159 lines
19 KiB
XML
159 lines
19 KiB
XML
<document id="C6772CDAADFE915D20A8CAA9AEFB0AE2" ID-DOI="10.11646/zootaxa.4097.1.1" ID-GBIF-Dataset="1bdc9f90-79e0-4265-b8b7-9057618b36db" ID-ISSN="1175-5326" ID-Zenodo-Dep="270988" ID-ZooBank="CD767562-3A19-4F4C-914D-80C50AD0D250" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459403589336" checkinUser="plazi" docAuthor="Baker, Alan N." docDate="2016" docId="03C57E0CA77FED27B0CDFC4DFCD9A2A5" docLanguage="en" docName="zootaxa.4097.1.1.pdf.imf" docOrigin="Zootaxa 4097 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiomusium acuferum Lyman 1875" docType="treatment" docVersion="5" lastPageNumber="4" masterDocId="FFFC0674A77CED24B05AFFDFFFB6A50A" masterDocTitle="An illustrated catalogue of type specimens of the bathyal brittlestar genera Ophiomusium Lyman and Ophiosphalma H. L. Clark (Echinodermata: Ophiuroidea)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="4" updateTime="1698186783575" updateUser="plazi">
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<mods:title id="1AEC981CA5E77096B2436782E19F89A9">An illustrated catalogue of type specimens of the bathyal brittlestar genera Ophiomusium Lyman and Ophiosphalma H. L. Clark (Echinodermata: Ophiuroidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="D9A71D2415261B3B6C6B99828686DEDC" type="personal">
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<mods:roleTerm id="A197906160A70CE969F87DE31AF607A0">Author</mods:roleTerm>
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<mods:namePart id="EFF6BF647195147D187778363D804E69">Baker, Alan N.</mods:namePart>
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<mods:typeOfResource id="82883E48E81E85A2B09195AFF1648AFE">text</mods:typeOfResource>
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<mods:title id="A04FDCD38731E5DD7556F3D78AE7090A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="66285459E577A3348575900EDCACD2E0">
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<mods:date id="EC1F7D64B120A22F8653F0129950F827">2016</mods:date>
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<mods:detail id="6EBBC3B69646C14814FCFB4FE181D971" type="volume">
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<mods:number id="08E1DE9EB6DE740DFFDF6094CA4B1DAE">4097</mods:number>
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</mods:detail>
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<mods:detail id="B8BF2FD0845BCE26C5ED58B2D08A91F7" type="issue">
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<mods:number id="1C0A972AB852EFD4F458621485CB993F">1</mods:number>
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<mods:classification id="7B447EE8C07D55A005C656F40809F168">journal article</mods:classification>
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<mods:identifier id="CF6FFE06C7A6BC6CCB1A2F7A48565C54" type="DOI">10.11646/zootaxa.4097.1.1</mods:identifier>
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<treatment id="03C57E0CA77FED27B0CDFC4DFCD9A2A5" ID-DOI="http://doi.org/10.5281/zenodo.6056417" ID-GBIF-Taxon="127639984" ID-Zenodo-Dep="6056417" LSID="urn:lsid:plazi:treatment:03C57E0CA77FED27B0CDFC4DFCD9A2A5" httpUri="http://treatment.plazi.org/id/03C57E0CA77FED27B0CDFC4DFCD9A2A5" lastPageNumber="4" pageId="3" pageNumber="4">
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<subSubSection id="C3769C91A77FED27B0CDFC4DFB29A106" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BD3CF1AA77FED27B0CDFC4DFEF9A6C6" blockId="3.[151,606,914,973]" pageId="3" pageNumber="4">
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<taxonomicName id="4C6CB499A77FED27B0CDFC4DFDE1A6A1" authority="Lyman, 1875" authorityName="Lyman" authorityYear="1875" box="[151,599,914,939]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="acuferum">
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<emphasis id="B9181308A77FED27B0CDFC4DFE01A6A1" bold="true" box="[151,439,914,939]" italics="true" pageId="3" pageNumber="4">Ophiomusium acuferum</emphasis>
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<bibRefCitation id="EFFDB2EBA77FED27B19FFC4CFDE1A6A1" author="Lyman" box="[453,599,914,939]" pageId="3" pageNumber="39" refString="Lyman, T. (1875) Zoological Results of the Hassler Expedition. 2. Ophiuridae and Astrophytidae. Illustrated Catalogue of the Museum of Comparative Zoology, Harvard University, 8, 1 - 34, 5 pls." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
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</taxonomicName>
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.
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<figureCitation id="1357D39FA77FED27B0CDFC6BFF49A6C7" box="[151,255,948,973]" captionStart="FIGURE 1" captionStartId="4.[151,250,1916,1938]" captionText="FIGURE 1. Ophiomusium acuferum MCZ 427: a, b, dorsal and ventral disc sectors; c, basal arm segments; d, e Ophiomusium sculptum USNM 21289: dorsal sector." pageId="3" pageNumber="4">Figures 1</figureCitation>
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a, b, c,
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</paragraph>
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<paragraph id="8BD3CF1AA77FED27B0CDFC29FB29A106" blockId="3.[151,1183,1014,1036]" box="[151,1183,1014,1036]" pageId="3" pageNumber="4">
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<taxonomicName id="4C6CB499A77FED27B0CDFC29FDAAA106" authority="Lyman, 1875" authorityName="Lyman" authorityYear="1875" box="[151,540,1014,1036]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="acuferum">
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<emphasis id="B9181308A77FED27B0CDFC29FE38A106" box="[151,398,1014,1036]" italics="true" pageId="3" pageNumber="4">Ophiomusium acuferum</emphasis>
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<bibRefCitation id="EFFDB2EBA77FED27B1CEFC28FDAAA106" author="Lyman" box="[404,540,1014,1036]" pageId="3" pageNumber="39" refString="Lyman, T. (1875) Zoological Results of the Hassler Expedition. 2. Ophiuridae and Astrophytidae. Illustrated Catalogue of the Museum of Comparative Zoology, Harvard University, 8, 1 - 34, 5 pls." type="journal article" year="1875">Lyman, 1875</bibRefCitation>
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</taxonomicName>
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;
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<bibRefCitation id="EFFDB2EBA77FED27B27CFC28FC96A106" author="Borges" box="[550,800,1014,1036]" pageId="3" pageNumber="38" refString="Borges, M. & Campos, L. de Siqueira (2011) Ophiomusium acuferum (Ophiolepididae and Ophiomisidium pulchellum (Ophiuridae) (Ophiuroidea: Echinodermata) redescription based on the Brazilian specimens. Zoologia, 28, 219 - 225. http: // dx. doi. org / 10.1590 / S 1984 - 46702011000200010" type="journal article" year="2011">Borges & Campos, 2011</bibRefCitation>
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.
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<taxonomicName id="4C6CB499A77FED27B377FC29FB29A106" authority="Verrill, 1899" authorityName="Verrill" authorityYear="1899" box="[813,1183,1014,1036]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="sculptum">
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<emphasis id="B9181308A77FED27B377FC29FBADA106" box="[813,1051,1014,1036]" italics="true" pageId="3" pageNumber="4">Ophiomusium sculptum</emphasis>
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<bibRefCitation id="EFFDB2EBA77FED27B47AFC29FB29A106" author="Verrill" box="[1056,1183,1014,1036]" pageId="3" pageNumber="40" refString="Verrill, A. E. (1899) Report on the Ophiuroidea collected by the Bahama Expedition in 1893. Bulletin of the Laboratories of Natural History of the State of Iowa, 5, 1 - 18, pls 1 - 8." type="journal article" year="1899">Verrill, 1899</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3769C91A77FED27B0CDFBE9FA31A009" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="8BD3CF1AA77FED27B0CDFBE9FD10A145" blockId="3.[151,1437,1078,1968]" box="[151,678,1078,1103]" pageId="3" pageNumber="4">
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<emphasis id="B9181308A77FED27B0CDFBE9FEB5A145" bold="true" box="[151,259,1078,1103]" pageId="3" pageNumber="4">
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<typeStatus id="54D771B8A77FED27B0CDFBE9FEB5A145" box="[151,259,1078,1103]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectionCode id="ED7D57DFA77FED27B153FBE8FEF3A145" LSID="urn:lsid:biocol.org:col:33791" box="[265,325,1079,1103]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="4">MCZ</collectionCode>
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427, off
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<collectingCountry id="F37B8F8AA77FED27B1EBFBE9FDABA145" box="[433,541,1078,1103]" name="Barbados" pageId="3" pageNumber="4">Barbados</collectingCountry>
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,
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<quantity id="4C9462FFA77FED27B27DFBE8FD14A145" box="[551,674,1079,1103]" metricMagnitude="2" metricUnit="m" metricValue="1.98" metricValueMax="2.19" metricValueMin="1.77" pageId="3" pageNumber="4" unit="m" value="198.0" valueMax="219.0" valueMin="177.0">
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177–
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<specimenCount id="9D6A0493A77FED27B205FBE8FD14A145" box="[607,674,1079,1103]" pageId="3" pageNumber="4" type="generic">219m</specimenCount>
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</quantity>
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.
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</paragraph>
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<paragraph id="8BD3CF1AA77FED27B09DFB85FA31A009" blockId="3.[151,1437,1078,1968]" pageId="3" pageNumber="4">
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Other material examined:
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<specimenCount id="9D6A0493A77FED27B1B5FB84FDCEA179" box="[495,632,1114,1139]" pageId="3" pageNumber="4" type="generic">3 specimens</specimenCount>
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<collectionCode id="ED7D57DFA77FED27B2D9FB84FD63A178" box="[643,725,1115,1138]" pageId="3" pageNumber="4">BMNH</collectionCode>
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<date id="FFD2E9DAA77FED27B2BBFB85FCEBA179" box="[737,861,1114,1139]" pageId="3" pageNumber="4" value="1992-02-22" valueMax="1992-02-16" valueMin="1992-02-22">92-2-22-16</date>
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-18,
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<collectingCountry id="F37B8F8AA77FED27B3CAFB85FC4FA179" box="[912,1017,1114,1139]" name="Barbados" pageId="3" pageNumber="4">Barbados</collectingCountry>
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,
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<quantity id="4C9462FFA77FED27B452FB84FBE5A178" box="[1032,1107,1115,1139]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="3" pageNumber="4" unit="m" value="133.0">133 m</quantity>
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;
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<specimenCount id="9D6A0493A77FED27B439FB84FB5BA179" box="[1123,1261,1114,1139]" pageId="3" pageNumber="4" type="generic">4 specimens</specimenCount>
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<collectionCode id="ED7D57DFA77FED27B4A2FB84FAF1A179" LSID="urn:lsid:biocol.org:col:34871" box="[1272,1351,1115,1139]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
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21289, Mayaguez Harbour,
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<collectingCountry id="F37B8F8AA77FED27B127FB5FFDB2A19D" box="[381,516,1150,1175]" name="Puerto Rico" pageId="3" pageNumber="4">Puerto Rico</collectingCountry>
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, 97–120 fms (as
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<taxonomicName id="4C6CB499A77FED27B296FBA0FCE7A19D" box="[716,849,1150,1175]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="sculptum">
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<emphasis id="B9181308A77FED27B296FBA0FCE7A19D" box="[716,849,1150,1175]" italics="true" pageId="3" pageNumber="4">
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<collectingCountry id="F37B8F8AA77FED27B296FBA0FD54A19C" box="[716,738,1151,1174]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
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. sculptum
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</emphasis>
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</taxonomicName>
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);
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<specimenCount id="9D6A0493A77FED27B337FBA0FC4EA19D" box="[877,1016,1150,1175]" pageId="3" pageNumber="4" type="generic">9 specimens</specimenCount>
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,
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<collectionCode id="ED7D57DFA77FED27B45DFBA0FBE0A19D" LSID="urn:lsid:biocol.org:col:34871" box="[1031,1110,1151,1175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
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E513, E514, E4375, E4539, E4540, E4538, University of Iowa Barbados-Antigua Expedition Stations 4, 7, 15, 28, 44 49. The Yale Peabody Museum in Boston was searched for the
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<typeStatus id="54D771B8A77FED27B217FB17FD30A1EA" box="[589,646,1224,1248]" pageId="3" pageNumber="4">types</typeStatus>
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of
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<taxonomicName id="4C6CB499A77FED27B2F3FB18FC9AA1D5" box="[681,812,1222,1247]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="sculptum">
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<emphasis id="B9181308A77FED27B2F3FB18FC9AA1D5" box="[681,812,1222,1247]" italics="true" pageId="3" pageNumber="4">
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<collectingCountry id="F37B8F8AA77FED27B2F3FB18FD76A1D4" box="[681,704,1223,1246]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
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. sculptum
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</emphasis>
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</taxonomicName>
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, but none were located. Dr Marc Eleaume, Paris Museum has recently informed me that a
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<typeStatus id="54D771B8A77FED27B1A8FB34FDF0A009" box="[498,582,1259,1283]" pageId="3" pageNumber="4" type="syntype">syntype</typeStatus>
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of this species in in their collection:
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<collectionCode id="ED7D57DFA77FED27B389FB34FB91A008" LSID="urn:lsid:biocol.org:col:34988" box="[979,1063,1259,1282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="4">MNHN</collectionCode>
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10284, ex-
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<emphasis id="B9181308A77FED27B4FBFB35FAB1A009" box="[1185,1287,1258,1283]" italics="true" pageId="3" pageNumber="4">Albatross</emphasis>
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expedition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3769C91A77FED27B09DFAD1FCD9A2A5" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="8BD3CF1AA77FED27B09DFAD1FC58A0D1" blockId="3.[151,1437,1078,1968]" pageId="3" pageNumber="4">
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<emphasis id="B9181308A77FED27B09DFAD1FEC9A02D" bold="true" box="[199,383,1294,1319]" pageId="3" pageNumber="4">Main Features:</emphasis>
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In the
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<typeStatus id="54D771B8A77FED27B186FAD1FDF6A02D" box="[476,576,1294,1319]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
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, the periphery of marginal disc plates is raised and knobbly. All other disc plates have one or more tubercles, those adjacent to the marginal plates with a group of 3–4 tubercles concentrated at their distal corners. Those inside this ring (primaries) have one or two central tubercles. First dorsal arm plate raised and knobbly and separates the radial shields. Dorsal sides of the lateral arm plates are also knobbly. Small dorsal arm plates lie in a median longitudinal depression. Interradial marginal plates large, V-shaped, protuberant, and knobbly. Otherwise, the surface of all plates is smooth (micro-reticulate).
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</paragraph>
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<paragraph id="8BD3CF1AA77FED27B09DFA39FD7CA34D" blockId="3.[151,1437,1078,1968]" pageId="3" pageNumber="4">Ventral surface smooth, 2 pairs of exposed tentacle pores, 1 circular scale. Closely adjoining oral papillae. Sinuous genital slits run to the distal end of the first lateral arm plate. Oral shield, 2 genital plates, and a large interradial plate only in the ventral interradial area.</paragraph>
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<paragraph id="8BD3CF1AA77FED27B09DF98DFB46A3B8" blockId="3.[151,1437,1078,1968]" pageId="3" pageNumber="4">Ventral arm plates throughout, but very small distally. 3 small arm spines. On basal arm segments a longer spine is present on the dorsal side of each lateral arm plate, each side of the median groove. Lower down on the arm near the ventral margin, are two small spines the lowermost of which has a bifurcate tip and hollow core.</paragraph>
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<paragraph id="8BD3CF1AA77FED27B09DF961FCD9A2A5" blockId="3.[151,1437,1078,1968]" pageId="3" pageNumber="4">
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This species is conspecific with
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<taxonomicName id="4C6CB499A77FED27B274F960FCF4A3DD" authority="Verrill 1899" authorityName="Verrill" authorityYear="1899" box="[558,834,1726,1751]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="sculptum">
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||
<emphasis id="B9181308A77FED27B274F960FD02A3DD" box="[558,692,1726,1751]" italics="true" pageId="3" pageNumber="4">
|
||
<collectingCountry id="F37B8F8AA77FED27B274F960FDF3A3DC" box="[558,581,1727,1750]" name="Iceland" pageId="3" pageNumber="4">O</collectingCountry>
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||
. sculptum
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||
</emphasis>
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||
<bibRefCitation id="EFFDB2EBA77FED27B2E1F961FCF4A3DD" author="Verrill" box="[699,834,1726,1751]" pageId="3" pageNumber="40" refString="Verrill, A. E. (1899) Report on the Ophiuroidea collected by the Bahama Expedition in 1893. Bulletin of the Laboratories of Natural History of the State of Iowa, 5, 1 - 18, pls 1 - 8." type="journal article" year="1899">Verrill 1899</bibRefCitation>
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||
</taxonomicName>
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as identified in the
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<collectionCode id="ED7D57DFA77FED27B445F960FBD9A3DD" LSID="urn:lsid:biocol.org:col:34871" box="[1055,1135,1727,1751]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
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collections, including one specimen identified by Verrill himself, and others identified by H. L. Clark. These specimens compare well with Verrill’s original description and figures, except for the first ventral arms plates shown in his
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<figureCitation id="1357D39FA77FED27B495F8D9FA87A215" box="[1231,1329,1798,1823]" captionStart="FIGURE 2" captionStartId="4.[151,250,1977,1999]" captionTargetBox="[213,1366,205,1886]" captionTargetId="figure@4.[201,1383,195,1896]" captionTargetPageId="4" captionText="FIGURE 2. Ophiomusium alecto USNM E 0005197: a, b, dorsal and ventral disc sectors. FIGURE 3. Ophiomusium altum RMNH V. Ech. O 2045: a, b dorsal and ventral disc sectors." httpUri="https://zenodo.org/record/270989/files/figure.png" pageId="3" pageNumber="4">Figure 2</figureCitation>
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, Plate II, which are unlike any
|
||
<taxonomicName id="4C6CB499A77FED27B1CDF8F5FD85A249" box="[407,563,1834,1859]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9181308A77FED27B1CDF8F5FD85A249" box="[407,563,1834,1859]" italics="true" pageId="3" pageNumber="4">Ophiomusium</emphasis>
|
||
</taxonomicName>
|
||
in that they are extremely small (25% of arm width) and the exposed tentacle pores are located in the middle of the ventral faces of the lateral arm plates. Also, Verrill did not mention in his description of the
|
||
<typeStatus id="54D771B8A77FED27B13EF8ACFE22A281" box="[356,404,1907,1931]" pageId="3" pageNumber="4">type</typeStatus>
|
||
the presence of dorsal arm spines, which are present in the specimens he identified in the
|
||
<collectionCode id="ED7D57DFA77FED27B0CDF848FF5DA2A5" LSID="urn:lsid:biocol.org:col:34871" box="[151,235,1943,1967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
|
||
. Verrill’s
|
||
<typeStatus id="54D771B8A77FED27B10EF847FE32A2BA" box="[340,388,1944,1968]" pageId="3" pageNumber="4">type</typeStatus>
|
||
specimen could not be located in the
|
||
<collectionCode id="ED7D57DFA77FED27B376F847FCDAA2A5" LSID="urn:lsid:biocol.org:col:34880" box="[812,876,1944,1967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="3" pageNumber="4">YPM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |