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<document ID-DOI="10.11646/zootaxa.5264.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836989" ID-ZooBank="9C61EF9D-1085-4EC6-B8A1-DCA65F768D76" IM.bibliography_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" approvalRequired="5" approvalRequired_for_textStreams="4" approvalRequired_for_treatments="1" checkinTime="1681730074228" checkinUser="plazi" docAuthor="Kostova, Rumyana &amp; Bekchiev, Rostislav" docDate="2023" docId="03C600089627A548CE85FEED0A95FE36" docLanguage="en" docName="zootaxa.5264.4.3.pdf" docOrigin="Zootaxa 5264 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus cardioderus subsp. balcanicus Jeannel 1927" docType="treatment" docVersion="3" lastPageNumber="511" masterDocId="FFFF78709622A54ECE12FF830B3FFFBF" masterDocTitle="Trechus (s. str.) “ subnotatus ” species group (Coleoptera: Carabidae) in Bulgaria" masterLastPageNumber="525" masterPageNumber="505" pageNumber="510" updateTime="1681831532550" updateUser="valdenar" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Trechus (s. str.) “ subnotatus ” species group (Coleoptera: Carabidae) in Bulgaria</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kostova, Rumyana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bekchiev, Rostislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>505</mods:start>
<mods:end>525</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.4.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.4.3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836989</mods:identifier>
<mods:identifier type="ZooBank">9C61EF9D-1085-4EC6-B8A1-DCA65F768D76</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36" httpUri="http://treatment.plazi.org/id/03C600089627A548CE85FEED0A95FE36" lastPageId="6" lastPageNumber="511" pageId="5" pageNumber="510">
<subSubSection box="[151,1354,366,393]" pageId="5" pageNumber="510" type="nomenclature">
<paragraph blockId="5.[151,1354,366,393]" box="[151,1354,366,393]" pageId="5" pageNumber="510">
<heading bold="true" box="[151,1354,366,393]" fontSize="11" level="1" pageId="5" pageNumber="510" reason="1">
<emphasis bold="true" box="[151,1354,366,393]" pageId="5" pageNumber="510">
3.
<taxonomicName ID-CoL="5829X" authority="Jeannel, 1927" authorityName="Jeannel" authorityYear="1927" box="[181,744,366,393]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="balcanicus">
<emphasis bold="true" box="[181,567,366,393]" italics="true" pageId="5" pageNumber="510">Trechus cardioderus balcanicus</emphasis>
<bibRefCitation author="Jeannel, R." box="[575,744,366,393]" pageId="5" pageNumber="510" refId="ref13152" refString="Jeannel, R. (1927) Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres. Deuxieme Livraison. Socieitei entomologique de France, Paris, 592 pp." type="book" year="1927">Jeannel, 1927</bibRefCitation>
</taxonomicName>
(stat. res.), revived from synonymy with
<taxonomicName authorityName="Csiki" authorityYear="1912" box="[1256,1354,367,393]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="irenis">
<emphasis bold="true" box="[1256,1354,367,393]" italics="true" pageId="5" pageNumber="510">T. irenis</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="510" type="description">
<paragraph blockId="5.[151,1437,439,969]" pageId="5" pageNumber="510">
The name
<taxonomicName authorityName="Jeannel" authorityYear="1927" box="[271,629,439,465]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="balcanicus">
<emphasis box="[271,629,439,465]" italics="true" pageId="5" pageNumber="510">Trechus cardioderus balcanicus</emphasis>
</taxonomicName>
is not supposed to be in use anymore since the established by
<bibRefCitation author="Moravec, P. &amp; Lompe, A." pageId="5" pageNumber="510" pagination="324 - 343" refId="ref13495" refString="Moravec, P. &amp; Lompe, A. (2003) Genus Trechus Clairville, 1806. In: Lobl, I. &amp; Smetana, A. (Eds.) Catalogue of Palaearctic Coleoptera. Vol. 1. Apollo Books, Stenstrup, pp. 22 - 23 + 324 - 343." type="book chapter" year="2003">Moravec &amp; Lompe (2003)</bibRefCitation>
homonymy of
<taxonomicName authorityName="Jeannel" authorityYear="1927" box="[514,804,475,501]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="balcanicus">
<emphasis box="[514,804,475,501]" italics="true" pageId="5" pageNumber="510">T. cardioderus balcanicus</emphasis>
</taxonomicName>
with
<taxonomicName authority="Frivaldszky, 1879" authorityName="Frivaldszky" authorityYear="1879" box="[869,1413,475,501]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="balcanicus" subGenus="Anophthalmus">
<emphasis box="[869,1205,475,501]" italics="true" pageId="5" pageNumber="510">T. (Anophthalmus) balcanicus</emphasis>
<bibRefCitation author="Frivaldszky, J." box="[1213,1413,475,501]" pageId="5" pageNumber="510" pagination="230 - 233" refId="ref12822" refString="Frivaldszky, J. (1879) Coleoptera nova ab Eduardo Merkl in M. Balkan inventa. Termeszetrajzi Fuzetek, 3, 230 - 233." type="journal article" year="1879">Frivaldszky, 1879</bibRefCitation>
</taxonomicName>
=
<taxonomicName authority="(Frivaldszky, 1879)" baseAuthorityName="Frivaldszky" baseAuthorityYear="1879" box="[151,790,511,537]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="balcanicus" subGenus="Paraduvalius">
<emphasis box="[151,252,511,537]" italics="true" pageId="5" pageNumber="510">Duvalius</emphasis>
(
<emphasis box="[269,420,511,537]" italics="true" pageId="5" pageNumber="510">Paraduvalius</emphasis>
)
<emphasis box="[437,558,511,537]" italics="true" pageId="5" pageNumber="510">balcanicus</emphasis>
(
<bibRefCitation author="Frivaldszky, J." box="[574,781,511,537]" pageId="5" pageNumber="510" pagination="230 - 233" refId="ref12822" refString="Frivaldszky, J. (1879) Coleoptera nova ab Eduardo Merkl in M. Balkan inventa. Termeszetrajzi Fuzetek, 3, 230 - 233." type="journal article" year="1879">Frivaldszky, 1879</bibRefCitation>
)
</taxonomicName>
. In accordance with the International Code of Zoological Nomenclature Art. 23.9.5. (
<bibRefCitation author="ICZN" box="[452,584,547,573]" pageId="5" pageNumber="510" refId="ref13773" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth edition. International Trust for Zoological Nomenclature. London. Available from: https: // www. iczn. org / the-code / the-code-online / (accessed 20 April 2022)" type="url" year="1999">ICZN 1999</bibRefCitation>
), as the taxa
<taxonomicName authorityName="Jeannel" authorityYear="1927" box="[720,1006,547,573]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="balcanicus">
<emphasis box="[720,1006,547,573]" italics="true" pageId="5" pageNumber="510">T. cardioderus balcanicus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Frivaldszky" baseAuthorityYear="1879" box="[1056,1210,547,573]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="balcanicus">
<emphasis box="[1056,1210,547,573]" italics="true" pageId="5" pageNumber="510">D. balcanicus</emphasis>
</taxonomicName>
were not considered congeneric after 1899, we must not automatically replace the junior homonym, and the case should be referred to the International Commission on Zoological Nomenclature. Until its decision, the prevailing usage of the name will remain.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="511" pageId="5" pageNumber="510" type="materials_examined">
<paragraph blockId="5.[151,1437,439,969]" pageId="5" pageNumber="510">
<emphasis bold="true" box="[199,455,690,717]" pageId="5" pageNumber="510">Type material: Type</emphasis>
:
<materialsCitation collectionCode="MNHN" collectorName="Jeannel" country="Bosnia and Herzegovina" location="Zepce" municipality="Apfelbeck" pageId="5" pageNumber="510" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[477,782,690,717]" pageId="5" pageNumber="510">
<collectingCountry box="[477,782,690,717]" name="Bosnia and Herzegovina" pageId="5" pageNumber="510">Bosnia and Herzegovina</collectingCountry>
</emphasis>
: 1³,
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCD4EFD37089CFD70" box="[860,931,692,719]" country="Bosnia and Herzegovina" municipality="Apfelbeck" name="Zepce" pageId="5" pageNumber="510">Žepče</location>
,
<collectingMunicipality box="[948,1068,691,717]" pageId="5" pageNumber="510">Apfelbeck</collectingMunicipality>
, 1903 (coll.
<collectorName box="[1228,1315,691,717]" pageId="5" pageNumber="510">Jeannel</collectorName>
,
<collectionCode box="[1334,1425,691,717]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="510">MNHN</collectionCode>
);
<typeStatus box="[151,275,727,753]" pageId="5" pageNumber="510">
<emphasis bold="true" box="[151,275,727,753]" pageId="5" pageNumber="510">Paratypes</emphasis>
</typeStatus>
</materialsCitation>
<emphasis bold="true" box="[275,280,727,753]" pageId="5" pageNumber="510">:</emphasis>
<materialsCitation box="[287,1256,726,755]" collectionCode="V, MNHN" collectorName="Apfelbeck &amp; Jeannel" country="Bosnia and Herzegovina" location="Srnetica" pageId="5" pageNumber="510" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[287,588,726,753]" pageId="5" pageNumber="510">
<collectingCountry box="[287,583,726,753]" name="Bosnia and Herzegovina" pageId="5" pageNumber="510">Bosnia and Herzegovina</collectingCountry>
:
</emphasis>
1³,
<emphasis box="[644,737,727,753]" italics="true" pageId="5" pageNumber="510">
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCC96FD5409DEFD4E" box="[644,737,727,753]" country="Bosnia and Herzegovina" name="Srnetica" pageId="5" pageNumber="510">Srnetica</location>
</emphasis>
planina,
<collectionCode box="[840,860,727,753]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="510" type="Museum">V</collectionCode>
.
<collectorName box="[870,987,727,753]" pageId="5" pageNumber="510">Apfelbeck</collectorName>
(coll.
<collectorName box="[1059,1146,727,753]" pageId="5" pageNumber="510">Jeannel</collectorName>
,
<collectionCode box="[1157,1248,727,753]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="510">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="MNHN" collectorName="Jeannel" country="Bosnia and Herzegovina" location="Kamenica" municipality="Apfelbeck" pageId="5" pageNumber="510" specimenCount="1" typeStatus="paratype">
1³,
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCB36FD540EA7FD4E" box="[1316,1432,727,753]" country="Bosnia and Herzegovina" municipality="Apfelbeck" name="Kamenica" pageId="5" pageNumber="510">Kamenica</location>
,
<collectingMunicipality box="[151,268,763,789]" pageId="5" pageNumber="510">Apfelbeck</collectingMunicipality>
(coll.
<collectorName box="[342,429,763,789]" pageId="5" pageNumber="510">Jeannel</collectorName>
,
<collectionCode box="[440,531,763,789]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="510">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[551,1339,763,791]" collectionCode="MNHN" country="Bosnia and Herzegovina" location="Kupres" pageId="5" pageNumber="510" specimenCount="1" typeStatus="paratype">
<collectorName box="[593,757,763,789]" pageId="5" pageNumber="510">H. Koprivnica</collectorName>
, près de
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCD4BFD780892FCAA" box="[857,941,763,789]" country="Bosnia and Herzegovina" name="Kupres" pageId="5" pageNumber="510">Kupres</location>
,
<collectorName box="[951,1068,763,789]" pageId="5" pageNumber="510">Apfelbeck</collectorName>
(coll.
<collectorName box="[1142,1229,763,789]" pageId="5" pageNumber="510">Jeannel</collectorName>
,
<collectionCode box="[1240,1331,763,789]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="510">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="MNHN" collectorName="Jeannel" country="Serbia" location="Stopica" municipality="Rojantsvo" pageId="5" pageNumber="510" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[1351,1429,762,788]" pageId="5" pageNumber="510">
<collectingCountry box="[1351,1429,762,788]" name="Serbia" pageId="5" pageNumber="510">Serbia</collectingCountry>
</emphasis>
: 1³,
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCEDAFC9C0A23FC86" box="[200,284,799,825]" country="Serbia" municipality="Rojantsvo" name="Stopica" pageId="5" pageNumber="510">Stopica</location>
pecina, à
<collectingMunicipality box="[398,512,799,825]" pageId="5" pageNumber="510">Rojantsvo</collectingMunicipality>
(coll.
<collectorName box="[584,671,799,825]" pageId="5" pageNumber="510">Jeannel</collectorName>
,
<collectionCode box="[682,773,799,825]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="510">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="5.[151,1437,439,969]" pageId="5" pageNumber="510">
<emphasis bold="true" box="[199,813,834,860]" pageId="5" pageNumber="510">
Other material examined:
<collectingCountry box="[525,634,834,860]" name="Bulgaria" pageId="5" pageNumber="510">Bulgaria</collectingCountry>
:
<collectingRegion box="[648,713,834,860]" pageId="5" pageNumber="510">Stara</collectingRegion>
planina
</emphasis>
<emphasis bold="true" box="[823,877,835,861]" pageId="5" pageNumber="510">Mts.</emphasis>
: 1³,
<collectingCounty box="[946,1011,835,861]" pageId="5" pageNumber="510">Kotel</collectingCounty>
, “Lednitsata”
<collectingMunicipality box="[1176,1236,835,861]" pageId="5" pageNumber="510">Cave</collectingMunicipality>
, “Zlosten”
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCB4BFCC00EA7FCE2" box="[1369,1432,835,861]" country="Bulgaria" county="Kotel" latitude="42.85172" longLatPrecision="1" longitude="26.55142" municipality="Cave" name="Place" pageId="5" pageNumber="510" stateProvince="Stara">Place</location>
,
<date box="[151,277,871,897]" pageId="5" pageNumber="510" value="2011-11-03">
<collectingDate box="[151,277,871,897]" pageId="5" pageNumber="510" value="2011-11-03">11.03.2011</collectingDate>
</date>
,
<collectorName box="[290,397,871,897]" pageId="5" pageNumber="510">B. Petrov</collectorName>
leg. (
<collectionCode box="[462,570,871,897]" pageId="5" pageNumber="510">NMNHS</collectionCode>
); 2³, BerkovitsaKom,
<date box="[845,974,871,897]" pageId="5" pageNumber="510" value="2016-07-08">
<collectingDate box="[845,974,871,897]" pageId="5" pageNumber="510" value="2016-07-08">07.08.2016</collectingDate>
</date>
,
<collectionCode box="[986,1008,871,897]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="510">R</collectionCode>
.
<collectorName box="[1020,1126,871,897]" pageId="5" pageNumber="510">Bekchiev</collectorName>
leg. (
<collectionCode box="[1191,1267,871,897]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="5" pageNumber="510">BFUS</collectionCode>
); 1 ³, Medven Vill., the river,
<geoCoordinate box="[332,457,906,933]" degrees="42.85172" direction="north" orientation="latitude" pageId="5" pageNumber="510" precision="1" value="42.85172">N42.85172</geoCoordinate>
<geoCoordinate box="[468,593,906,933]" degrees="26.55142" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="1" value="26.55142">E26.55142</geoCoordinate>
,
<date box="[608,737,907,933]" pageId="5" pageNumber="510" value="2019-08-08">
<collectingDate box="[608,737,907,933]" pageId="5" pageNumber="510" value="2019-08-08">08.08.2019</collectingDate>
</date>
,
<collectionCode box="[752,774,907,933]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="510">R</collectionCode>
.
<collectorName box="[788,881,907,933]" pageId="5" pageNumber="510">Kostova</collectorName>
leg. (
<collectionCode box="[952,1028,907,933]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="5" pageNumber="510">BFUS</collectionCode>
);
<materialsCitation collectingDate="2007-10-07" collectionCode="NMNHS" collectorName="B. Gueorguiev" country="Bulgaria" location="Place" pageId="5" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Ruy Mt.">
<emphasis bold="true" box="[1051,1164,907,933]" pageId="5" pageNumber="510">
<collectingRegion box="[1051,1159,907,933]" pageId="5" pageNumber="510">Ruy Mt.</collectingRegion>
:
</emphasis>
2³,
<specimenCount box="[1227,1262,907,933]" pageId="5" pageNumber="510" type="female">1♀</specimenCount>
, 2 km N from Lomnitsa Vill., “Transko zhdrelo”
<location LSID="urn:lsid:plazi:treatment:03C600089627A548CE85FEED0A95FE36:8EB0E7C59627A54BCC0EFC2C0964FC76" box="[540,603,943,969]" country="Bulgaria" name="Place" pageId="5" pageNumber="510" stateProvince="Ruy Mt.">Place</location>
, 9
<date box="[642,771,943,969]" pageId="5" pageNumber="510" value="2007-10-07">
<collectingDate box="[642,771,943,969]" pageId="5" pageNumber="510" value="2007-10-07">10.07.2007</collectingDate>
</date>
,
<collectorName box="[783,947,943,969]" pageId="5" pageNumber="510">B. Guéorguiev</collectorName>
leg. (
<collectionCode box="[1010,1115,943,969]" pageId="5" pageNumber="510">NMNHS</collectionCode>
)
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836998" ID-Zenodo-Dep="7836998" httpUri="https://zenodo.org/record/7836998/files/figure.png" pageId="5" pageNumber="510" startId="5.[151,250,1885,1909]" targetBox="[151,1436,993,1861]" targetPageId="5">
<paragraph blockId="5.[151,1436,1885,1981]" pageId="5" pageNumber="510">
<emphasis bold="true" box="[151,275,1885,1909]" pageId="5" pageNumber="510">FIGURE 3.</emphasis>
Variability in the median lobe of the aedeagus of
<taxonomicName authority="Jeann." authorityName="Jeannel" authorityYear="1927" box="[780,1175,1885,1909]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="balcanicus">
<emphasis box="[780,1104,1885,1909]" italics="true" pageId="5" pageNumber="510">Trechus cardioderus balcanicus</emphasis>
Jeann.
</taxonomicName>
in lateral and dorsal view (a,b—East Stara Planina Mts, Medven Vill.; c,d—West Stara Planina Mts, BerkovitsaKom; e,f—Ruy Mt., Lomnitsa Vill., “Transko zhdrelo” Place). Scale: 0.2 mm.
</paragraph>
</caption>
<paragraph blockId="6.[151,1436,150,393]" pageId="6" pageNumber="511">
<emphasis bold="true" box="[199,275,151,177]" pageId="6" pageNumber="511">Notes:</emphasis>
Relatively small: the body length of males is
<quantity box="[815,983,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.355" metricValueMax="4.76" metricValueMin="3.95" pageId="6" pageNumber="511" unit="mm" value="4.355" valueMax="4.76" valueMin="3.95">3.954.76 mm</quantity>
. The bodys dorsal surface is brown or dark brown and unicolourous. Reduced wings. Median lobe of the aedeagus: the apex in the lateral view is strongly elongated and curved downwards; the elongation is also visible dorsally, with the apical lamella separated from the main part of the median lobe by constriction, occasionally rather deep (
<figureCitation box="[951,1020,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1885,1909]" captionTargetBox="[151,1436,993,1861]" captionTargetId="figure-305@5.[151,1436,993,1861]" captionTargetPageId="5" captionText="FIGURE 3. Variability in the median lobe of the aedeagus of Trechus cardioderus balcanicus Jeann. in lateral and dorsal view (a,b—East Stara Planina Mts, Medven Vill.; c,d—West Stara Planina Mts, BerkovitsaKom; e,f—Ruy Mt., Lomnitsa Vill., “Transko zhdrelo” Place). Scale: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7836998" httpUri="https://zenodo.org/record/7836998/files/figure.png" pageId="6" pageNumber="511">Fig. 3</figureCitation>
). The studied material from
<collectingCountry box="[1340,1436,259,285]" name="Bulgaria" pageId="6" pageNumber="511">Bulgaria</collectingCountry>
fully corresponds to the description and male genitalia drawing of
<bibRefCitation author="Jeannel, R." box="[938,1169,294,321]" pageId="6" pageNumber="511" refId="ref13152" refString="Jeannel, R. (1927) Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres. Deuxieme Livraison. Socieitei entomologique de France, Paris, 592 pp." type="book" year="1927">Jeannel (1927: 455)</bibRefCitation>
and to the drawing of
<bibRefCitation author="Nonveiller, G. &amp; Pavicevic, D. &amp; Popovic, M." box="[151,461,331,357]" pageId="6" pageNumber="511" pagination="5 - 25" refId="ref13626" refString="Nonveiller, G., Pavicevic, D. &amp; Popovic, M. (1994) Les especes du genre Trechus actuellement connues de Serbie (Coleoptera, Carabidae). Bulletin de la Societe entomologique de France, 99 (1), 5 - 25. https: // doi. org / 10.3406 / bsef. 1994.17032" type="journal article" year="1994">
Nonveiller
<emphasis box="[278,335,331,357]" italics="true" pageId="6" pageNumber="511">et al.</emphasis>
(1994: 14)
</bibRefCitation>
, as well as to the material of the
<taxonomicName authorityName="Jeannel" authorityYear="1927" box="[831,1121,331,357]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="balcanicus">
<emphasis box="[831,1121,331,357]" italics="true" pageId="6" pageNumber="511">T. cardioderus balcanicus</emphasis>
</taxonomicName>
<typeStatus box="[1129,1177,331,357]" pageId="6" pageNumber="511">type</typeStatus>
series in the collection of
<collectionCode box="[182,204,367,393]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
. Jeannel at
<collectionCode box="[332,421,367,393]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>