395 lines
60 KiB
XML
395 lines
60 KiB
XML
<document id="86EA4ED5AACBD1909DF8AAB41B76E656" ID-CLB-Dataset="57601" ID-DOI="10.11646/zootaxa.5278.2.1" ID-GBIF-Dataset="2d4bcf32-8354-41cd-a2fd-05e47458c178" ID-ISSN="1175-5326" ID-Zenodo-Dep="7905995" ID-ZooBank="697AC989-141D-4A89-8F02-5A4644E303EA" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683528451479" checkinUser="plazi" docAuthor="Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak & Toda, Masanori J." docDate="2023" docId="03C687D9FFC1FFF8A5C6FAA9820BE1F9" docLanguage="en" docName="zootaxa.5278.2.1.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colocasiomyia luciphila Zhang & Gao & Takano & Yafuso & Suwito & Meleng & Toda 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="225" masterDocId="FFFFFFA1FFD6FFE0A551FFBF817EE53C" masterDocTitle="Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species" masterLastPageNumber="238" masterPageNumber="201" pageNumber="224" updateTime="1699515332173" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C60D879A0A94215F4720060A64D94AFE">Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species</mods:title>
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<mods:namePart id="7899246CE81543C471E92357DB0121B3">Zhang, Guang</mods:namePart>
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<mods:namePart id="DDDB0C751F0D13531662E8E8566C34A0">Gao, Jian-Jun</mods:namePart>
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<mods:affiliation id="055260E36230258410351BCFBE1AF682">Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology (Institute of Biodiversity), Yunnan University, Kunming, 650091 China & gaojj @ ynu. edu. cn; https: // orcid. org / 0000 - 0003 - 4636 - 2130</mods:affiliation>
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<mods:nameIdentifier id="8F71DAA4F366FEE0E81E17793CFB4BAF" type="email">gaojj@ynu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="3BDB4515B7DD2BA5D756D2F48EF9B3EC">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="298A595C4F3BDC5769603161DD6130DB">Yafuso, Masako</mods:namePart>
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<mods:affiliation id="8DA970C036394E6D139C84766F969996">Uema 1 - 2 - 21, Naha, Okinawa 902 - 0073, Japan. yafusom @ juno. ocn. ne. jp; https: // orcid. org / 0000 - 0001 - 9737 - 7792</mods:affiliation>
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<mods:namePart id="8DEEE67FF56CBF28D7EA262C32AFA56C">Suwito, Awit</mods:namePart>
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<mods:affiliation id="10639F4B87393A4DD1C9418416C7518E">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. sungaicola @ gmail. com; https: // orcid. org / 0000 - 0003 - 2600 - 2488</mods:affiliation>
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<mods:affiliation id="F957415D08B0D19126D3F00EDB2A6EEA">Research Development and Innovation Division, Forest Department Sarawak, KM 10, Jalan Datuk Amar Kalong Ningkan, 93250, Kuching, Sarawak, Malaysia. paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597</mods:affiliation>
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<mods:date id="7E90DDB739CFC9FEE7471E3BE91632A6">2023</mods:date>
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<treatment id="03C687D9FFC1FFF8A5C6FAA9820BE1F9" ID-DOI="http://doi.org/10.5281/zenodo.7907980" ID-GBIF-Taxon="209588450" ID-Zenodo-Dep="7907980" LSID="urn:lsid:plazi:treatment:03C687D9FFC1FFF8A5C6FAA9820BE1F9" httpUri="http://treatment.plazi.org/id/03C687D9FFC1FFF8A5C6FAA9820BE1F9" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224">
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3)
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<taxonomicName id="4C6F4D4CFFC1FFF7A596FAA983D5E00D" ID-CoL="BDNM8" authority="Zhang & Toda" authorityName="Zhang & Gao & Takano & Yafuso & Suwito & Meleng & Toda" authorityYear="2023" box="[199,683,1302,1329]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="luciphila" status="sp. nov.">
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<emphasis id="B91BEADDFFC1FFF7A596FAA98092E00D" bold="true" box="[199,492,1302,1329]" italics="true" pageId="23" pageNumber="224">Colocasiomyia luciphila</emphasis>
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Zhang & Toda
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,
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<taxonomicNameLabel id="A22857A6FFC1FFF7A7E9FAA88268E00D" box="[696,790,1303,1329]" pageId="23" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3756544FFC1FFF7A5C6FA848016E069" box="[151,360,1339,1365]" pageId="23" pageNumber="224" type="description">
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<paragraph id="8BD036CFFFC1FFF7A5C6FA848016E069" blockId="23.[151,790,1302,1365]" box="[151,360,1339,1365]" pageId="23" pageNumber="224">
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(
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<figureCitation id="13542A4AFFC1FFF7A5CEFA84805BE069" box="[159,293,1339,1365]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStart-9="FIGURE 12" captionStartId-0="4.[151,250,1939,1963]" captionStartId-1="5.[151,250,1430,1454]" captionStartId-2="6.[151,250,1863,1887]" captionStartId-3="7.[151,250,1814,1838]" captionStartId-4="8.[151,250,1921,1945]" captionStartId-5="9.[151,250,1213,1237]" captionStartId-6="10.[151,250,1943,1967]" captionStartId-7="11.[151,250,1926,1950]" captionStartId-8="12.[151,250,1971,1995]" captionStartId-9="16.[151,250,1932,1956]" captionTargetBox-0="[151,1433,634,1915]" captionTargetBox-1="[151,1435,181,1406]" captionTargetBox-2="[162,1420,181,1839]" captionTargetBox-3="[151,1435,181,1789]" captionTargetBox-4="[151,1436,885,1898]" captionTargetBox-5="[151,1435,309,1190]" captionTargetBox-6="[190,1397,181,1919]" captionTargetBox-7="[260,1328,181,1900]" captionTargetBox-8="[155,1436,597,1948]" captionTargetBox-9="[169,1419,420,1909]" captionTargetId-0="figure-203@4.[151,1436,633,1915]" captionTargetId-1="figure-20@5.[151,1436,181,1406]" captionTargetId-2="figure-19@6.[157,1429,181,1839]" captionTargetId-3="figure-20@7.[151,1436,181,1790]" captionTargetId-4="figure-310@8.[151,1436,885,1898]" captionTargetId-5="figure-91@9.[151,1436,309,1190]" captionTargetId-6="figure-19@10.[190,1397,181,1919]" captionTargetId-7="figure-20@11.[260,1328,181,1901]" captionTargetId-8="figure-179@12.[151,1436,597,1948]" captionTargetId-9="figure-108@16.[169,1419,417,1909]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="16" captionText-0="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." captionText-1="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." captionText-2="FIGURE 4. Head parts (caudal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: dl a, dorsolateral tentorial apodeme; dm a, dorsomedial tentorial apodeme; i ocp s, inner occipital seta; o vt s, outer vertical seta; spc s, supracervical seta. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." captionText-3="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." captionText-4="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." captionText-5="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." captionText-6="FIGURE 8. Right wings of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, anterodistal portion (scale bars: 0.1 mm); 2, the whole (scale bars: 0.5 mm). Abbreviations: dm-m, discal-medial crossvein; M4, 4th branch of media." captionText-7="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." captionText-8="FIGURE 10. Abdominal sternites of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female.Abbreviation: s, sternite. Scale bars: 0.1 mm." captionText-9="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906001" figureDoi-1="http://doi.org/10.5281/zenodo.7906005" figureDoi-2="http://doi.org/10.5281/zenodo.7906007" figureDoi-3="http://doi.org/10.5281/zenodo.7906011" figureDoi-4="http://doi.org/10.5281/zenodo.7906013" figureDoi-5="http://doi.org/10.5281/zenodo.7906015" figureDoi-6="http://doi.org/10.5281/zenodo.7906019" figureDoi-7="http://doi.org/10.5281/zenodo.7906021" figureDoi-8="http://doi.org/10.5281/zenodo.7906023" figureDoi-9="http://doi.org/10.5281/zenodo.7906029" httpUri-0="https://zenodo.org/record/7906001/files/figure.png" httpUri-1="https://zenodo.org/record/7906005/files/figure.png" httpUri-2="https://zenodo.org/record/7906007/files/figure.png" httpUri-3="https://zenodo.org/record/7906011/files/figure.png" httpUri-4="https://zenodo.org/record/7906013/files/figure.png" httpUri-5="https://zenodo.org/record/7906015/files/figure.png" httpUri-6="https://zenodo.org/record/7906019/files/figure.png" httpUri-7="https://zenodo.org/record/7906021/files/figure.png" httpUri-8="https://zenodo.org/record/7906023/files/figure.png" httpUri-9="https://zenodo.org/record/7906029/files/figure.png" pageId="23" pageNumber="224">Figs 2–12C</figureCitation>
|
||
,
|
||
<figureCitation id="13542A4AFFC1FFF7A461FA84801EE069" box="[304,352,1339,1365]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="23" pageNumber="224">14C</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFC1FFF7A5C6FACF827FE0B4" box="[151,769,1392,1417]" pageId="23" pageNumber="224" type="reference_group">
|
||
<paragraph id="8BD036CFFFC1FFF7A5C6FACF827FE0B4" blockId="23.[151,769,1392,1417]" box="[151,769,1392,1417]" pageId="23" pageNumber="224">
|
||
<treatmentCitationGroup id="AB7F11E1FFC1FFF7A5C6FACF827FE0B4" box="[151,769,1392,1417]" pageId="23" pageNumber="224">
|
||
<taxonomicName id="4C6F4D4CFFC1FFF7A5C6FACF8382E0B4" ID-CoL="5ZMRT" authority=": Fartyal et al., 2013: 769" authorityName="Fartyal" authorityPageNumber="769" authorityYear="2013" box="[151,764,1392,1417]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="undefined-5">
|
||
<emphasis id="B91BEADDFFC1FFF7A5C6FACF8051E0B4" box="[151,303,1392,1416]" italics="true" pageId="23" pageNumber="224">Colocasiomyia</emphasis>
|
||
sp.5
|
||
<treatmentCitation id="0ACE10DEFFC1FFF7A434FACF8382E0B4" author="Fartyal, R. S. & Gao, J. J. & Toda, M. J. & Hu, Y. G. & Takano, K. T. & Suwito, A. & Katoh, T. & Takigahira, T. & Yin, J. T." box="[357,764,1392,1417]" page="769" pageId="23" pageNumber="224" year="2013">
|
||
aff.
|
||
<emphasis id="B91BEADDFFC1FFF7A4DFFACF8087E0B4" box="[398,505,1392,1416]" italics="true" pageId="23" pageNumber="224">nepalensis</emphasis>
|
||
:
|
||
<bibRefCitation id="EFFE4B3EFFC1FFF7A757FACF8382E0B4" author="Fartyal, R. S. & Gao, J. J. & Toda, M. J. & Hu, Y. G. & Takano, K. T. & Suwito, A. & Katoh, T. & Takigahira, T. & Yin, J. T." box="[518,764,1392,1417]" pageId="23" pageNumber="224" pagination="763 - 782" refId="ref20189" refString="Fartyal, R. S., Gao, J. J., Toda, M. J., Hu, Y. G., Takano, K. T., Suwito, A., Katoh, T., Takigahira, T. & Yin, J. T. (2013) Colocasiomyia (Diptera: Drosophilidae) revised phylogenetically, with a new species group having peculiar lifecycles on monsteroid (Araceae) host plants. Systematic Entomology, 38, 763 - 782. https: // doi. org / 10.1111 / syen. 12027" type="journal article" year="2013">
|
||
Fartyal
|
||
<emphasis id="B91BEADDFFC1FFF7A705FACE83F9E0B4" box="[596,647,1392,1416]" italics="true" pageId="23" pageNumber="224">et al.</emphasis>
|
||
, 2013: 769
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFC1FFF7A5C6FA0985F9E395" pageId="23" pageNumber="224" type="diagnosis">
|
||
<paragraph id="8BD036CFFFC1FFF7A5C6FA0985F9E395" blockId="23.[151,1436,1462,2029]" pageId="23" pageNumber="224">
|
||
<emphasis id="B91BEADDFFC1FFF7A5C6FA09806CE0EC" bold="true" box="[151,274,1462,1488]" pageId="23" pageNumber="224">Diagnosis.</emphasis>
|
||
Antennal, first flagellomere approximately 2.4 times as long as pedicel; arista approximately 1.6 times as long as first flagellomere (
|
||
<figureCitation id="13542A4AFFC1FFF7A4ECFA64836BE0C9" box="[445,533,1499,1525]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,181,1406]" captionTargetId="figure-20@5.[151,1436,181,1406]" captionTargetPageId="5" captionText="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906005" httpUri="https://zenodo.org/record/7906005/files/figure.png" pageId="23" pageNumber="224">Fig. 3C</figureCitation>
|
||
). Fore tarsomere II with 2 pegs; smaller peg slightly longer than 1/2 of larger one (
|
||
<figureCitation id="13542A4AFFC1FFF7A5CEFA40818AE325" box="[159,244,1535,1561]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="23" pageNumber="224">Fig. 9C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A5A5F9AE8182E323" box="[244,252,1553,1567]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A5A5F9AE8182E323" attach="left" box="[244,252,1553,1567]" fontSize="6" pageId="23" pageNumber="224">1</subScript>
|
||
</figureCitation>
|
||
). Male sternites III and IV longer than wide; V+VI concave on distal margin (
|
||
<figureCitation id="13542A4AFFC1FFF7A135FA4085B8E325" box="[1124,1222,1535,1561]" captionStart="FIGURE 10" captionStartId="12.[151,250,1971,1995]" captionTargetBox="[155,1436,597,1948]" captionTargetId="figure-179@12.[151,1436,597,1948]" captionTargetPageId="12" captionText="FIGURE 10. Abdominal sternites of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female.Abbreviation: s, sternite. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906023" httpUri="https://zenodo.org/record/7906023/files/figure.png" pageId="23" pageNumber="224">Fig. 10C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A196F9AE85B1E323" box="[1223,1231,1553,1567]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A196F9AE85B1E323" attach="right" box="[1223,1231,1553,1567]" fontSize="6" pageId="23" pageNumber="224">1</subScript>
|
||
</figureCitation>
|
||
). Female sternites III–VI longer than wide; VII much wider than long, slightly convex on distal margin (
|
||
<figureCitation id="13542A4AFFC1FFF7A122F99C85A6E301" box="[1139,1240,1571,1597]" captionStart="FIGURE 10" captionStartId="12.[151,250,1971,1995]" captionTargetBox="[155,1436,597,1948]" captionTargetId="figure-179@12.[151,1436,597,1948]" captionTargetPageId="12" captionText="FIGURE 10. Abdominal sternites of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female.Abbreviation: s, sternite. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906023" httpUri="https://zenodo.org/record/7906023/files/figure.png" pageId="23" pageNumber="224">Fig. 10C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A189F98A859EE37F" box="[1240,1248,1589,1603]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A189F98A859EE37F" attach="left" box="[1240,1248,1589,1603]" fontSize="6" pageId="23" pageNumber="224">2</subScript>
|
||
</figureCitation>
|
||
). Surstylus with 6–7 stout, longer, peg-like teeth nearly uniform in thickness (Fig. 11-1C
|
||
<subScript id="17EB348AFFC1FFF7A6E2F9E682B9E35B" attach="left" box="[947,967,1625,1639]" fontSize="6" pageId="23" pageNumber="224">1,2</subScript>
|
||
). Pregonites apically roundish, smooth on ventral margin (Fig. 11-1C
|
||
<subScript id="17EB348AFFC1FFF7A495F9C280AAE3B7" attach="left" box="[452,468,1661,1675]" fontSize="6" pageId="23" pageNumber="224">4a</subScript>
|
||
). Epigynium with 2–6 small sensilla per side near lower posterior margin (
|
||
<figureCitation id="13542A4AFFC1FFF7A071F9D484FAE3B9" box="[1312,1412,1643,1669]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="23" pageNumber="224">Fig. 12C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A0D5F9C284F2E3B7" box="[1412,1420,1661,1675]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A0D5F9C284F2E3B7" attach="left" box="[1412,1420,1661,1675]" fontSize="6" pageId="23" pageNumber="224">1</subScript>
|
||
</figureCitation>
|
||
). Distal, narrow portion of hypogynial valve half as long as basal, broad portion (
|
||
<figureCitation id="13542A4AFFC1FFF7A145F9308505E395" box="[1044,1147,1679,1705]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="23" pageNumber="224">Fig. 12C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFC1FFF8A596F90D8554E75D" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224" type="description">
|
||
<paragraph id="8BD036CFFFC1FFF7A596F90D8466E261" blockId="23.[151,1436,1462,2029]" pageId="23" pageNumber="224">
|
||
<emphasis id="B91BEADDFFC1FFF7A596F90D802FE3F0" bold="true" box="[199,337,1714,1740]" pageId="23" pageNumber="224">Description</emphasis>
|
||
(♁ and
|
||
<specimenCount id="9D69FD46FFC1FFF7A4E6F90C80B5E3F1" box="[439,459,1715,1741]" pageId="23" pageNumber="224" type="female">♀</specimenCount>
|
||
). Head. Antennal pedicel grayish brown. Distance between antennal scapes as wide as scape diameter (
|
||
<figureCitation id="13542A4AFFC1FFF7A428F96880A4E3CB" box="[377,474,1751,1783]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="23" pageNumber="224">
|
||
Fig. 2C
|
||
<subScript id="17EB348AFFC1FFF7A483F95680A4E3CB" attach="left" box="[466,474,1769,1783]" fontSize="6" pageId="23" pageNumber="224">2</subScript>
|
||
</figureCitation>
|
||
). Facial carina narrower than 1st flagellomere (
|
||
<figureCitation id="13542A4AFFC1FFF7A15CF9688510E3CB" box="[1037,1134,1751,1783]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="23" pageNumber="224">
|
||
Fig. 2C
|
||
<subScript id="17EB348AFFC1FFF7A137F9568510E3CB" attach="left" box="[1126,1134,1769,1783]" fontSize="6" pageId="23" pageNumber="224">2</subScript>
|
||
</figureCitation>
|
||
). Supracervical setae 6–8 and postoculars 13–18 per side (
|
||
<figureCitation id="13542A4AFFC1FFF7A75CF9448316E229" box="[525,616,1787,1813]" captionStart="FIGURE 4" captionStartId="6.[151,250,1863,1887]" captionTargetBox="[162,1420,181,1839]" captionTargetId="figure-19@6.[157,1429,181,1839]" captionTargetPageId="6" captionText="FIGURE 4. Head parts (caudal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: dl a, dorsolateral tentorial apodeme; dm a, dorsomedial tentorial apodeme; i ocp s, inner occipital seta; o vt s, outer vertical seta; spc s, supracervical seta. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906007" httpUri="https://zenodo.org/record/7906007/files/figure.png" pageId="23" pageNumber="224">Fig. 4C</figureCitation>
|
||
). Cibarial projections at anterolateral corners slightly longer than width of anterior margin; medial sensilla approximately 3–4 per side (
|
||
<figureCitation id="13542A4AFFC1FFF7A6D6F8A0829EE205" box="[903,992,1822,1849]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="23" pageNumber="224">Fig. 5C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A6B1F88E8296E203" box="[992,1000,1841,1855]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A6B1F88E8296E203" attach="left" box="[992,1000,1841,1855]" fontSize="6" pageId="23" pageNumber="224">2</subScript>
|
||
</figureCitation>
|
||
). Prementum and neighboring lateral membrane with 7–8 setae per side (
|
||
<figureCitation id="13542A4AFFC1FFF7A775F8FC8304E261" box="[548,634,1858,1885]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="23" pageNumber="224">Fig. 5C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A728F8EA83FFE25F" box="[633,641,1877,1891]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A728F8EA83FFE25F" attach="left" box="[633,641,1877,1891]" fontSize="6" pageId="23" pageNumber="224">2</subScript>
|
||
</figureCitation>
|
||
). Labellum with 11–13 pseudotracheae per side (
|
||
<figureCitation id="13542A4AFFC1FFF7A1FAF8FC847EE261" box="[1195,1280,1858,1885]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="23" pageNumber="224">Fig. 5C</figureCitation>
|
||
<figureCitation id="13542A4AFFC1FFF7A051F8EA8476E25F" box="[1280,1288,1877,1891]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="23" pageNumber="224">
|
||
<subScript id="17EB348AFFC1FFF7A051F8EA8476E25F" attach="left" box="[1280,1288,1877,1891]" fontSize="6" pageId="23" pageNumber="224">1</subScript>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFC1FFF7A596F8D88256E299" blockId="23.[151,1436,1462,2029]" pageId="23" pageNumber="224">
|
||
Thorax. Prescutellar, acrostichal setae only slightly longer than acrostichal setulae (
|
||
<figureCitation id="13542A4AFFC1FFF7A106F8D885D3E2BD" box="[1111,1197,1895,1921]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="23" pageNumber="224">Fig. 6C</figureCitation>
|
||
). Only one prominent katepisternal seta longer than acrostichal setulae (
|
||
<figureCitation id="13542A4AFFC1FFF7A792F8348265E299" box="[707,795,1931,1957]" captionStart="FIGURE 7" captionStartId="9.[151,250,1213,1237]" captionTargetBox="[151,1435,309,1190]" captionTargetId="figure-91@9.[151,1436,309,1190]" captionTargetPageId="9" captionText="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906015" httpUri="https://zenodo.org/record/7906015/files/figure.png" pageId="23" pageNumber="224">Fig. 7C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFC1FFF7A596F8108244E2D1" blockId="23.[151,1436,1462,2029]" pageId="23" pageNumber="224">
|
||
Legs grayish brown, except for all tarsi pale grayish yellow (
|
||
<figureCitation id="13542A4AFFC1FFF7A63BF81082AFE2F5" box="[874,977,1967,1993]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="23" pageNumber="224">Fig. 14C</figureCitation>
|
||
). Fore, mid and hind tarsomere Is as long as II+III, II+III+IV and II+III+IV+V, respectively (
|
||
<figureCitation id="13542A4AFFC1FFF7A785F86C8253E2D1" box="[724,813,2003,2029]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="23" pageNumber="224">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFCEFFF8A596FF28819EE5E9" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
Abdomen. Tergites dark grayish brown (
|
||
<figureCitation id="13542A4AFFCEFFF8A7CDFF288278E58D" box="[668,774,151,177]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="24" pageNumber="225">Fig. 14C</figureCitation>
|
||
). Female tergite VII mid-dorsally separated into 2 lateral plates.
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFCEFFF8A596FF608040E47D" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
Male terminalia. Epandrium with approximately 4 long setae only on lateral to ventral portion (Fig. 11-1C
|
||
<subScript id="17EB348AFFCEFFF8A0D5FF4E84F2E5C3" attach="left" box="[1412,1420,241,255]" fontSize="6" pageId="24" pageNumber="225">1</subScript>
|
||
). Cercus entirely pubescent, round on ventral margin, with 28–29 setae (Fig. 11-1C
|
||
<subScript id="17EB348AFFCEFFF8A172FEAA8555E41F" attach="left" box="[1059,1067,277,291]" fontSize="6" pageId="24" pageNumber="225">1</subScript>
|
||
). Phallal sheath partly pubescent (Fig. 11-1C
|
||
<subScript id="17EB348AFFCEFFF8A448FE868053E47B" attach="left" box="[281,301,313,327]" fontSize="6" pageId="24" pageNumber="225">4,b</subScript>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFCEFFF8A596FEF48554E459" blockId="24.[151,1437,151,1221]" box="[199,1066,331,357]" pageId="24" pageNumber="225">Female terminalia. Perineal membrane without patch of dense, distinct warts.</paragraph>
|
||
<paragraph id="8BD036CFFFCEFFF8A596FED0851EE491" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
Measurements (
|
||
<typeStatus id="54D4886DFFCEFFF8A429FED080A5E4B5" box="[376,475,367,393]" pageId="24" pageNumber="225" type="holotype">holotype</typeStatus>
|
||
/range in 1♁ and
|
||
<specimenCount id="9D69FD46FFCEFFF8A7CAFED083C0E4B5" box="[667,702,367,393]" pageId="24" pageNumber="225" type="female">3♀</specimenCount>
|
||
<typeStatus id="54D4886DFFCEFFF8A795FED0824CE4B5" box="[708,818,367,393]" pageId="24" pageNumber="225" type="paratype">paratypes</typeStatus>
|
||
, in mm): BL = 1.51/♁ 1.59,
|
||
<specimenCount id="9D69FD46FFCEFFF8A13AFED085FEE4B5" box="[1131,1152,367,393]" pageId="24" pageNumber="225" type="female">♀</specimenCount>
|
||
1.40–1.92; ThL = 0.60/♁ 0.60,
|
||
<specimenCount id="9D69FD46FFCEFFF8A587FE2C8195E491" box="[214,235,403,429]" pageId="24" pageNumber="225" type="female">♀</specimenCount>
|
||
0.48–0.75; WL = 1.22/♁ 1.23,
|
||
<specimenCount id="9D69FD46FFCEFFF8A71CFE2C831CE491" box="[589,610,403,429]" pageId="24" pageNumber="225" type="female">♀</specimenCount>
|
||
1.06–1.55; WW = 0.57/♁ 0.61,
|
||
<specimenCount id="9D69FD46FFCEFFF8A69CFE2C829CE491" box="[973,994,403,429]" pageId="24" pageNumber="225" type="female">♀</specimenCount>
|
||
0.50–0.73.
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFCEFFF8A596FE088554E75D" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
Indices (
|
||
<typeStatus id="54D4886DFFCEFFF8A478FE0880F2E4ED" box="[297,396,439,465]" pageId="24" pageNumber="225" type="holotype">holotype</typeStatus>
|
||
/range in 1♁ and
|
||
<specimenCount id="9D69FD46FFCEFFF8A709FE088302E4ED" box="[600,636,439,465]" pageId="24" pageNumber="225" type="female">3♀</specimenCount>
|
||
, or less if noted,
|
||
<typeStatus id="54D4886DFFCEFFF8A619FE0882C8E4ED" box="[840,950,439,465]" pageId="24" pageNumber="225" type="paratype">paratypes</typeStatus>
|
||
, in ratio): FW/HW = 0.48/0.49–0.55; ch/o = 0.29/0.33–0.38; prorb = 0.86/0.82–1.08; rcorb = 0.32/0.29–0.40; vb = 0.17/0.18–0.25 (
|
||
<specimenCount id="9D69FD46FFCEFFF8A1F0FE6485BDE4C9" box="[1185,1219,475,501]" pageId="24" pageNumber="225" type="female">3♀</specimenCount>
|
||
); dcl = 0.39/0.35– 0.40 (1♁ and
|
||
<specimenCount id="9D69FD46FFCEFFF8A46CFE40801FE725" box="[317,353,511,537]" pageId="24" pageNumber="225" type="female">2♀</specimenCount>
|
||
); presctl = 0.20/0.21–0.25 (1♁ and
|
||
<specimenCount id="9D69FD46FFCEFFF8A65BFE408250E725" box="[778,814,511,537]" pageId="24" pageNumber="225" type="female">2♀</specimenCount>
|
||
); sctl = 0.56/0.52–0.60; orbito = 0.45/0.33–0.47; dcp = 0.54/0.49–0.59; sctlp = 0.93/0.88–1.03; C = 1.57/1.49–1.64; 4c = 1.61/1.39–1.64; 4v = 2.39/2.20–2.63; 5x = 2.61/2.62–2.75; ac = 3.27/2.49–3.09; M = 0.90/0.78–0.92; C3F = 0.60/0.49–0.72.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFCEFFF8A596FDD48084E605" pageId="24" pageNumber="225" type="materials_examined">
|
||
<paragraph id="8BD036CFFFCEFFF8A596FDD48065E795" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
<materialsCitation id="3B073C92FFCEFFF8A596FDD48065E795" ID-GBIF-Occurrence="4101310303" collectingDate="2012-01-25" collectingDateMax="2012-01-26" collectingDateMin="2012-01-25" collectionCode="RDID" collectorName="M. J. Toda" country="Malaysia" location="Lambir Hills National Park" pageId="24" pageNumber="225" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B91BEADDFFCEFFF8A596FDD48045E7B9" bold="true" box="[199,315,619,645]" pageId="24" pageNumber="225">
|
||
<typeStatus id="54D4886DFFCEFFF8A596FDD48048E7B9" box="[199,310,619,645]" pageId="24" pageNumber="225" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
♁ (
|
||
<collectionCode id="ED7EAE0AFFCEFFF8A43AFDD480CBE7B9" box="[363,437,619,645]" pageId="24" pageNumber="225">RDID</collectionCode>
|
||
), “
|
||
<location id="8EB06014FFCEFFF8A485FDD4826CE7B9" LSID="urn:lsid:plazi:treatment:03C687D9FFC1FFF8A5C6FAA9820BE1F9:8EB06014FFCEFFF8A485FDD4826CE7B9" box="[468,786,619,645]" country="Malaysia" name="Lambir Hills National Park" pageId="24" pageNumber="225">Lambir Hills National Park</location>
|
||
, Sarawak,
|
||
<collectingCountry id="F378765FFFCEFFF8A6C1FDD48287E7B9" box="[912,1017,619,645]" name="Malaysia" pageId="24" pageNumber="225">Malaysia</collectingCountry>
|
||
,
|
||
<date id="FFD1100FFFCEFFF8A156FDD585E0E7B9" box="[1031,1182,618,645]" pageId="24" pageNumber="225" value="2012-01-25" valueMax="2012-01-26" valueMin="2012-01-25">
|
||
<collectingDate id="EF95E9E7FFCEFFF8A156FDD585E0E7B9" box="[1031,1182,618,645]" pageId="24" pageNumber="225" value="2012-01-25" valueMax="2012-01-26" valueMin="2012-01-25">25–26.i.2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="522E4ED8FFCEFFF8A1FAFDD48449E7B9" box="[1195,1335,619,645]" pageId="24" pageNumber="225">ex light trap</collectingMethod>
|
||
(LT-35),
|
||
<collectorName id="269A5319FFCEFFF8A5C6FD308072E795" box="[151,268,655,681]" pageId="24" pageNumber="225">M.J. Toda</collectorName>
|
||
”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD036CFFFCEFFF8A596FD0C8084E605" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
<materialsCitation id="3B073C92FFCEFFF8A596FD0C82F5E7F1" ID-GBIF-Occurrence="4101310313" box="[199,907,690,717]" collectingDate="2012-01-25" collectingDateMax="2012-01-26" collectingDateMin="2012-01-25" collectionCode="SEHU" collectorName="M. J. Toda" country="Malaysia" location="Lambir Hills National Park" pageId="24" pageNumber="225" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
|
||
<emphasis id="B91BEADDFFCEFFF8A596FD0C8038E7F1" bold="true" box="[199,326,691,717]" pageId="24" pageNumber="225">
|
||
<typeStatus id="54D4886DFFCEFFF8A596FD0C803FE7F1" box="[199,321,691,717]" pageId="24" pageNumber="225" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="F378765FFFCEFFF8A41EFD0C80C4E7F1" box="[335,442,691,717]" name="Malaysia" pageId="24" pageNumber="225">Malaysia</collectingCountry>
|
||
:
|
||
<specimenCount id="9D69FD46FFCEFFF8A496FD0D8095E7F1" box="[455,491,690,717]" pageId="24" pageNumber="225" type="female">5♀</specimenCount>
|
||
, same data as the holotype (
|
||
<collectionCode id="ED7EAE0AFFCEFFF8A665FD0C82FDE7F1" box="[820,899,691,717]" pageId="24" pageNumber="225">SEHU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D69FD46FFCEFFF8A6C8FD0C82C2E7F1" box="[921,956,691,717]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
|
||
, same except for
|
||
<date id="FFD1100FFFCEFFF8A1DAFD0C845FE7F1" box="[1163,1313,691,717]" pageId="24" pageNumber="225" value="2012-01-26" valueMax="2012-01-27" valueMin="2012-01-26">26–27.i.2012</date>
|
||
,
|
||
<specimenCount id="9D69FD46FFCEFFF8A07EFD0C842CE7F1" box="[1327,1362,691,717]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
|
||
, same except for
|
||
<date id="FFD1100FFFCEFFF8A440FD688003E7CD" box="[273,381,727,753]" pageId="24" pageNumber="225" value="2012-01-23">23.i.2012</date>
|
||
, ex florescent light,
|
||
<materialsCitation id="3B073C92FFCEFFF8A735FD688589E7CD" ID-GBIF-Occurrence="4101310317" box="[612,1271,726,753]" collectingDate="2012-01-25" collectionCode="RDID" country="Malaysia" pageId="24" pageNumber="225" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D69FD46FFCEFFF8A735FD6883F9E7CD" box="[612,647,727,753]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
|
||
, same except for
|
||
<date id="FFD1100FFFCEFFF8A600FD6982C3E7CD" box="[849,957,726,753]" pageId="24" pageNumber="225" value="2012-01-25">
|
||
<collectingDate id="EF95E9E7FFCEFFF8A600FD6982C3E7CD" box="[849,957,726,753]" pageId="24" pageNumber="225" value="2012-01-25">25.i.2012</collectingDate>
|
||
</date>
|
||
, ex florescent light (
|
||
<collectionCode id="ED7EAE0AFFCEFFF8A1F4FD68858EE7CD" box="[1189,1264,727,753]" pageId="24" pageNumber="225">RDID</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B073C92FFCEFFF8A055FD6780C3E629" ID-GBIF-Occurrence="4101310311" collectingDate="2012-01-25" collectingDateMax="2012-01-26" collectingDateMin="2012-01-25" collectionCode="RDID" collectorName="M. J. Toda" country="Malaysia" location="Lambir Hills National Park" pageId="24" pageNumber="225" specimenCount="1" typeStatus="paratype">1♁ (#05225), same data as the holotype</materialsCitation>
|
||
,
|
||
<specimenCount id="9D69FD46FFCEFFF8A498FD448092E629" box="[457,492,763,789]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
|
||
(#05223), same except for
|
||
<date id="FFD1100FFFCEFFF8A674FD4482C7E629" box="[805,953,763,789]" pageId="24" pageNumber="225" value="2012-01-23" valueMax="2012-01-24" valueMin="2012-01-23">23–24.i.2012</date>
|
||
(Lot #229),
|
||
<materialsCitation id="3B073C92FFCEFFF8A118FD448088E605" ID-GBIF-Occurrence="4101310310" collectingDate="2012-01-24" collectingDateMax="2012-01-25" collectingDateMin="2012-01-24" collectionCode="KIZ" country="Malaysia" pageId="24" pageNumber="225" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D69FD46FFCEFFF8A118FD448512E629" box="[1097,1132,763,789]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
|
||
(#05224), same except for
|
||
<date id="FFD1100FFFCEFFF8A5C6FCA18055E605" box="[151,299,798,825]" pageId="24" pageNumber="225" value="2012-01-24" valueMax="2012-01-25" valueMin="2012-01-24">
|
||
<collectingDate id="EF95E9E7FFCEFFF8A5C6FCA18055E605" box="[151,299,798,825]" pageId="24" pageNumber="225" value="2012-01-24" valueMax="2012-01-25" valueMin="2012-01-24">24–25.i.2012</collectingDate>
|
||
</date>
|
||
(Lot #230) (
|
||
<collectionCode id="ED7EAE0AFFCEFFF8A4EBFCA08090E605" box="[442,494,799,825]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="24" pageNumber="225">KIZ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFCEFFF8A596FCFD832EE661" box="[199,592,834,861]" pageId="24" pageNumber="225" type="distribution">
|
||
<paragraph id="8BD036CFFFCEFFF8A596FCFD832EE661" blockId="24.[151,1437,151,1221]" box="[199,592,834,861]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91BEADDFFCEFFF8A596FCFD801FE660" bold="true" box="[199,353,834,860]" pageId="24" pageNumber="225">Distribution.</emphasis>
|
||
<collectingCountry id="F378765FFFCEFFF8A436FCFC80B0E661" box="[359,462,835,861]" name="Malaysia" pageId="24" pageNumber="225">Malaysia</collectingCountry>
|
||
(
|
||
<collectingCountry id="F378765FFFCEFFF8A48CFCFC833AE661" box="[477,580,835,861]" name="Malaysia" pageId="24" pageNumber="225">Sarawak</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFCEFFF8A596FCD88083E19D" pageId="24" pageNumber="225" type="discussion">
|
||
<paragraph id="8BD036CFFFCEFFF8A596FCD88083E19D" blockId="24.[151,1437,151,1221]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91BEADDFFCEFFF8A596FCD88045E6BD" bold="true" box="[199,315,871,897]" pageId="24" pageNumber="225">Remarks.</emphasis>
|
||
This species resembles
|
||
<taxonomicName id="4C6F4D4CFFCEFFF8A70BFCD8838AE6BD" box="[602,756,871,897]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="nepalensis">
|
||
<emphasis id="B91BEADDFFCEFFF8A70BFCD88313E6BD" box="[602,621,871,897]" italics="true" pageId="24" pageNumber="225">C</emphasis>
|
||
.
|
||
<emphasis id="B91BEADDFFCEFFF8A72FFCD8838AE6BD" box="[638,756,871,897]" italics="true" pageId="24" pageNumber="225">nepalensis</emphasis>
|
||
</taxonomicName>
|
||
in having only one prominent, katepisternal seta and two pegs less different in length on the fore tarsomere II, but can be distinguished from it by the arista 1.6 times as long as first flagellomere (in
|
||
<taxonomicName id="4C6F4D4CFFCEFFF8A4B9FC1083FFE6F5" box="[488,641,943,969]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="nepalensis">
|
||
<emphasis id="B91BEADDFFCEFFF8A4B9FC108085E6F5" box="[488,507,943,969]" italics="true" pageId="24" pageNumber="225">C</emphasis>
|
||
.
|
||
<emphasis id="B91BEADDFFCEFFF8A75AFC1083FFE6F5" box="[523,641,943,969]" italics="true" pageId="24" pageNumber="225">nepalensis</emphasis>
|
||
</taxonomicName>
|
||
: “2.5× the length of flagellomere I”;
|
||
<bibRefCitation id="EFFE4B3EFFCEFFF8A162FC1085A2E6F5" author="Grimaldi, D. A." box="[1075,1244,943,969]" pageId="24" pageNumber="225" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
|
||
), cibarial medial sensilla in parallel rows as wide as sensilla campaniformia (narrower than the latter; “
|
||
<figureCitation id="13542A4AFFCEFFF8A11FFC6C85ECE6D1" box="[1102,1170,979,1005]" captionStart="FIGURE 7" captionStartId="9.[151,250,1213,1237]" captionTargetBox="[151,1435,309,1190]" captionTargetId="figure-91@9.[151,1436,309,1190]" captionTargetPageId="9" captionText="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906015" httpUri="https://zenodo.org/record/7906015/files/figure.png" pageId="24" pageNumber="225">Fig. 7</figureCitation>
|
||
” in
|
||
<bibRefCitation id="EFFE4B3EFFCEFFF8A191FC6C8418E6D1" author="Grimaldi, D. A." box="[1216,1382,979,1005]" pageId="24" pageNumber="225" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
|
||
), the facial carina narrower than 1st flagellomere (wider than the latter; “
|
||
<figureCitation id="13542A4AFFCEFFF8A6C7FC4882A4E12D" box="[918,986,1015,1041]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="24" pageNumber="225">Fig. 6</figureCitation>
|
||
” in
|
||
<bibRefCitation id="EFFE4B3EFFCEFFF8A15DFC4885CAE12D" author="Grimaldi, D. A." box="[1036,1204,1015,1041]" pageId="24" pageNumber="225" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
|
||
), and the absence of membranous aedeagus (“Distiphallus fusiform, apically with fine spicules, covered with membrane”). On the other hand, the following characters suggest some relatedness to the
|
||
<taxonomicName id="4C6F4D4CFFCEFFF8A605FB8082A6E165" authorityName="Zhang & Gao & Takano & Yafuso & Suwito & Meleng & Toda" authorityYear="2023" box="[852,984,1087,1113]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="oligochaeta">
|
||
<emphasis id="B91BEADDFFCEFFF8A605FB8082A6E165" box="[852,984,1087,1113]" italics="true" pageId="24" pageNumber="225">oligochaeta</emphasis>
|
||
</taxonomicName>
|
||
species subgroup described below: only one prominent, katepisternal seta; prescutellar, acrostichal setae less differentiated; and male sternites III and IV and female III–VI longer than wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756544FFCEFFF8A596FB14820BE1F9" box="[199,885,1195,1221]" pageId="24" pageNumber="225" type="etymology">
|
||
<paragraph id="8BD036CFFFCEFFF8A596FB14820BE1F9" blockId="24.[151,1437,151,1221]" box="[199,885,1195,1221]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91BEADDFFCEFFF8A596FB148030E1F9" bold="true" box="[199,334,1195,1221]" pageId="24" pageNumber="225">Etymology.</emphasis>
|
||
Referring to its nature of being attracted to light.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |