168 lines
18 KiB
XML
168 lines
18 KiB
XML
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<mods:title id="F2BE13A226C097AF671B2DBFF2EF0D36">The first Triassic beaded lacewing (Neuroptera: Berothidae) from Central Asia with redescription of Mesoberotha superbа (Riek, 1955)</mods:title>
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<mods:namePart id="68285DF8BF336822982B421CA41F0A95">Khramov, Alexander V.</mods:namePart>
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<mods:date id="98D40C958CB5FDD0221DE21346B41148">2023</mods:date>
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<taxonomicName id="4C6CB692FF95883A5A82FA8BFD83871C" ID-CoL="6RC7X" authority="(Riek, 1955)" baseAuthorityName="Riek" baseAuthorityYear="1955" box="[151,571,1369,1396]" class="Insecta" family="Mesoberothidae" genus="Mesoberotha" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="288" phylum="Arthropoda" rank="species" species="superba">
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<emphasis id="B9181103FF95883A5A82FA8BFD83871C" bold="true" box="[151,571,1369,1396]" pageId="1" pageNumber="288">
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<emphasis id="B9181103FF95883A5A82FA8BFE25871B" bold="true" box="[151,413,1369,1396]" italics="true" pageId="1" pageNumber="288">Mesoberotha superba</emphasis>
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(
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<bibRefCitation id="EFFDB0E0FF95883A5BB9FA8BFD8B871C" author="Riek, E. F." box="[428,563,1369,1395]" pageId="1" pageNumber="288" pagination="654 - 690" refId="ref4687" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3, 654 - 690. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek, 1955</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3769E9AFF95883A5A82FAACFF5387F7" box="[151,235,1406,1432]" pageId="1" pageNumber="288" type="description">
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<paragraph id="8BD3CD11FF95883A5A82FAACFF5387F7" blockId="1.[151,571,1369,1432]" box="[151,235,1406,1432]" pageId="1" pageNumber="288">
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(
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<figureCitation id="1357D194FF95883A5A8AFAACFF5887F7" box="[159,224,1406,1432]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[245,1342,179,1913]" captionTargetId="figure-24@2.[245,1342,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Forewing of Mesoberotha superba (Riek, 1955) (Berothidae) from the Upper Triassic of Australia; holotype C1404, C2098, part and counterpart, aligned in the same direction, photographs and drawing. Photos by Jakub Prokop. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8249421" httpUri="https://zenodo.org/record/8249421/files/figure.png" pageId="1" pageNumber="288">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3769E9AFF95883A5A82FA10FEB98796" pageId="1" pageNumber="288" type="reference_group">
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<paragraph id="8BD3CD11FF95883A5A82FA10FEB98796" blockId="1.[151,1437,1474,1529]" pageId="1" pageNumber="288">
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<treatmentCitationGroup id="AB7CEA3FFF95883A5A82FA10FEB98796" pageId="1" pageNumber="288">
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<treatmentCitation id="0ACDEB00FF95883A5A82FA10FDFD87B5" author="Riek, E. F." box="[151,581,1474,1498]" page="676" pageId="1" pageNumber="288" year="1955">
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<taxonomicName id="4C6CB692FF95883A5A82FA10FDFD87B5" authority="Riek" authorityName="Riek" authorityPageNumber="675 - 676" authorityYear="1955" box="[151,581,1474,1498]" class="Insecta" family="Berothidae" genus="Proberotha" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="288" phylum="Arthropoda" rank="species" species="superba">
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<emphasis id="B9181103FF95883A5A82FA10FEDC87B5" box="[151,356,1474,1498]" italics="true" pageId="1" pageNumber="288">Proberotha superba</emphasis>
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<bibRefCitation id="EFFDB0E0FF95883A5B7EFA10FDFD87B5" author="Riek, E. F." box="[363,581,1474,1498]" pageId="1" pageNumber="288" pagination="654 - 690" refId="ref4687" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3, 654 - 690. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek, 1955: 675–676</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, fig. 18. Refs.
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<bibRefCitation id="EFFDB0E0FF95883A58CAFA10FBF887B5" author="Ponomarenko, A. G. & Shcherbakov, D. E." box="[735,1088,1474,1498]" pageId="1" pageNumber="288" pagination="203" refId="ref4600" refString="Ponomarenko, A. G. & Shcherbakov, D. E. (2004) New Lacewings (Neuroptera) from the Terminal Permian and Basal Triassic of Siberia. Paleontological Journal, 38, S 197 - S 203." type="journal article" year="2004">Ponomarenko, Shcherbakov, 2004</bibRefCitation>
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;
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<bibRefCitation id="EFFDB0E0FF95883A5E5EFA10FA9487B5" author="Makarkin, V. N. & Yang, Q. & Ren, D." box="[1099,1324,1474,1498]" pageId="1" pageNumber="288" refId="ref4453" refString="Makarkin, V. N., Yang, Q. & Ren, D. (2011) Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae. ZooKeys, 130, 199 - 125. https: // doi. org / 10.3897 / zookeys. 130.1418" type="book" year="2011">
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Makarkin
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<emphasis id="B9181103FF95883A5EA0FA11FB5287B5" box="[1205,1258,1474,1498]" italics="true" pageId="1" pageNumber="288">et al.</emphasis>
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, 2011
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</bibRefCitation>
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||
;
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<bibRefCitation id="EFFDB0E0FF95883A5F2DFA10FF458796" author="Khramov, A. V." pageId="1" pageNumber="288" pagination="26 - 35" refId="ref4129" refString="Khramov, A. V. (2015) Jurassic Beaded Lacewings (Insecta: Neuroptera: Berothidae) from Kazakhstan and Mongolia. Paleontological Journal, 49, 26 - 35. https: // doi. org / 10.1134 / S 0031030115010062" type="journal article" year="2015">Khramov, 2015</bibRefCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3769E9AFF95883A5A82F9F5FC2C850E" pageId="1" pageNumber="288" type="description">
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<paragraph id="8BD3CD11FF95883A5A82F9F5FC2C850E" blockId="1.[151,1437,1575,1998]" pageId="1" pageNumber="288">
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<emphasis id="B9181103FF95883A5A82F9F5FE91842E" bold="true" box="[151,297,1575,1601]" pageId="1" pageNumber="288">Description.</emphasis>
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Forewing
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long,
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<quantity id="4C9460F4FF95883A5833F9F5FDC4842D" box="[550,636,1575,1602]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="288" unit="mm" value="3.5">3.5 mm</quantity>
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||
wide, trichosors visible; recurrent humeral vein absent; costal area with a hump in the proximal half, narrowed toward wing base; subcostal veinlets strongly bended forward, many of them deeply forked; at least one crossvein discernible between Sc and R1; Sc+R1 rather short, bended downward; Rs with 8 branches, 3rd and 4th of them approaching each other neat the base, being connected by short crossvein; outer gradate series of crossveins consisting of six crossveins, middle gradate series of crossveins consists of four crossveins, MA with small terminal fork; MP bifurcated slightly distal to origin of MA, but proximal to origin of Rs1; MP1 and MP2 deeply forked terminally, CuA pectinate, with four branches, 1st of them bifurcated near the base; CuP bifurcated proximal to origin of MA, anterior branch of CuP deeply bifurcated terminally; A1 and A2 with four short terminal branches each, A3 with small terminal fork.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3769E9AFF95883A5AD2F8B9FE1C85A1" pageId="1" pageNumber="288" type="discussion">
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<paragraph id="8BD3CD11FF95883A5AD2F8B9FE1C85A1" blockId="1.[151,1437,1575,1998]" pageId="1" pageNumber="288">
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<emphasis id="B9181103FF95883A5AD2F8B9FEF085EA" bold="true" box="[199,328,1899,1925]" pageId="1" pageNumber="288">Comments</emphasis>
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||
.
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<materialsCitation id="3B04C74CFF95883A5B40F8B9FE1885A1" collectionCode="R" pageId="1" pageNumber="288" specimenCode="R1" specimenCount="1" typeStatus="holotype">
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While distal portions of Sc and
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<specimenCode id="DBCA656AFF95883A58A6F8B9FD6C85EA" box="[691,724,1899,1925]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="288">R1</specimenCode>
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are only partially preserved in the
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<typeStatus id="54D773B3FF95883A5E43F8B9FB0385EA" box="[1110,1211,1899,1925]" pageId="1" pageNumber="288">holotype</typeStatus>
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, these veins seem to be fused, as Riek suggested in the original description (1955). Apart from some missing crossveins, Riekʼs drawing turned out to be correct
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</materialsCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF139D99FF9688395A82F845FEB88598" ID-DOI="http://doi.org/10.5281/zenodo.8249421" ID-Zenodo-Dep="8249421" httpUri="https://zenodo.org/record/8249421/files/figure.png" pageId="2" pageNumber="289" startId="2.[151,250,1943,1967]" targetBox="[245,1342,179,1913]" targetPageId="2" targetType="figure">
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<paragraph id="8BD3CD11FF9688395A82F845FEB88598" blockId="2.[151,1437,1943,2040]" pageId="2" pageNumber="289">
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<emphasis id="B9181103FF9688395A82F845FEAF85C0" bold="true" box="[151,279,1943,1967]" pageId="2" pageNumber="289">FIGURE 1.</emphasis>
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Forewing of
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<taxonomicName id="4C6CB692FF9688395BB9F84AFCA385C0" authority="(Riek, 1955)" baseAuthorityName="Riek" baseAuthorityYear="1955" box="[428,795,1943,1968]" class="Insecta" family="Mesoberothidae" genus="Mesoberotha" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="289" phylum="Arthropoda" rank="species" species="superba">
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<emphasis id="B9181103FF9688395BB9F84AFD3585DF" box="[428,653,1944,1968]" italics="true" pageId="2" pageNumber="289">Mesoberotha superba</emphasis>
|
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(
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<bibRefCitation id="EFFDB0E0FF968839588BF84AFCAC85C0" author="Riek, E. F." box="[670,788,1943,1968]" pageId="2" pageNumber="289" pagination="654 - 690" refId="ref4687" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3, 654 - 690. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek, 1955</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
(
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||
<taxonomicName id="4C6CB692FF9688395939F84AFC2685DF" authorityName="Handlirsch" authorityYear="1906" box="[812,926,1944,1968]" class="Insecta" family="Berothidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="289" phylum="Arthropoda" rank="family">Berothidae</taxonomicName>
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||
) from the Upper Triassic of Australia; holotype C1404, C2098, part and counterpart, aligned in the same direction, photographs and drawing. Photos by Jakub Prokop. Scale bar 1 mm.
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||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3769E9AFF9788385AD2FF45FF408372" pageId="3" pageNumber="290" type="materials_examined">
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<paragraph id="8BD3CD11FF9788385AD2FF45FD0182BA" blockId="3.[151,1436,150,285]" pageId="3" pageNumber="290">
|
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<emphasis id="B9181103FF9788385AD2FF45FEC082DF" bold="true" box="[199,376,150,177]" pageId="3" pageNumber="290">Type material.</emphasis>
|
||
<materialsCitation id="3B04C74CFF9788385B97FF45FD0D82BA" country="Australia" location="Museum" municipality="South Brisbane" pageId="3" pageNumber="290" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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||
Almost complete forewing,
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||
<typeStatus id="54D773B3FF97883858D3FF45FC9082DE" box="[710,808,151,177]" pageId="3" pageNumber="290">holotype</typeStatus>
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||
№ C1404 (part), C2098 (counterpart), now housed at
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<collectingRegion id="49A803F3FF9788385A82FF69FEA382BA" box="[151,283,187,213]" country="Australia" name="Queensland" pageId="3" pageNumber="290">Queensland</collectingRegion>
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<location id="8EB39BCAFF9788385B37FF69FE3F82BA" LSID="urn:lsid:plazi:treatment:03C57C07FF9588385A82FA8BFF408372:8EB39BCAFF9788385B37FF69FE3F82BA" box="[290,391,187,213]" country="Australia" municipality="South Brisbane" name="Museum" pageId="3" pageNumber="290" stateProvince="Queensland">Museum</location>
|
||
,
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||
<collectingMunicipality id="6BB7576BFF9788385B87FF69FDF982BA" box="[402,577,187,213]" pageId="3" pageNumber="290">South Brisbane</collectingMunicipality>
|
||
,
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||
<collectingCountry id="F37B8D81FF978838585EFF69FD0D82BA" box="[587,693,187,213]" name="Australia" pageId="3" pageNumber="290">Australia</collectingCountry>
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||
</materialsCitation>
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||
.
|
||
</paragraph>
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||
<paragraph id="8BD3CD11FF9788385AD2FF0DFF408372" blockId="3.[151,1436,150,285]" pageId="3" pageNumber="290">
|
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<materialsCitation id="3B04C74CFF9788385AD2FF0DFF4B8372" country="Australia" county="Carnian" location="Mount Crosby" municipality="Mount Crosby Formation" pageId="3" pageNumber="290" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<emphasis id="B9181103FF9788385AD2FF0DFE4D8297" bold="true" box="[199,501,222,249]" pageId="3" pageNumber="290">
|
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<typeStatus id="54D773B3FF9788385AD2FF0DFEBA8296" box="[199,258,223,249]" pageId="3" pageNumber="290">Type</typeStatus>
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||
locality and horizon
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||
</emphasis>
|
||
.
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||
<location id="8EB39BCAFF9788385817FF0DFD1D8296" LSID="urn:lsid:plazi:treatment:03C57C07FF9588385A82FA8BFF408372:8EB39BCAFF9788385817FF0DFD1D8296" box="[514,677,223,249]" country="Australia" county="Carnian" municipality="Mount Crosby Formation" name="Mount Crosby" pageId="3" pageNumber="290" stateProvince="Queensland">Mount Crosby</location>
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||
, north of
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||
<location id="8EB39BCAFF978838591BFF0DFCD28296" LSID="urn:lsid:plazi:treatment:03C57C07FF9588385A82FA8BFF408372:8EB39BCAFF978838591BFF0DFCD28296" box="[782,874,223,249]" country="Australia" county="Carnian" municipality="Mount Crosby Formation" name="Ipswich" pageId="3" pageNumber="290" stateProvince="Queensland">Ipswich</location>
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||
,
|
||
<collectingRegion id="49A803F3FF9788385961FF0DFC438296" box="[884,1019,223,249]" country="Australia" name="Queensland" pageId="3" pageNumber="290">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37B8D81FF9788385E11FF0DFBD68296" box="[1028,1134,223,249]" name="Australia" pageId="3" pageNumber="290">Australia</collectingCountry>
|
||
;
|
||
<collectingMunicipality id="6BB7576BFF9788385E6DFF0DFA208296" box="[1144,1432,223,249]" pageId="3" pageNumber="290">Mount Crosby Formation</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62B2B59DFF9788385A82FED1FF4B8372" box="[151,243,259,285]" pageId="3" pageNumber="290">Carnian</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |