="24" pageNumber="187">type</typeStatus>
|
||
locality, but no additional specimens were found. Nevertheless, we describe it because of its very distinct general appearance. This species was only found in Chiba-ken. A distribution map of examined specimens of this species in
|
||
<collectingCountry id="F375F3BDAC12FF9A4FFAFC4517B1FC18" box="[178,238,958,981]" name="Japan" pageId="24" pageNumber="187">Japan</collectingCountry>
|
||
is illustrated in
|
||
<figureCitation id="1359AFA8AC12FF9A4EDCFC4516A4FC18" box="[404,507,958,981]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="24" pageNumber="187">Figure 22</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |