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<document id="68ECEB289DE203C0540615572D62D1A1" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F06EFFFAFF42FE79FEEDFCAF" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Coprophanaeoides) cribrosus Genier &amp; Arnaud, new species" docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="13" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="116206BDC0103252F12A33620FF6D2F2">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:namePart id="115FFCF091AD1A0125B8437CAFC12961">François Génier</mods:namePart>
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<mods:namePart id="A9372F378A13F95E2EA2FCF8742215C0">Patrick Arnaud</mods:namePart>
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<mods:title id="C11808D6E2B0E22B78532487A868541E">Zootaxa</mods:title>
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<mods:part id="DFD4072A015531799CE5DC7730F734DE">
<mods:date id="18FAED41F922F0B648976D1C670E12FC">2016</mods:date>
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<mods:number id="F83BBBF8DC7AF6A952E5BF9E63B446E1">4099</mods:number>
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<mods:identifier id="163F60FEC1D9763CDF1F96E57F80EB85" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
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<treatment id="03C68782F06EFFFAFF42FE79FEEDFCAF" ID-DOI="http://doi.org/10.5281/zenodo.5780258" ID-GBIF-Taxon="127631977" ID-Zenodo-Dep="5780258" LSID="urn:lsid:plazi:treatment:03C68782F06EFFFAFF42FE79FEEDFCAF" httpUri="http://treatment.plazi.org/id/03C68782F06EFFFAFF42FE79FEEDFCAF" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C375651FF06EFFFBFF42FE79FB8FFE7E" box="[151,1129,404,430]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BD03694F06EFFFBFF42FE79FB8FFE7E" blockId="12.[151,1129,404,463]" box="[151,1129,404,430]" pageId="12" pageNumber="13">
<heading id="D09881F8F06EFFFBFF42FE79FB8FFE7E" bold="true" box="[151,1129,404,430]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B91BEA86F06EFFFBFF42FE79FB8FFE7E" bold="true" box="[151,1129,404,430]" pageId="12" pageNumber="13">
4.
<taxonomicName id="4C6F4D17F06EFFFBFF61FE79FC2AFE7E" ID-CoL="34R2C" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[180,972,404,430]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cribrosus" status="sp. nov." subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F06EFFFBFF61FE79FD02FE7E" bold="true" box="[180,740,404,430]" italics="true" pageId="12" pageNumber="13">Dendropaemon (Coprophanaeoides) cribrosus</emphasis>
Génier &amp; Arnaud
</taxonomicName>
,
<taxonomicNameLabel id="A22857FDF06EFFFBFC0CFE79FB8FFE7E" box="[985,1129,404,430]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06EFFFBFF42FE5BFE67FE1F" box="[151,385,438,463]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BD03694F06EFFFBFF42FE5BFE67FE1F" blockId="12.[151,1129,404,463]" box="[151,385,438,463]" pageId="12" pageNumber="13">
(
<figureCitation id="13542A11F06EFFFBFF4AFE5BFF08FE1F" box="[159,238,438,463]" captionStart="FIGURES 1 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="12" pageNumber="13">Figs. 4</figureCitation>
, 4647, 156)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06EFFFBFF42FE13FC8DFDC7" box="[151,875,510,535]" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BD03694F06EFFFBFF42FE13FC8DFDC7" blockId="12.[151,1437,510,2011]" box="[151,875,510,535]" pageId="12" pageNumber="13">
<emphasis id="B91BEA86F06EFFFBFF42FE13FECBFDC7" bold="true" box="[151,301,510,535]" pageId="12" pageNumber="13">
<typeStatus id="54D48836F06EFFFBFF42FE13FF34FDC7" box="[151,210,510,535]" pageId="12" pageNumber="13">Type</typeStatus>
locality
</emphasis>
. Ubajara (
<geoCoordinate id="EE5B5053F06EFFFBFE71FE13FDE7FDC7" box="[420,513,510,535]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="-3.8333333">03°50S</geoCoordinate>
<geoCoordinate id="EE5B5053F06EFFFBFDDDFE13FD89FDC7" box="[520,623,510,535]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="-40.933334">40°56W</geoCoordinate>
,
<quantity id="4C979B71F06EFFFBFDAFFE12FD59FDC7" box="[634,703,511,535]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="12" pageNumber="13" unit="m" value="820.0">820m</quantity>
), Ceara,
<collectingCountry id="F3787604F06EFFFBFCF4FE13FC81FDC7" box="[801,871,510,535]" name="Brazil" pageId="12" pageNumber="13">Brasil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06EFFFBFF12FDCFFEB5FD1B" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BD03694F06EFFFBFF12FDCFFEB5FD1B" blockId="12.[151,1437,510,2011]" pageId="12" pageNumber="13">
<emphasis id="B91BEA86F06EFFFBFF12FDCFFEA7FDEA" bold="true" box="[199,321,546,571]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Differs from nearly all other species in the genus by its long elytral pilosity combined with the sharply carinate lateral edge of the pronotal lateral depressions. The much less pilose dorsum, especially the nearly glabrous eighth elytral interval will separate
<taxonomicName id="4C6F4D17F06EFFFBFD74FD81FCD6FD53" box="[673,816,618,643]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cribrosus">
<emphasis id="B91BEA86F06EFFFBFD74FD81FCD6FD53" box="[673,816,618,643]" italics="true" pageId="12" pageNumber="13">D. cribrosus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6F4D17F06EFFFBFCAEFD81FBCCFD53" box="[891,1066,618,643]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosissimus">
<emphasis id="B91BEA86F06EFFFBFCAEFD81FBCCFD53" box="[891,1066,618,643]" italics="true" pageId="12" pageNumber="13">D. pilosissimus</emphasis>
</taxonomicName>
and the simply broadly arcuate clypeal edge will distinguish it from
<taxonomicName id="4C6F4D17F06EFFFBFDE0FD62FD57FD77" box="[565,689,654,679]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="furtadoi">
<emphasis id="B91BEA86F06EFFFBFDE0FD62FD57FD77" box="[565,689,654,679]" italics="true" pageId="12" pageNumber="13">D. furtadoi</emphasis>
</taxonomicName>
. See diagnosis under
<taxonomicName id="4C6F4D17F06EFFFBFC7DFD62FBCAFD77" box="[936,1068,655,679]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="carinifer">
<emphasis id="B91BEA86F06EFFFBFC7DFD62FBCAFD77" box="[936,1068,655,679]" italics="true" pageId="12" pageNumber="13">D. carinifer</emphasis>
</taxonomicName>
for characters separating it from its sister species.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06EFFFAFF12FD3BFCC7FD54" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8BD03694F06EFFFAFF12FD3BFF13FE25" blockId="12.[151,1437,510,2011]" lastBlockId="13.[151,1437,151,895]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B91BEA86F06EFFFBFF12FD3BFEBFFD3F" bold="true" box="[199,345,726,751]" pageId="12" pageNumber="13">Description.</emphasis>
<collectingCountry id="F3787604F06EFFFBFE8AFD3BFE7FFD3F" box="[351,409,726,751]" name="Malta" pageId="12" pageNumber="13">Male</collectingCountry>
<typeStatus id="54D48836F06EFFFBFE4AFD3BFDE7FD3F" box="[415,513,726,751]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F06EFFFBFDC5FD3BFDB3FD3F" box="[528,597,726,751]" captionStart="FIGURES 1 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="12" pageNumber="13">Fig. 4</figureCitation>
).
<emphasis id="B91BEA86F06EFFFBFDBFFD3BFD41FD3F" bold="true" box="[618,679,726,751]" pageId="12" pageNumber="13">Body</emphasis>
. Body moderately large, length 13.0 mm, maximum width 7.0 mm; body subrectangular in dorsal view; dorsum slightly convex.
<emphasis id="B91BEA86F06EFFFBFC82FD17FC7EFCC3" bold="true" box="[855,920,762,787]" pageId="12" pageNumber="13">Color</emphasis>
. Dorsal surface dark brown to black, glossy, with green metallic sheen; head black along anterior edge of clypeus, metallic green on remaining surface; pronotum with green metallic sheen except disc and irregular area laterally; elytra with uniform green metallic sheen; ventrum with faint greenish metallic sheen; pygidium with green metallic sheen; legs with coppery and greenish metallic sheen on femora and tibiae.
<emphasis id="B91BEA86F06EFFFBFD73FC67FD03FC73" bold="true" box="[678,741,906,931]" pageId="12" pageNumber="13">Head</emphasis>
. Clypeus broadly arcuate, anterior portion slightly upturned; clypeal teeth acutely triangular; clypeal median emargination v-shaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae anteriorly and small setiferous tubercles posteriorly; clypeogenal suture well-defined, bluntly carinate internally; genal surface with small setiferous tubercles, lacking distinct transverse carina, simply convex; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge slightly trilobate in frontal view; eyes large in dorsal view, interocular ratio 3.8.
<emphasis id="B91BEA86F06EFFFBFCA3FB6BFC09FB4F" bold="true" box="[886,1007,1158,1183]" pageId="12" pageNumber="13">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.5; disc of pronotum minutely punctate basally with large confluent setiferous puncture medially changing into dense squamose and setiferous rugulae anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin unmodified lateral to eye; anterior portion with a slightly tri-sinuous carina notched medially; anterior angles surface finely granulate, similar to lateral margin along posterior edge of anterior margin; lateral fossae oval, bordered laterally by a sharp carina and anteriorly by a blunt tubercle; lateral portions strongly explanate; pronotal basal fossae ill-defined, slightly concave; posterior margin well-defined and crenulate, with several long setae.
<emphasis id="B91BEA86F06EFFFBFCE3FA6FFC64FA4B" bold="true" box="[822,898,1410,1435]" pageId="12" pageNumber="13">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.1; elytral base distinctly marginate; elytral striae 14 moderately wide basally and fine apically, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures fine, well-defined and setiferous, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae slightly convex, minutely punctate and with few larger setiferous punctures along striae, surface glossy.
<emphasis id="B91BEA86F06EFFFBFD8AF9DBFCC4F99F" bold="true" box="[607,802,1590,1615]" pageId="12" pageNumber="13">Thoracic sterna</emphasis>
. Proepisternal carina absent; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, v-shaped.
<emphasis id="B91BEA86F06EFFFBFC1BF9B7FBE0F9A3" bold="true" box="[974,1030,1626,1651]" pageId="12" pageNumber="13">Legs</emphasis>
. Profemur posterior surface flat and glabrous internally, posterointernal margin rather thin, uneven, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle bluntly lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with some ill-defined irregular punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur internal edge nearly straight and lateral edge arcuate, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on more than half the length. Metatibia rather slender, gradually widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with ill-defined microsculpture. Metatarsus 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
<emphasis id="B91BEA86F06FFFFAFC3FFEA1FB00FEB5" bold="true" box="[1002,1254,332,357]" pageId="13" pageNumber="14">Abdominal sternites</emphasis>
. Sternites 36 longitudinally flat; sternites 46 with 13 unaligned rows of setae laterally, narrowly glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium minutely punctate on disc.
<emphasis id="B91BEA86F06FFFFAFF03FE5AFE62FE00" bold="true" box="[214,388,439,464]" pageId="13" pageNumber="14">
<collectingCountry id="F3787604F06FFFFAFF03FE5AFEF4FE00" box="[214,274,439,464]" name="Malta" pageId="13" pageNumber="14">Male</collectingCountry>
genitalia
</emphasis>
(Figs. 4647). Parameres simply rounded apically in dorsal view; surface smooth, glossy apically.
</paragraph>
<paragraph id="8BD03694F06FFFFAFF12FE12FA98FDC7" blockId="13.[151,1437,151,895]" box="[199,1406,511,536]" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFF12FE12FE90FDC8" bold="true" box="[199,374,511,536]" pageId="13" pageNumber="14">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF06FFFFAFE50FDEDFE39FDC8" box="[389,479,511,536]" pageId="13" pageNumber="14" type="male">4 males</specimenCount>
,
<specimenCount id="9D69FD1DF06FFFFAFE3EFE12FDBFFDC8" box="[491,601,511,536]" pageId="13" pageNumber="14" type="female">6 females</specimenCount>
). Length: male 11.013.0 (12.3±0.9), female 12.513.5 (13.1±0.4) mm.
</paragraph>
<paragraph id="8BD03694F06FFFFAFF12FDC9FCC7FD54" blockId="13.[151,1437,151,895]" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFF12FDC9FE43FDED" bold="true" box="[199,421,548,573]" pageId="13" pageNumber="14">
Primary
<typeStatus id="54D48836F06FFFFAFEE1FDC9FE81FDED" box="[308,359,548,573]" pageId="13" pageNumber="14">type</typeStatus>
data
</emphasis>
.
<typeStatus id="54D48836F06FFFFAFE60FDC9FDFBFDED" box="[437,541,548,573]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
male (CPFA): [
<collectingCountry id="F3787604F06FFFFAFD07FDC9FCDBFDEC" box="[722,829,548,572]" name="Brazil" pageId="13" pageNumber="14">BRASIL</collectingCountry>
: CEARA/ Ubajara,
<quantity id="4C979B71F06FFFFAFBF5FDC9FB86FDEC" box="[1056,1120,548,572]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="13" pageNumber="14" unit="m" value="820.0">820m</quantity>
/
<geoCoordinate id="EE5B5053F06FFFFAFBA4FDC9FB2BFDED" box="[1137,1229,548,573]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-3.8333333">03°50S</geoCoordinate>
<geoCoordinate id="EE5B5053F06FFFFAFB00FDC9FADDFDED" box="[1237,1339,548,573]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="-40.933334">40°56W</geoCoordinate>
/
<date id="FFD11054F06FFFFAFA98FDC9FA71FDED" box="[1357,1431,548,573]" pageId="13" pageNumber="14" value="1995-01">I.1995</date>
, Miglioli, forêt/ primaire, piège lumineux]; [WORLD/ SCARAB./ DATABASE/ WSD00016742]; [
<typeStatus id="54D48836F06FFFFAFB28FDA5FA75FDB0" box="[1277,1427,584,608]" pageId="13" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C6F4D17F06FFFFAFF42FD81FEA2FD55" box="[151,324,620,645]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F06FFFFAFF42FD81FEA2FD55" box="[151,324,620,645]" italics="true" pageId="13" pageNumber="14">Dendropaemon</emphasis>
</taxonomicName>
/
<taxonomicName id="4C6F4D17F06FFFFAFE86FD81FE5AFD55" box="[339,444,620,645]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cribrosus" status="sp. nov.">
<emphasis id="B91BEA86F06FFFFAFE86FD81FE5AFD55" box="[339,444,620,645]" italics="true" pageId="13" pageNumber="14">cribrosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857FDF06FFFFAFE16FD81FE13FD54" box="[451,501,620,644]" pageId="13" pageNumber="14" rank="species">n.sp.</taxonomicNameLabel>
/ Génier &amp; Arnaud, 2014].
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06FFFFAFF12FD62FCB1FD1C" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BD03694F06FFFFAFF12FD62FCB1FD1C" blockId="13.[151,1437,151,895]" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFF12FD62FE4AFD78" bold="true" box="[199,428,655,680]" pageId="13" pageNumber="14">Material examined</emphasis>
.
<collectingCountry id="F3787604F06FFFFAFE69FD62FDC1FD78" box="[444,551,655,680]" name="Brazil" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFE69FD62FDC1FD78" bold="true" box="[444,551,655,680]" pageId="13" pageNumber="14">BRAZIL</emphasis>
</collectingCountry>
:
<collectionCode id="ED7EAE51F06FFFFAFDECFD62FD7DFD78" box="[569,667,655,680]" pageId="13" pageNumber="14">CEARÁ</collectionCode>
, Ubajara, elev.
<quantity id="4C979B71F06FFFFAFC85FD7DFC7CFD77" box="[848,922,656,680]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="13" pageNumber="14" unit="m" value="820.0">820 m</quantity>
(
<geoCoordinate id="EE5B5053F06FFFFAFC79FD62FC1EFD78" box="[940,1016,655,680]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-3.8333333">3°50'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F06FFFFAFBD2FD62FB8BFD78" box="[1031,1133,655,680]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="-40.933334">40°56'W</geoCoordinate>
),
<date id="FFD11054F06FFFFAFB51FD62FB2AFD78" box="[1156,1228,655,680]" pageId="13" pageNumber="14" value="1995-01">i.1995</date>
, coll. Miglioli—
<specimenCount id="9D69FD1DF06FFFFAFA5AFD62FF17FD1D" pageId="13" pageNumber="14" type="female">6 females</specimenCount>
,
<specimenCount id="9D69FD1DF06FFFFAFF29FD59FEB4FD1D" box="[252,338,692,717]" pageId="13" pageNumber="14" type="male">4 males</specimenCount>
(incl.
<typeStatus id="54D48836F06FFFFAFE4EFD59FDE6FD1D" box="[411,512,692,717]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, 8
<typeStatus id="54D48836F06FFFFAFDCAFD59FD6AFD1C" box="[543,652,692,716]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
) (
<collectionCode id="ED7EAE51F06FFFFAFD76FD59FD0AFD1C" box="[675,748,692,716]" pageId="13" pageNumber="14">CPFA</collectionCode>
,
<collectionCode id="ED7EAE51F06FFFFAFD22FD59FCAAFD1C" box="[759,844,692,716]" pageId="13" pageNumber="14">PMOC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06FFFFAFF12FD3AFC24FCC4" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BD03694F06FFFFAFF12FD3AFB7CFD20" blockId="13.[151,1437,151,895]" box="[199,1178,727,752]" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFF12FD3AFEA1FD20" bold="true" box="[199,327,727,752]" pageId="13" pageNumber="14">Etymology</emphasis>
.
<taxonomicName id="4C6F4D17F06FFFFAFE81FD3AFE25FD20" box="[340,451,727,752]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="117" phylum="Arthropoda" rank="species" species="cribrosus">
<emphasis id="B91BEA86F06FFFFAFE81FD3AFE25FD20" box="[340,451,727,752]" italics="true" pageId="13" pageNumber="14">Cribrosus</emphasis>
</taxonomicName>
, an adjective relating to the more heavily punctate pronotal disc.
</paragraph>
<paragraph id="8BD03694F06FFFFAFF12FD11FC24FCC4" blockId="13.[151,1437,151,895]" box="[199,962,764,789]" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFF12FD11FE9BFCC5" bold="true" box="[199,381,764,789]" pageId="13" pageNumber="14">Natural history</emphasis>
. All known specimens were collected at light traps.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06FFFFAFF12FCF2FEEDFCAF" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BD03694F06FFFFAFF12FCF2FEEDFCAF" blockId="13.[151,1437,151,895]" pageId="13" pageNumber="14">
<emphasis id="B91BEA86F06FFFFAFF12FCF2FED2FCE8" bold="true" box="[199,308,799,824]" pageId="13" pageNumber="14">Remarks</emphasis>
. Females differs in lacking the media tubercle of the pronotal carina and by their more widely medially glabrous sternites 14. Variation, little aside size and the extent of green metallic sheen on head and pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>