465 lines
91 KiB
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465 lines
91 KiB
XML
<document ID-DOI="10.1093/zoolinnean/zlac082" ID-GBIF-Dataset="f791f8ed-ae19-41a1-b5cc-b9b2cc89dec4" ID-ISSN="0024-4082" ID-Zenodo-Dep="7814127" ID-ZooBank="99842C3A-879F-4552-96B7-204D79CF3EDF" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="31" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="13" approvalRequired_for_textStreams="11" approvalRequired_for_treatments="6" checkinTime="1680597737982" checkinUser="plazi" docAuthor="Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam & Fikáček, Martin" docDate="2023" docId="03C2CE00FFF7FF8E3D7C0ECEFD47457E" docLanguage="en" docName="zlac082.pdf" docOrigin="Zoological Journal of the Linnean Society 197 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlac082" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Karumbe Arriaga-Varela, Tomaszewska, Szawaryn, Robertson, Seidel, Ślipiński & Fikáček, 2023, GEN. NOV." docType="treatment" docVersion="3" lastPageNumber="1099" masterDocId="FFFBB678FFF9FF9B3E190B0AFFBD416E" masterDocTitle="The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence" masterLastPageNumber="1115" masterPageNumber="1078" pageNumber="1092" updateTime="1681148941709" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence</mods:title>
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<mods:namePart>Arriaga-Varela, Emmanuel</mods:namePart>
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<mods:namePart>Tomaszewska, Wioletta</mods:namePart>
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<mods:namePart>Szawaryn, Karol</mods:namePart>
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<mods:namePart>Robertson, James</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Seidel, Matthias</mods:namePart>
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<mods:namePart>Ślipiński, Adam</mods:namePart>
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<mods:namePart>Fikáček, Martin</mods:namePart>
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<mods:date>2023</mods:date>
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<mods:number>2022-11-24</mods:number>
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<mods:number>197</mods:number>
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<mods:url>http://dx.doi.org/10.1093/zoolinnean/zlac082</mods:url>
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<subSubSection pageId="14" pageNumber="1092" type="nomenclature">
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<paragraph blockId="14.[869,1401,1475,1531]" pageId="14" pageNumber="1092">
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<heading box="[869,1401,1475,1500]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="1092" reason="2">
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<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[869,986,1476,1499]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1113" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[869,986,1476,1499]" italics="true" pageId="14" pageNumber="1092">KARUMBE</emphasis>
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</taxonomicName>
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ARRIAGA- VARELA, TOMASZEWSKA &
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</heading>
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<heading box="[1011,1258,1506,1531]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="1092" reason="4">
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SZAWARYN
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<emphasis bold="true" box="[1139,1258,1506,1530]" pageId="14" pageNumber="1092">
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<taxonomicNameLabel box="[1139,1258,1506,1530]" pageId="14" pageNumber="1092" rank="genus">GEN. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection pageId="14" pageNumber="1092" type="description">
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<paragraph blockId="14.[1068,1201,1545,1570]" box="[1068,1201,1545,1570]" pageId="14" pageNumber="1092">
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(
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<figureCitation box="[1077,1192,1545,1570]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStart-3="Figure 9" captionStart-4="Figure 10" captionStart-5="Figure 11" captionStartId-0="15.[145,223,1742,1764]" captionStartId-1="16.[164,242,1280,1302]" captionStartId-2="17.[146,226,1180,1202]" captionStartId-3="18.[164,244,1685,1707]" captionStartId-4="19.[145,224,1601,1623]" captionStartId-5="20.[164,243,1731,1753]" captionTargetBox-0="[321,1262,200,1690]" captionTargetBox-1="[323,1279,211,1237]" captionTargetBox-2="[325,1255,199,1114]" captionTargetBox-3="[164,1444,196,1646]" captionTargetBox-4="[145,1425,195,1561]" captionTargetBox-5="[244,1364,194,1692]" captionTargetId-0="figure-73@15.[302,1268,197,1702]" captionTargetId-1="figure-333@16.[319,1285,197,1240]" captionTargetId-2="figure-337@17.[302,1268,195,1140]" captionTargetId-3="figure-85@18.[164,1444,196,1646]" captionTargetId-4="figure-122@19.[145,1425,195,1561]" captionTargetId-5="figure-26@20.[244,1364,194,1692]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionTargetPageId-5="20" captionText-0="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." captionText-1="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." captionText-2="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." captionText-3="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." captionText-4="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." captionText-5="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi-0="http://doi.org/10.5281/zenodo.7814141" figureDoi-1="http://doi.org/10.5281/zenodo.7814143" figureDoi-2="http://doi.org/10.5281/zenodo.7814145" figureDoi-3="http://doi.org/10.5281/zenodo.7814149" figureDoi-4="http://doi.org/10.5281/zenodo.7814151" figureDoi-5="http://doi.org/10.5281/zenodo.7814155" httpUri-0="https://zenodo.org/record/7814141/files/figure.png" httpUri-1="https://zenodo.org/record/7814143/files/figure.png" httpUri-2="https://zenodo.org/record/7814145/files/figure.png" httpUri-3="https://zenodo.org/record/7814149/files/figure.png" httpUri-4="https://zenodo.org/record/7814151/files/figure.png" httpUri-5="https://zenodo.org/record/7814155/files/figure.png" pageId="14" pageNumber="1092">FIGS 6–11</figureCitation>
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)
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</paragraph>
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<paragraph blockId="14.[827,1444,1585,1638]" pageId="14" pageNumber="1092">
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<emphasis box="[827,1161,1585,1607]" italics="true" pageId="14" pageNumber="1092">Zoobank registration:</emphasis>
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urn: lsid: zoobank. org:act:
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<uuid box="[903,1421,1616,1638]" pageId="14" pageNumber="1092">50E490DD-3714-457E-9E27-69DABA334956</uuid>
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:
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="14" pageNumber="1092" type="type_taxon">
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<paragraph blockId="14.[827,1442,1662,1715]" pageId="14" pageNumber="1092">
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||
<emphasis box="[827,1201,1662,1684]" italics="true" pageId="14" pageNumber="1092">
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||
<typeStatus box="[827,883,1663,1684]" pageId="14" pageNumber="1092">Type</typeStatus>
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species:
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<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[999,1201,1662,1684]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1092" phylum="Arthropoda" rank="species" species="geiseri" status="sp. nov.">Karumbe geiseri</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[1211,1302,1663,1684]" pageId="14" pageNumber="1092" rank="species">sp. nov.</taxonomicNameLabel>
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by present designation.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="16" lastPageNumber="1094" pageId="14" pageNumber="1092" type="diagnosis">
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||
<paragraph blockId="14.[827,1444,1753,1898]" lastBlockId="16.[163,780,1428,1910]" lastPageId="16" lastPageNumber="1094" pageId="14" pageNumber="1092">
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<emphasis box="[827,1097,1753,1775]" italics="true" pageId="14" pageNumber="1092">
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Diagnosis:
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<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[984,1097,1753,1774]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1092" phylum="Arthropoda" rank="genus">Karumbe</taxonomicName>
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</emphasis>
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can be distinguished from other genera in
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<taxonomicName authorityName="BRETHES" authorityYear="1925" box="[1016,1246,1784,1806]" class="Insecta" family="Cerasommatidiidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1092" phylum="Arthropoda" rank="family">Cerasommatidiidae</taxonomicName>
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by the following combination of characters: dorsal surfaces of head, pronotum and elytra homogeneously covered by moderately long suberect setae (
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<figureCitation box="[1253,1384,1876,1898]" captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="14" pageNumber="1092">Figs 6A–F</figureCitation>
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,
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<figureCitation box="[1402,1435,1876,1897]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="14" pageNumber="1092">9E</figureCitation>
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,
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<figureCitation box="[145,191,1827,1848]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="15" pageNumber="1093">10C</figureCitation>
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,
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||
<figureCitation box="[207,288,1827,1848]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="15" pageNumber="1093">11B, D</figureCitation>
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); lateral margins of pronotum with an internal subparallel carina demarcating a wide area, and anterior pronotal margin bordered with internally crenulate bordering line (
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<figureCitation box="[1261,1355,1826,1848]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="15" pageNumber="1093">Figs 9C</figureCitation>
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,
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<figureCitation box="[1371,1418,1827,1848]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="15" pageNumber="1093">10C</figureCitation>
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||
,
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||
<figureCitation box="[809,856,1857,1878]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="15" pageNumber="1093">11B</figureCitation>
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); prosternal process moderately to distinctly wide, possessing long lateral carinae extending anteriorly much beyond prosternal process (
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<figureCitation box="[686,773,1428,1450]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="16" pageNumber="1094">Figs 7F</figureCitation>
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,
|
||
<figureCitation box="[163,196,1459,1480]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">9D</figureCitation>
|
||
,
|
||
<figureCitation box="[211,258,1459,1480]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="16" pageNumber="1094">10B</figureCitation>
|
||
,
|
||
<figureCitation box="[273,319,1459,1480]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="16" pageNumber="1094">11C</figureCitation>
|
||
); trochanters flattened and expanded posteriorly to cover the tibio-tarsal joint (
|
||
<figureCitation box="[656,772,1489,1511]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="16" pageNumber="1094">Figs 7I, K</figureCitation>
|
||
,
|
||
<figureCitation box="[163,190,1520,1541]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">9F</figureCitation>
|
||
,
|
||
<figureCitation box="[204,279,1520,1541]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="16" pageNumber="1094">10E, F</figureCitation>
|
||
,
|
||
<figureCitation box="[292,337,1520,1541]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="16" pageNumber="1094">11F</figureCitation>
|
||
); abdominal ventrite 1 with complete laterally, semicircular postcoxal lines (
|
||
<figureCitation box="[612,706,1551,1573]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="16" pageNumber="1094">Figs 7M</figureCitation>
|
||
,
|
||
<figureCitation box="[719,764,1551,1572]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="16" pageNumber="1094">11F</figureCitation>
|
||
); aedeagus with tegmen fused at base with penis (
|
||
<figureCitation box="[172,299,1612,1634]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="16" pageNumber="1094">Fig. 8B, E</figureCitation>
|
||
). The crenulate bordering line of the anterior margin of the pronotum (
|
||
<figureCitation box="[562,651,1643,1665]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">Figs 9C</figureCitation>
|
||
,
|
||
<figureCitation box="[665,711,1643,1664]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="16" pageNumber="1094">10C</figureCitation>
|
||
,
|
||
<figureCitation box="[724,770,1643,1664]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="16" pageNumber="1094">11B</figureCitation>
|
||
) (simple in
|
||
<taxonomicName box="[291,489,1673,1694]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[291,489,1673,1694]" italics="true" pageId="16" pageNumber="1094">Cerasommatidia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[505,586,1674,1695]" class="Insecta" family="Mycetaeidae" genus="Yamuy" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[505,586,1674,1695]" italics="true" pageId="16" pageNumber="1094">Yamuy</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[647,763,1673,1694]" class="Insecta" family="Eupsilobiidae" genus="Mahavelo" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[647,763,1673,1694]" italics="true" pageId="16" pageNumber="1094">MahaƲelo</emphasis>
|
||
</taxonomicName>
|
||
), distinctly and comparatively widely bordered pronotal base and the homogeneous moderately long suberect setae on dorsal surface are peculiar to this genus (tiny inconspicuous setae are present in
|
||
<taxonomicName box="[194,389,1827,1848]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[194,389,1827,1848]" italics="true" pageId="16" pageNumber="1094">Cerasommatidia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[449,563,1827,1848]" class="Insecta" family="Eupsilobiidae" genus="Mahavelo" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[449,563,1827,1848]" italics="true" pageId="16" pageNumber="1094">MahaƲelo</emphasis>
|
||
</taxonomicName>
|
||
, while
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[650,730,1827,1848]" class="Insecta" family="Mycetaeidae" genus="Yamuy" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[650,730,1827,1848]" italics="true" pageId="16" pageNumber="1094">Yamuy</emphasis>
|
||
</taxonomicName>
|
||
has two kinds of vestiture: tiny decumbent hairs and sparse, long erect setae).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814141" ID-Zenodo-Dep="7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="15" pageNumber="1093" startId="15.[145,223,1742,1764]" targetBox="[321,1262,200,1690]" targetPageId="15">
|
||
<paragraph blockId="15.[145,1426,1742,1794]" pageId="15" pageNumber="1093">
|
||
<emphasis bold="true" box="[145,248,1742,1764]" pageId="15" pageNumber="1093">Figure 6.</emphasis>
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[259,355,1742,1764]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1093" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[259,355,1742,1764]" italics="true" pageId="15" pageNumber="1093">Karumbe</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[361,450,1742,1764]" pageId="15" pageNumber="1093" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
A, B,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[510,693,1742,1764]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1093" phylum="Arthropoda" rank="species" species="brethesi" status="sp. nov.">
|
||
<emphasis box="[510,693,1742,1764]" italics="true" pageId="15" pageNumber="1093">Karumbe brethesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[699,778,1742,1764]" pageId="15" pageNumber="1093" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: A, habitus dorsal. B, habitus lateral. C, D,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[1222,1391,1742,1764]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1093" phylum="Arthropoda" rank="species" species="geiseri" status="sp. nov.">
|
||
<emphasis box="[1222,1391,1742,1764]" italics="true" pageId="15" pageNumber="1093">Karumbe geiseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="15" pageNumber="1093" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: C, habitus dorsal. D, habitus lateral. E, F,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[611,801,1772,1794]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1093" phylum="Arthropoda" rank="species" species="pakaluki" status="sp. nov.">
|
||
<emphasis box="[611,801,1772,1794]" italics="true" pageId="15" pageNumber="1093">Karumbe pakaluki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[806,882,1772,1794]" pageId="15" pageNumber="1093" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814143" ID-Zenodo-Dep="7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="16" pageNumber="1094" startId="16.[164,242,1280,1302]" targetBox="[323,1279,211,1237]" targetPageId="16">
|
||
<paragraph blockId="16.[163,1443,1280,1390]" pageId="16" pageNumber="1094">
|
||
<emphasis bold="true" box="[164,268,1280,1302]" pageId="16" pageNumber="1094">Figure 7.</emphasis>
|
||
Morphology of
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[433,530,1280,1302]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[433,530,1280,1302]" italics="true" pageId="16" pageNumber="1094">Karumbe</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[536,626,1280,1302]" pageId="16" pageNumber="1094" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
A–E, G, J–K, M,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[804,1000,1280,1302]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="species" species="pakaluki" status="sp. nov.">
|
||
<emphasis box="[804,1000,1280,1302]" italics="true" pageId="16" pageNumber="1094">Karumbe pakaluki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1007,1084,1280,1302]" pageId="16" pageNumber="1094" rank="species">sp. nov.</taxonomicNameLabel>
|
||
F, I, L,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[1160,1331,1280,1302]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="species" species="geiseri" status="sp. nov.">
|
||
<emphasis box="[1160,1331,1280,1302]" italics="true" pageId="16" pageNumber="1094">Karumbe geiseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1337,1413,1280,1302]" pageId="16" pageNumber="1094" rank="species">sp. nov.</taxonomicNameLabel>
|
||
H,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[163,347,1309,1331]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1094" phylum="Arthropoda" rank="species" species="brethesi" status="sp. nov.">
|
||
<emphasis box="[163,347,1309,1331]" italics="true" pageId="16" pageNumber="1094">Karumbe brethesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[353,429,1309,1331]" pageId="16" pageNumber="1094" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="21" lastPageNumber="1099" pageId="16" pageNumber="1094" type="description">
|
||
<paragraph blockId="16.[827,1443,1428,1910]" pageId="16" pageNumber="1094">
|
||
<emphasis box="[827,968,1429,1450]" italics="true" pageId="16" pageNumber="1094">Description:</emphasis>
|
||
Length
|
||
<quantity box="[1075,1244,1428,1450]" metricMagnitude="-3" metricUnit="m" metricValue="1.175" metricValueMax="1.45" metricValueMin="0.9" pageId="16" pageNumber="1094" unit="mm" value="1.175" valueMax="1.45" valueMin="0.9">0.90–1.45 mm</quantity>
|
||
. Body short oval and moderately convex, 1.25–1.30 times as long as wide, around 2.0 times as long as high. Coloration light to dark brown, usually with paler legs, antennae and palpi (
|
||
<figureCitation box="[898,1008,1550,1573]" captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="16" pageNumber="1094">Fig. 6A–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="16.[827,1443,1428,1910]" lastBlockId="17.[145,761,1312,1886]" lastPageId="17" lastPageNumber="1095" pageId="16" pageNumber="1094">
|
||
Head with dorsal surface homogeneously covered with moderately long setae (
|
||
<figureCitation box="[1166,1251,1612,1634]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">Fig. 9A</figureCitation>
|
||
). Clypeus large, rectangular (
|
||
<figureCitation box="[982,1067,1643,1665]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">Fig. 9A</figureCitation>
|
||
). Frontoclypeal suture distinct, straight. Ventral antennal grooves short, not extending to posterior eye level (
|
||
<figureCitation box="[1075,1154,1704,1726]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">Fig. 9A</figureCitation>
|
||
). Antenna approximately 0.35–0.40 of body length; antennal club (
|
||
<figureCitation box="[1289,1376,1735,1757]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="16" pageNumber="1094">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation box="[1389,1434,1735,1756]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="16" pageNumber="1094">11A</figureCitation>
|
||
) one quarter of total antennal length; antennomeres 1–5 longer than wide, with antennomere 5 longer than neighbouring antennomeres 4 and 6; antennomeres 6–9 variable. Eyes comparatively large, moderately prominent (
|
||
<figureCitation box="[965,1056,1888,1910]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="16" pageNumber="1094">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation box="[1070,1102,1888,1909]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="16" pageNumber="1094">9A</figureCitation>
|
||
). Galea large, densely setose at apex (
|
||
<figureCitation box="[244,325,1312,1334]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Fig. 7C</figureCitation>
|
||
). Lacinia weakly narrower than galea with dense setae at apex and inner margin. Labium with mentum widest near midlength or basal third; terminal palpomere short and rounded to elongate and acuminate, 1.2–2.2 times as long as wide at base (
|
||
<figureCitation captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Figs 7D</figureCitation>
|
||
,
|
||
<figureCitation box="[189,219,1466,1487]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="17" pageNumber="1095">9B</figureCitation>
|
||
,
|
||
<figureCitation box="[232,277,1466,1487]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="17" pageNumber="1095">10A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814145" ID-Zenodo-Dep="7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="17" pageNumber="1095" startId="17.[146,226,1180,1202]" targetBox="[325,1255,199,1114]" targetPageId="17">
|
||
<paragraph blockId="17.[145,1425,1180,1261]" pageId="17" pageNumber="1095">
|
||
<emphasis bold="true" box="[146,253,1180,1202]" pageId="17" pageNumber="1095">Figure 8.</emphasis>
|
||
Terminalia of
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[414,512,1180,1202]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1095" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[414,512,1180,1202]" italics="true" pageId="17" pageNumber="1095">Karumbe</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[519,612,1180,1202]" pageId="17" pageNumber="1095" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
A–C,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[675,849,1180,1202]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1095" phylum="Arthropoda" rank="species" species="geiseri" status="sp. nov.">
|
||
<emphasis box="[675,849,1180,1202]" italics="true" pageId="17" pageNumber="1095">Karumbe geiseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[856,937,1180,1202]" pageId="17" pageNumber="1095" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[634,831,1209,1231]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1095" phylum="Arthropoda" rank="species" species="pakaluki" status="sp. nov.">
|
||
<emphasis box="[634,831,1209,1231]" italics="true" pageId="17" pageNumber="1095">Karumbe pakaluki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[838,917,1209,1231]" pageId="17" pageNumber="1095" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G,
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[845,1029,1239,1261]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1095" phylum="Arthropoda" rank="species" species="brethesi" status="sp. nov.">
|
||
<emphasis box="[845,1029,1239,1261]" italics="true" pageId="17" pageNumber="1095">Karumbe brethesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1036,1114,1239,1261]" pageId="17" pageNumber="1095" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="17.[145,761,1312,1886]" lastBlockId="17.[809,1426,1312,1886]" pageId="17" pageNumber="1095">
|
||
Prothorax. Pronotum 2.1–2.2 times as wide as long, widest at base and strongly convergent anteriorly, 1.7– 1.9 times wider at widest part than on front angles (
|
||
<figureCitation box="[154,245,1588,1610]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="17" pageNumber="1095">Figs 9C</figureCitation>
|
||
,
|
||
<figureCitation box="[259,305,1588,1609]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="17" pageNumber="1095">10C</figureCitation>
|
||
,
|
||
<figureCitation box="[320,367,1588,1609]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="17" pageNumber="1095">11B</figureCitation>
|
||
). Anterior margin bordered with densely crenulate bordering line; base of pronotum sinuate with margin comparatively widely bordered with less distinctly crenulate, well-developed bordering line. Pronotal sides scarcely rounded, area between lateral edge and carina weakly to distinctly concave, wide and continuously weakly narrowing posteriorly; hind angles with small, oblique indentation to receive humeral corner of elytron. Pronotal disc moderately convex, with sparse punctures provided with moderately long, thick setae. Prosternum with anterior margin scarcely arcuate posteriorly (
|
||
<figureCitation captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Figs 7F</figureCitation>
|
||
,
|
||
<figureCitation box="[852,884,1373,1394]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="17" pageNumber="1095">9D</figureCitation>
|
||
,
|
||
<figureCitation box="[899,945,1373,1394]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="17" pageNumber="1095">10B</figureCitation>
|
||
,
|
||
<figureCitation box="[960,1006,1373,1394]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="17" pageNumber="1095">11C</figureCitation>
|
||
); prosternal process comparatively wide, at apex about 0.7–0.9 of width of procoxal cavity, with raised lateral carinae extending anteriorly well beyond level of anterior margin of procoxal cavities; with area between carinae weakly concave; prosternal process extending posteriorly to hind level of procoxae. Hypomeron with deep and long, weakly sinuate antennal grooves (
|
||
<figureCitation box="[1019,1101,1588,1610]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Figs 7F</figureCitation>
|
||
,
|
||
<figureCitation box="[1114,1146,1588,1609]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="17" pageNumber="1095">9D</figureCitation>
|
||
,
|
||
<figureCitation box="[1159,1204,1588,1609]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="17" pageNumber="1095">10B</figureCitation>
|
||
,
|
||
<figureCitation box="[1216,1262,1588,1609]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="17" pageNumber="1095">11C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="17.[809,1426,1312,1886]" lastBlockId="19.[809,1424,1728,1903]" lastPageId="19" lastPageNumber="1097" pageId="17" pageNumber="1095">
|
||
Pterothorax.Scutellar shield small, transverse, weakly rounded at apex (
|
||
<figureCitation box="[1004,1084,1649,1671]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Fig. 7G</figureCitation>
|
||
). Mesoventrite with intercoxal process smooth, almost flat with anterior raised border incomplete medially (
|
||
<figureCitation box="[1066,1149,1711,1733]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Figs 7I</figureCitation>
|
||
,
|
||
<figureCitation box="[1165,1193,1711,1732]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="17" pageNumber="1095">9F</figureCitation>
|
||
,
|
||
<figureCitation box="[1208,1254,1711,1732]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="17" pageNumber="1095">10E</figureCitation>
|
||
,
|
||
<figureCitation box="[1269,1315,1711,1732]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="17" pageNumber="1095">11C</figureCitation>
|
||
), 1.1–1.4 times as wide as mesocoxal diameter. Metaventrite as long as abdominal ventrites 1–3 together; with remnants of short discrimen (
|
||
<figureCitation box="[1148,1221,1803,1825]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="17" pageNumber="1095">Fig. 7I</figureCitation>
|
||
); anterior margin bordered with carina weekly widening towards lateral corners, at intercoxal process bordering line weakly crenulate; central area with small to large and deep setiferous punctures. Anterior part of metanepisternum with small outer blunt projection (
|
||
<figureCitation box="[594,675,1863,1885]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="18" pageNumber="1096">Fig. 7I</figureCitation>
|
||
). Elytra
|
||
<quantity box="[827,997,1801,1823]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.35" metricValueMin="0.65" pageId="18" pageNumber="1096" unit="mm" value="1.0" valueMax="1.35" valueMin="0.65">0.65–1.35 mm</quantity>
|
||
long, about as long as wide, 2.7–3.1 times as long and 1.4 times as wide as pronotum, with lateral margins scarcely to not explanate (
|
||
<figureCitation box="[1292,1428,1863,1885]" captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="18" pageNumber="1096">Fig. 6A, C, E</figureCitation>
|
||
). Surface with moderately dense and deep setiferous punctures (
|
||
<figureCitation box="[287,381,1758,1780]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="19" pageNumber="1097">Figs 9E</figureCitation>
|
||
,
|
||
<figureCitation box="[398,447,1758,1779]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="19" pageNumber="1097">11D</figureCitation>
|
||
), deeper and larger than those on pronotum, bearing moderately long suberect setae; with additional intermixed shallow foveolate punctures. Epipleura moderately wide, narrowing to apex, complete at apex, with internal bordering line narrow, present from the level of mid coxae to apex. Hindwings well developed, without anal lobe, with single reduced anal vein, and median fleck present, undivided (
|
||
<figureCitation box="[940,1025,1820,1842]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="19" pageNumber="1097">Fig. 7H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814149" ID-Zenodo-Dep="7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="18" pageNumber="1096" startId="18.[164,244,1685,1707]" targetBox="[164,1444,196,1646]" targetPageId="18">
|
||
<paragraph blockId="18.[164,1444,1685,1766]" pageId="18" pageNumber="1096">
|
||
<emphasis bold="true" box="[164,271,1685,1707]" pageId="18" pageNumber="1096">Figure 9.</emphasis>
|
||
Morphology of
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[445,637,1685,1707]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1096" phylum="Arthropoda" rank="species" species="brethesi" status="sp. nov.">
|
||
<emphasis box="[445,637,1685,1707]" italics="true" pageId="18" pageNumber="1096">Karumbe brethesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[645,724,1685,1707]" pageId="18" pageNumber="1096" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814151" ID-Zenodo-Dep="7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="19" pageNumber="1097" startId="19.[145,224,1601,1623]" targetBox="[145,1425,195,1561]" targetPageId="19">
|
||
<paragraph blockId="19.[145,1425,1601,1681]" pageId="19" pageNumber="1097">
|
||
<emphasis bold="true" box="[145,263,1601,1623]" pageId="19" pageNumber="1097">Figure 10.</emphasis>
|
||
Morphology of
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[431,602,1601,1623]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1097" phylum="Arthropoda" rank="species" species="geiseri" status="sp. nov.">
|
||
<emphasis box="[431,602,1601,1623]" italics="true" pageId="19" pageNumber="1097">Karumbe geiseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[609,685,1601,1623]" pageId="19" pageNumber="1097" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="19.[809,1424,1728,1903]" lastBlockId="21.[145,762,197,802]" lastPageId="21" lastPageNumber="1099" pageId="19" pageNumber="1097">
|
||
Legs. Trochanters roundly or angulately produced posteriorly (
|
||
<figureCitation box="[967,1100,1881,1903]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="19" pageNumber="1097">Figs 9D, F</figureCitation>
|
||
,
|
||
<figureCitation box="[1118,1200,1881,1902]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="19" pageNumber="1097">10E, F</figureCitation>
|
||
,
|
||
<figureCitation box="[1218,1265,1881,1902]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="19" pageNumber="1097">11F</figureCitation>
|
||
). Meso- and metatrochanters with posterior margin as a laminar expansion that covers tibio-tarsal joint when retracted. Femora flattened (mid and hind femora more distinctly than fore femora) with grooves for tibiae present throughout whole length. Claws with distinct quadrate tooth at base (
|
||
<figureCitation box="[303,381,350,372]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="21" pageNumber="1099">Fig. 7L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814155" ID-Zenodo-Dep="7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="20" pageNumber="1098" startId="20.[164,243,1731,1753]" targetBox="[244,1364,194,1692]" targetPageId="20">
|
||
<paragraph blockId="20.[164,1444,1731,1841]" pageId="20" pageNumber="1098">
|
||
<emphasis bold="true" box="[164,283,1731,1753]" pageId="20" pageNumber="1098">Figure 11.</emphasis>
|
||
Morphology of
|
||
<taxonomicName authority="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček, 2023" authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[452,650,1731,1753]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1098" phylum="Arthropoda" rank="species" species="pakaluki" status="sp. nov.">
|
||
<emphasis box="[452,650,1731,1753]" italics="true" pageId="20" pageNumber="1098">Karumbe pakaluki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[657,734,1731,1753]" pageId="20" pageNumber="1098" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="21.[145,762,197,802]" pageId="21" pageNumber="1099">
|
||
Abdomen with ventrite 1 anteriorly with marginal carina forming distinct, rounded and complete laterally postcoxal lines, extending posteriorly to nearly midlength of ventrite 1 (
|
||
<figureCitation box="[419,511,473,495]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="21" pageNumber="1099">Figs 7M</figureCitation>
|
||
,
|
||
<figureCitation box="[524,567,473,494]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="21" pageNumber="1099">11F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="21.[145,762,197,802]" pageId="21" pageNumber="1099">
|
||
Male genitalia (
|
||
<figureCitation box="[392,530,504,526]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="21" pageNumber="1099">Fig. 8B, E</figureCitation>
|
||
). Aedeagus with penis short and stout, sclerotized, curved apically, asymmetric, with ramificate apex (
|
||
<figureCitation box="[611,743,565,587]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="21" pageNumber="1099">Fig. 8B, E</figureCitation>
|
||
). Tegmen fused to penis, submembranous, throneshaped, short or basally elongate; tegminal strut short, reduced.
|
||
</paragraph>
|
||
<paragraph blockId="21.[145,762,197,802]" pageId="21" pageNumber="1099">
|
||
Female genitalia. Spermatheca moderately large, elongate-reniform, submembranous; sperm duct short; accessory gland small membranous, of irregular shape (
|
||
<figureCitation box="[153,289,780,802]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="21" pageNumber="1099">Fig. 8C, F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="21" pageNumber="1099" type="etymology">
|
||
<paragraph blockId="21.[145,761,840,892]" pageId="21" pageNumber="1099">
|
||
<emphasis box="[145,396,840,861]" italics="true" pageId="21" pageNumber="1099">
|
||
Etymology:
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[291,396,840,861]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1099" phylum="Arthropoda" rank="genus">Karumbe</taxonomicName>
|
||
</emphasis>
|
||
is the Guaraní word for ‘turtle’. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="21" pageNumber="1099" type="synonymic_list">
|
||
<paragraph blockId="21.[145,760,920,972]" pageId="21" pageNumber="1099">
|
||
<emphasis box="[145,578,920,941]" italics="true" pageId="21" pageNumber="1099">
|
||
Species included:
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[366,578,920,941]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1099" phylum="Arthropoda" rank="species" species="brethesi">Karumbe brethesi</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[595,707,920,941]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1099" phylum="Arthropoda" rank="species" species="geiseri">
|
||
<emphasis box="[595,707,920,941]" italics="true" pageId="21" pageNumber="1099">K. geiseri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis box="[145,283,950,972]" italics="true" pageId="21" pageNumber="1099">
|
||
<taxonomicName authorityName="Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček" authorityYear="2023" box="[145,279,950,972]" class="Insecta" family="Mycetaeidae" genus="Karumbe" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="1099" phylum="Arthropoda" rank="species" species="pakaluki">K. pakaluki</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[145,762,1018,1040]" pageId="21" pageNumber="1099" type="distribution">
|
||
<paragraph blockId="21.[145,762,1018,1040]" box="[145,762,1018,1040]" pageId="21" pageNumber="1099">
|
||
<emphasis box="[145,291,1018,1039]" italics="true" pageId="21" pageNumber="1099">Distribution:</emphasis>
|
||
<collectingCountry box="[303,375,1018,1040]" name="Brazil" pageId="21" pageNumber="1099">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry box="[384,485,1018,1040]" name="Grenada" pageId="21" pageNumber="1099">Grenada</collectingCountry>
|
||
,
|
||
<collectingCountry box="[494,610,1018,1040]" name="Venezuela" pageId="21" pageNumber="1099">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation box="[624,747,1018,1040]" captionStart="Figure 20" captionStartId="34.[162,240,1081,1103]" captionTargetBox="[162,1442,200,1042]" captionTargetId="figure-327@34.[162,1442,200,1042]" captionTargetPageId="34" captionText="Figure 20. Geographical distribution of Cerasommatidiidae spp. A, distribution of species in eastern Caribbean region. B, distribution of species in southeastern Brazil. C, distribution of species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.7814187" httpUri="https://zenodo.org/record/7814187/files/figure.png" pageId="21" pageNumber="1099">Fig. 20A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |