452 lines
54 KiB
XML
452 lines
54 KiB
XML
<document id="FB9E20B8F7F975104614ABBBB358DFED" ID-CLB-Dataset="46403" ID-DOI="10.5281/zenodo.206004" ID-GBIF-Dataset="cf42d2dc-94a9-4c2b-91c5-6059e0283e0a" ID-ISSN="1175-5326" ID-Zenodo-Dep="206004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460386165159" checkinUser="plazi" docAuthor="Hoffman, Richard L., Martinez, Daniela & D, Eduardo Florez" docDate="2011" docId="03984758FFBC8219FF35F95EFC8E816A" docLanguage="en" docName="zt03015p060.pdf" docOrigin="Zootaxa 3015" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Psammodesmus bryophorus Hoffman, Martinez & D, 2011, new species" docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="FFA13F20FFBD821EFFA2FF99FFFB876D" masterDocTitle="A new Colombian species in the milliped genus Psammodesmus, symbiotic host for bryophytes (Polydesmida: Platyrhacidae)" masterLastPageNumber="60" masterPageNumber="52" pageNumber="53" updateTime="1698297596396" updateUser="ExternalLinkService">
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<mods:mods id="CB8DFFB3326AB348B4F65E10367A24AF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2A6EFC870BB1BB782FED25F09B9C7034">
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<mods:title id="5B967A703B4204EF98D5756D92AEDA12">A new Colombian species in the milliped genus Psammodesmus, symbiotic host for bryophytes (Polydesmida: Platyrhacidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="750298BD7B6587B32CB3581EC1DB9D08" type="personal">
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<mods:role id="95707DE6CECF11C81B6B66794F9B8D99">
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<mods:roleTerm id="FF3543EFA39AB2E041BE0A30B79EFA1A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C6489E66DA726C1AFF991ED8362F19F7">Hoffman, Richard L.</mods:namePart>
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</mods:name>
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<mods:name id="71054A8CEC2FA3A7A9EE36AA3CC7682B" type="personal">
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<mods:role id="84931A88C34D368A04834F4831DF0332">
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<mods:roleTerm id="76F9F4A461D064BD85898EAD355472F7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA436FF4179E3E75E1DCB4CCC1DA037C">Martinez, Daniela</mods:namePart>
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</mods:name>
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<mods:name id="6556CFF7E064AB5386374A82CB4F3941" type="personal">
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<mods:role id="F84300F87B7C9EAF2ECB3AD468F8C2AE">
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<mods:roleTerm id="5B34DF51F3728E61C4046FA831F1B000">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="205683DB961CCB9E005A3F6C3158A971">D, Eduardo Florez</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8254881AD171EFDC2886F562BF0251B8">text</mods:typeOfResource>
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<mods:relatedItem id="323934FB047A29C74C7F827398B900FE" type="host">
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<mods:titleInfo id="48DD57391717D8CC1D0870D24F628AEE">
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<mods:title id="292B04AC967D8FB25EB988CE7A7CD089">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B1E3F6302790494868D2318E2BD274C7">
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<mods:date id="96B639D65B915435EB3D7CA9CC662951">2011</mods:date>
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<mods:detail id="19C2E98656B73D526C85F7C2E2E1AEC5" type="volume">
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<mods:number id="A7E0426043B2CA2E8DBDD3B1430BD252">3015</mods:number>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="F1A172E4DD3E2DD69A8AD33143892942">journal article</mods:classification>
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<mods:identifier id="3C1C25B2FD6B5D112A06444F37C240F6" type="CLB-Dataset">46403</mods:identifier>
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<mods:identifier id="B08841F1BD2855F9DC9F010BC871CA0E" type="DOI">10.5281/zenodo.206004</mods:identifier>
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<mods:identifier id="E7B8F66FD13176620C1C5C74338D9F2A" type="GBIF-Dataset">cf42d2dc-94a9-4c2b-91c5-6059e0283e0a</mods:identifier>
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<mods:identifier id="B24423AE3E5C299322D56374D70025AE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="65E7659234566110BC1961AAD914F1B6" type="Zenodo-Dep">206004</mods:identifier>
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<treatment id="03984758FFBC8219FF35F95EFC8E816A" ID-DOI="http://doi.org/10.5281/zenodo.6195192" ID-GBIF-Taxon="119415513" ID-Zenodo-Dep="6195192" LSID="urn:lsid:plazi:treatment:03984758FFBC8219FF35F95EFC8E816A" httpUri="http://treatment.plazi.org/id/03984758FFBC8219FF35F95EFC8E816A" lastPageId="7" lastPageNumber="59" pageId="1" pageNumber="53">
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<subSubSection id="C32BA5C5FFBC821FFF35F95EFE32806F" pageId="1" pageNumber="53" type="nomenclature">
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<paragraph id="8B8EF64EFFBC821FFF35F95EFD72818C" blockId="1.[151,649,1735,1794]" box="[151,649,1735,1761]" pageId="1" pageNumber="53">
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<heading id="D0C64122FFBC821FFF35F95EFD72818C" bold="true" box="[151,649,1735,1761]" fontSize="11" level="1" pageId="1" pageNumber="53" reason="1">
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<emphasis id="B9452A5CFFBC821FFF35F95EFD72818C" bold="true" box="[151,649,1735,1761]" pageId="1" pageNumber="53">
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<taxonomicName id="4C318DCDFFBC821FFF35F95EFE11818C" ID-CoL="8Q8YT" box="[151,490,1735,1761]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="bryophorus" status="sp. nov.">
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<emphasis id="B9452A5CFFBC821FFF35F95EFE11818C" bold="true" box="[151,490,1735,1761]" italics="true" pageId="1" pageNumber="53">Psammodesmus bryophorus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2769727FFBC821FFE5BF95EFD72818C" box="[505,649,1735,1761]" pageId="1" pageNumber="53" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8EF64EFFBC821FFF35F973FE32806F" blockId="1.[151,649,1735,1794]" box="[151,457,1770,1794]" pageId="1" pageNumber="53">
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<figureCitation id="130AEACBFFBC821FFF35F973FF07806F" box="[151,252,1770,1794]" captionStart="FIGURE 1" captionStartId="2.[151,250,1066,1089]" captionTargetBox="[191,1377,664,951]" captionTargetId="figure@2.[151,1436,574,1044]" captionTargetPageId="2" captionText="FIGURE 1. Psammodesmus bryophorus, n. sp. Holotype, dorsal aspect (photo by Melody Cartwright. VMNH)." httpUri="https://zenodo.org/record/206005/files/figure.png" pageId="1" pageNumber="53">Figures 1</figureCitation>
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–6, 11, 12, 14, 15.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32BA5C5FFBC821CFF35F8ABFEE48671" lastPageId="2" lastPageNumber="54" pageId="1" pageNumber="53" type="materials_examined">
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<paragraph id="8B8EF64EFFBC821CFF35F8ABFB6387B9" blockId="1.[151,1436,1842,2011]" lastBlockId="2.[151,1436,151,541]" lastPageId="2" lastPageNumber="54" pageId="1" pageNumber="53">
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<emphasis id="B9452A5CFFBC821FFF35F8ABFEFD8026" bold="true" box="[151,262,1842,1867]" pageId="1" pageNumber="53">Material.</emphasis>
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Male
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<typeStatus id="548A48ECFFBC821FFEF1F8ABFE4E8027" box="[339,437,1842,1866]" pageId="1" pageNumber="53" type="holotype">holotype</typeStatus>
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(ICN-MD 316) from
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<collectingCountry id="F326B6DEFFBC821FFD15F8ABFCAB8027" box="[695,848,1842,1866]" name="Colombia" pageId="1" pageNumber="53">COLOMBIA</collectingCountry>
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, Departamento Nariňo, Municipio Barbacoas, Corregimiento Altaquer, Reserva Natural Río Ñambí on Rt. 10, ca
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<quantity id="4CC95BABFFBC821FFCF5F8CFFC5A8003" box="[855,929,1878,1902]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="53" unit="km" value="50.0">50 km</quantity>
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. (airline) south of Barbacoas,
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<quantity id="4CC95BABFFBC821FFB4DF8CFFABF8003" box="[1263,1348,1878,1902]" metricMagnitude="5" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="53" unit="km" value="165.0">165 km</quantity>
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west of Pasto (
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<figureCitation id="130AEACBFFBC821FFF46F8E3FECC80FF" box="[228,311,1914,1938]" captionStart="FIGURE 16" captionStartId="6.[151,249,877,900]" captionTargetBox="[157,1424,202,845]" captionTargetId="figure@6.[151,1436,193,856]" captionTargetPageId="6" captionText="FIGURE 16. Southwestern Colombia, showing location of type locality (on left), boxed area in south central Nariño Department shown on right for closer detail." httpUri="https://zenodo.org/record/206007/files/figure.png" pageId="1" pageNumber="53">Fig. 16</figureCitation>
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),
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<quantity id="4CC95BABFFBC821FFEEFF8E3FE2080FF" box="[333,475,1914,1938]" metricMagnitude="7" metricUnit="m" metricValue="1.300145" pageId="1" pageNumber="53" unit="m" value="1.300145E7">13001450 m</quantity>
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[
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<geoCoordinate id="EE059089FFBC821FFE4EF8E3FDB480FF" box="[492,591,1914,1938]" direction="north" orientation="latitude" pageId="1" pageNumber="53" precision="925" value="1.3">1° 18’ N</geoCoordinate>
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&
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<geoCoordinate id="EE059089FFBC821FFDD7F8E3FD1680FF" box="[629,749,1914,1938]" direction="west" orientation="longitude" pageId="1" pageNumber="53" precision="925" value="-78.083336">78° 05’ W</geoCoordinate>
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,
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<date id="FF8FD08EFFBC821FFD5BF8E3FC6080FF" box="[761,923,1914,1938]" pageId="1" pageNumber="53" value="2010-04-16">16 April 2010</date>
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, E. Flórez and students in Animal Taxonomy course of the Universidad Nacional de
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<collectingCountry id="F326B6DEFFBC821FFDEEF807FD4580DB" box="[588,702,1950,1974]" name="Colombia" pageId="1" pageNumber="53">Colombia</collectingCountry>
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. Female
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<typeStatus id="548A48ECFFBC821FFC80F806FC7980DA" box="[802,898,1951,1975]" pageId="1" pageNumber="53" type="paratype">paratype</typeStatus>
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(ICN-MD 317) with same data except
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<date id="FF8FD08EFFBC821FFA9BF807FF0080B7" pageId="1" pageNumber="53" value="2009-10-16">16 October 2009</date>
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and male
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<typeStatus id="548A48ECFFBC821FFED1F85AFE2880B6" box="[371,467,1987,2011]" pageId="1" pageNumber="53" type="paratype">paratype</typeStatus>
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(ICN-MD 318) with same data except
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<date id="FF8FD08EFFBC821FFC31F85BFBA980B7" box="[915,1106,1986,2010]" pageId="1" pageNumber="53" value="2009-10-10">10 October 2009</date>
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are deposited in the Virginia Museum of Natural History.
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<specimenCount id="9D373DC7FFBF821CFE79FF0EFDCF87C2" box="[475,564,151,175]" pageId="2" pageNumber="54" type="male">14 male</specimenCount>
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and
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<specimenCount id="9D373DC7FFBF821CFDCEFF0EFD3687C2" box="[620,717,151,175]" pageId="2" pageNumber="54" type="female">1 female</specimenCount>
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<typeStatus id="548A48ECFFBF821CFD76FF01FCC487DD" box="[724,831,152,176]" pageId="2" pageNumber="54" type="paratype">paratypes</typeStatus>
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are preserved in the Arachnological Collection of the Instituto de Ciencias Naturales, Universidad Nacional de
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<collectingCountry id="F326B6DEFFBF821CFCB8FF25FC7787B9" box="[794,908,188,212]" name="Colombia" pageId="2" pageNumber="54">Colombia</collectingCountry>
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. (
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<collectionCode id="ED206E8BFFBF821CFC3DFF25FC2A87B9" box="[927,977,188,212]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="2" pageNumber="54">ICN</collectionCode>
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- MD 319 - 336).
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</paragraph>
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<paragraph id="8B8EF64EFFBF821CFF65FF79FEE48671" blockId="2.[151,1436,151,541]" pageId="2" pageNumber="54">
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<emphasis id="B9452A5CFFBF821CFF65FF79FEF68795" bold="true" box="[199,269,224,248]" pageId="2" pageNumber="54">Name</emphasis>
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: Greek
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<emphasis id="B9452A5CFFBF821CFECAFF79FE518795" box="[360,426,224,248]" italics="true" pageId="2" pageNumber="54">bryon</emphasis>
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(moss) +
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<emphasis id="B9452A5CFFBF821CFDB9FF79FD8F8795" box="[539,628,224,248]" italics="true" pageId="2" pageNumber="54">phoreus</emphasis>
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(bearer), with reference to the singular symbiotic relationship noted for this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32BA5C5FFBF821CFF65FEBEFC4886C1" pageId="2" pageNumber="54" type="diagnosis">
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<paragraph id="8B8EF64EFFBF821CFF65FEBEFC4886C1" blockId="2.[151,1436,151,541]" pageId="2" pageNumber="54">
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<emphasis id="B9452A5CFFBF821CFF65FEBEFEBA862D" bold="true" box="[199,321,295,320]" pageId="2" pageNumber="54">Diagnosis.</emphasis>
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A member of the
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<taxonomicName id="4C318DCDFFBF821CFDAEFEB0FD5A862D" box="[524,673,296,320]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="fasciolatus">
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<emphasis id="B9452A5CFFBF821CFDAEFEB0FD5A862D" box="[524,673,296,320]" italics="true" pageId="2" pageNumber="54">P. fasciolatus</emphasis>
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</taxonomicName>
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group: apical lamina of the gonotelopodite suboval, not triangularly produced ventrad; paranota narrow, directed posteriad, widely separated, elevated above level of middorsum; metaterga smooth, with paramedian oblique white lines continued parallel on prozona; anterior margin of paranota reflexed dorsad, the edge modified, appearing as a tightly twisted cord.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32BA5C5FFBF821BFF65FE22FBD480D2" lastPageId="5" lastPageNumber="57" pageId="2" pageNumber="54" type="materials_examined">
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<paragraph id="8B8EF64EFFBF821CFF65FE22FD5C8570" blockId="2.[151,1436,151,541]" pageId="2" pageNumber="54">
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<emphasis id="B9452A5CFFBF821CFF65FE22FEC086BE" bold="true" box="[199,315,443,467]" pageId="2" pageNumber="54">
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<typeStatus id="548A48ECFFBF821CFF65FE22FECD86BE" box="[199,310,443,467]" pageId="2" pageNumber="54" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult male, ca.
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<quantity id="4CC95BABFFBF821CFE54FE23FDBE86BF" box="[502,581,442,466]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="54" unit="mm" value="59.0">59 mm</quantity>
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long, widths of segments across paranota: collum,
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<quantity id="4CC95BABFFBF821CFB24FE23FB2486BF" box="[1158,1247,442,466]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="54" unit="mm" value="5.3">5.3 mm</quantity>
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, 2nd–4th,
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<quantity id="4CC95BABFFBF821CFA9DFE23FA6C86BF" box="[1343,1431,442,466]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="54" unit="mm" value="8.8">8.8 mm</quantity>
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||
, 6th,
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||
<quantity id="4CC95BABFFBF821CFF62FE78FEE28694" box="[192,281,481,505]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="54" unit="mm" value="9.1">9.1 mm</quantity>
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||
, 8th–14th, 9.0 mm, 16th,
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||
<quantity id="4CC95BABFFBF821CFD80FE78FD808694" box="[546,635,481,505]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="2" pageNumber="54" unit="mm" value="8.6">8.6 mm</quantity>
|
||
, 18th,
|
||
<quantity id="4CC95BABFFBF821CFD1DFE78FCE18694" box="[703,794,481,505]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="54" unit="mm" value="6.2">6.2 mm</quantity>
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||
. Body appearing slender, with large elevated and widely separated paranota, W/L ratio ca. 15% (
|
||
<figureCitation id="130AEACBFFBF821CFDF6FD9CFD6E8570" box="[596,661,517,541]" captionStart="FIGURE 1" captionStartId="2.[151,250,1066,1089]" captionTargetBox="[191,1377,664,951]" captionTargetId="figure@2.[151,1436,574,1044]" captionTargetPageId="2" captionText="FIGURE 1. Psammodesmus bryophorus, n. sp. Holotype, dorsal aspect (photo by Melody Cartwright. VMNH)." httpUri="https://zenodo.org/record/206005/files/figure.png" pageId="2" pageNumber="54">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4EA6C6FFBF821CFF35FBB3FAF0832D" box="[151,1291,1066,1089]" httpUri="https://zenodo.org/record/206005/files/figure.png" pageId="2" pageNumber="54" targetBox="[191,1377,664,951]" targetPageId="2">
|
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<paragraph id="8B8EF64EFFBF821CFF35FBB3FAF0832D" blockId="2.[151,1291,1066,1089]" box="[151,1291,1066,1089]" pageId="2" pageNumber="54">
|
||
<emphasis id="B9452A5CFFBF821CFF35FBB3FEE8832D" bold="true" box="[151,275,1066,1089]" pageId="2" pageNumber="54">FIGURE 1.</emphasis>
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||
<taxonomicName id="4C318DCDFFBF821CFEBBFBB3FDCD832D" box="[281,566,1066,1088]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="bryophorus" status="sp. nov.">
|
||
<emphasis id="B9452A5CFFBF821CFEBBFBB3FDCD832D" box="[281,566,1066,1088]" italics="true" pageId="2" pageNumber="54">Psammodesmus bryophorus</emphasis>
|
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</taxonomicName>
|
||
,
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||
<emphasis id="B9452A5CFFBF821CFDE0FBB2FD80832D" bold="true" box="[578,635,1067,1088]" pageId="2" pageNumber="54">
|
||
<taxonomicNameLabel id="A2769727FFBF821CFDE0FBB2FD80832D" box="[578,635,1067,1088]" pageId="2" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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Holotype, dorsal aspect (photo by Melody Cartwright. VMNH).
|
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</paragraph>
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</caption>
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<paragraph id="8B8EF64EFFBF821CFF65FBF4FE798278" blockId="2.[151,1437,1133,2027]" pageId="2" pageNumber="54">Entire dorsum piceous to dull black, ventral surfaces including legs reddish-brown. Posteror edge of collum, all margins of paranota, mandibles and epiproct ivory white. Metaterga with two oblique ivory lines, continuous or not across stricture, closer together and parallel on prozona. Anterior margin of collum with 4–5 prominent paramedian ovoid tubercles, and ca. 6-6 smaller subapical tubercles, both series white, paramedian stripes of collum small and broken up.</paragraph>
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<paragraph id="8B8EF64EFFBF821CFF65FAB8FE7F817A" blockId="2.[151,1437,1133,2027]" pageId="2" pageNumber="54">
|
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Head
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<quantity id="4CC95BABFFBF821CFEA8FAB8FE9A8254" box="[266,353,1313,1337]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="54" unit="mm" value="4.1">4.1 mm</quantity>
|
||
across genal apices. Labrum glabrous, deeply offset from level of clypeus, with 7-7 compound setae, facial setae: 4-4 labral, 2-2 clypeal, 1-1 subantennal, 1-1 epicranial. Genae with basal depression, distal convexity granular. Median triangular frontal field smooth, supra-antennal epicranial surface coarsely granular, posteriormost surface smooth and polished. Median epicranial sulcus deep and prominent. Interantennal space
|
||
<quantity id="4CC95BABFFBF821CFA90FA14FA6382C8" box="[1330,1432,1421,1445]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="54" unit="mm" value="0.6">0.60 mm</quantity>
|
||
. Antennae robust, ca
|
||
<quantity id="4CC95BABFFBF821CFEDCFA2DFE4482A1" box="[382,447,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="54" unit="mm" value="9.0">9 mm</quantity>
|
||
long, antennomeres set with sparse long setae. 5th and 6th without defined sensory pit on outer distal margin, but that area with a field of small possibly sensory setae; 7th without sensory modification; four apical sensory cones.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBF821CFF65F9BAFE078132" blockId="2.[151,1437,1133,2027]" pageId="2" pageNumber="54">
|
||
Collum
|
||
<quantity id="4CC95BABFFBF821CFE86F9BAFE878156" box="[292,380,1571,1595]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="54" unit="mm" value="5.2">5.2 mm</quantity>
|
||
wide, medially strongly arched with two paramedian smooth convexities, paranota abruptly narrowed laterad, subtriangular.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBF821CFF65F9F5FD988182" blockId="2.[151,1437,1133,2027]" pageId="2" pageNumber="54">Prozona ornamented with a network of very fine, short, transversely arranged ridges, imparting a somewhat aspirate texture. Stricture deep around segments but without defining edges. Metaterga essentially smooth, without transverse series of tubercles except collum and segments 2 and 3 with a row of indistinct tubercles along caudal margin; limbus broad, edge not laciniate.</paragraph>
|
||
<paragraph id="8B8EF64EFFBF821CFF65F962FB518086" blockId="2.[151,1437,1133,2027]" pageId="2" pageNumber="54">All paranota posterior to collum elevated above level of middorsum (Fig. 6), most prominently on anterior and posteriormost segments, those of segments 2 and 3 secondarily decurved horizontally distally. Paranota widely separated, except on segments 2–4 increasingly deflected posteriad, the anterior corners diminished into part of a continuous arc, caudal edges convex. Anterior edges abruptly reflexed nearly vertically, set off by a depression from paranotal surface, the actual white edge thickened and coarsely convoluted (Fig. 2), resembling a tightly twisted cord; posterior edges less strongly reflexed but notably thickened and provided with 4–6 acute dentations (Fig. 3). Ozopores removed from lateral edge by about 2 or 3 times diameter of the peritreme (Fig. 2).</paragraph>
|
||
<paragraph id="8B8EF64EFFBE821DFF35F881FF3580DC" blockId="3.[151,1437,1814,1969]" pageId="3" pageNumber="55">
|
||
<emphasis id="B9452A5CFFBE821DFF35F881FEC18043" bold="true" box="[151,314,1816,1839]" pageId="3" pageNumber="55">FIGURES 2–6.</emphasis>
|
||
<emphasis id="B9452A5CFFBE821DFEE2F880FD998042" box="[320,610,1817,1839]" italics="true" pageId="3" pageNumber="55">
|
||
<taxonomicName id="4C318DCDFFBE821DFEE2F880FDA58042" box="[320,606,1817,1839]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="bryophorus" status="sp. nov.">Psammodesmus bryophorus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B9452A5CFFBE821DFDC0F880FD608043" bold="true" box="[610,667,1817,1838]" pageId="3" pageNumber="55">
|
||
<taxonomicNameLabel id="A2769727FFBE821DFDC0F880FD608043" box="[610,667,1817,1838]" pageId="3" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
2. Left paranotum of 13th segment, with part of the anterior margin enlarged at upper left. 3. Left paranotum of 14th segment. 4. Left paranotum of 18th segment. 5. Epiproct. 6. Left side of midbody segment, posterior view, showing elevated paranotum. Figs.7–10.
|
||
<taxonomicName id="4C318DCDFFBE821DFCB9F8C2FB6E801C" authority="Silvestri." authorityName="Silvestri." box="[795,1173,1883,1905]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBE821DFCB9F8C2FBC9801C" box="[795,1074,1883,1905]" italics="true" pageId="3" pageNumber="55">Psammodesmus fasciolatus</emphasis>
|
||
Silvestri.
|
||
</taxonomicName>
|
||
7. Left paranotum of 13th segment. 8. Left paranotuk of 14th segment. 9. Left paranotum of 18th segment. 10. Left side of midbody segment, posterior view.
|
||
</paragraph>
|
||
<caption id="DF4EA6C6FFB9821AFF35F903FB3681BD" httpUri="https://zenodo.org/record/206006/files/figure.png" pageId="4" pageNumber="56" targetBox="[218,1356,188,1620]" targetPageId="4">
|
||
<paragraph id="8B8EF64EFFB9821AFF35F903FB3681BD" blockId="4.[151,1437,1690,1744]" pageId="4" pageNumber="56">
|
||
<emphasis id="B9452A5CFFB9821AFF35F903FEA881DD" bold="true" box="[151,339,1690,1713]" pageId="4" pageNumber="56">FIGURES 11–12.</emphasis>
|
||
<taxonomicName id="4C318DCDFFB9821AFEF9F902FD8381DC" box="[347,632,1691,1713]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="bryophorus" status="sp. nov.">
|
||
<emphasis id="B9452A5CFFB9821AFEF9F902FD8381DC" box="[347,632,1691,1713]" italics="true" pageId="4" pageNumber="56">Psammodesmus bryophorus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9452A5CFFB9821AFD25F902FD3981DD" bold="true" box="[647,706,1691,1712]" pageId="4" pageNumber="56">
|
||
<taxonomicNameLabel id="A2769727FFB9821AFD25F902FD3981DD" box="[647,706,1691,1712]" pageId="4" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
11. Gonopods in situ, view view. 12. Left gonopod, mesal aspect. Fig. 13.
|
||
<taxonomicName id="4C318DCDFFB9821AFF1FF923FD8981BD" authority="Silvestri. Right" authorityName="Silvestri. Right" box="[189,626,1722,1744]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFB9821AFF1FF923FE2881BD" box="[189,467,1722,1744]" italics="true" pageId="4" pageNumber="56">Psammodesmus fasciolatus</emphasis>
|
||
Silvestri. Right
|
||
</taxonomicName>
|
||
gonopod of holotype, mesal aspect (left gonopod missing).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8EF64EFFB9821AFF65F96EFC2080F2" blockId="4.[151,1436,1783,2024]" pageId="4" pageNumber="56">Epiproct (Fig. 5). Paraprocts generally flat, ornamented with about six vertical small ridges each bearing a single row of tiny tubercles; basal third of surface abruptly angled outward to lie in a plane with the hypoproct; latter subtriangular, with two prominent digitiform paramedian processes. Legs attached to strongly elevated, densely setose podosterna, condyles not produced as subcoxal spines. Sides of metazona nearly smooth, at most with sparse indistinct tubercles. Legs uniformly invested with sparse, long, erect setae.</paragraph>
|
||
<paragraph id="8B8EF64EFFB9821BFF65F832FDA7879A" blockId="4.[151,1436,1783,2024]" lastBlockId="5.[151,1436,151,535]" lastPageId="5" lastPageNumber="57" pageId="4" pageNumber="56">
|
||
Gonopod aperture (
|
||
<figureCitation id="130AEACBFFB9821AFE01F832FE0880AE" box="[419,499,1963,1987]" captionStart="FIGURES 11 – 12" captionStartId="4.[151,264,1690,1713]" captionTargetBox="[218,1356,188,1620]" captionTargetId="figure@4.[151,1436,162,1669]" captionTargetPageId="4" captionText="FIGURES 11 – 12. Psammodesmus bryophorus, n. sp. 11. Gonopods in situ, view view. 12. Left gonopod, mesal aspect. Fig. 13. Psammodesmus fasciolatus Silvestri. Right gonopod of holotype, mesal aspect (left gonopod missing)." httpUri="https://zenodo.org/record/206006/files/figure.png" pageId="4" pageNumber="56">Fig. 11</figureCitation>
|
||
) transversely oval, not intruding onto stricture, lateral ends and posterior edge elevated. Gonopods stout, contiguous in situ (
|
||
<figureCitation id="130AEACBFFB9821AFDD7F856FD3D808A" box="[629,710,1999,2023]" captionStart="FIGURES 11 – 12" captionStartId="4.[151,264,1690,1713]" captionTargetBox="[218,1356,188,1620]" captionTargetId="figure@4.[151,1436,162,1669]" captionTargetPageId="4" captionText="FIGURES 11 – 12. Psammodesmus bryophorus, n. sp. 11. Gonopods in situ, view view. 12. Left gonopod, mesal aspect. Fig. 13. Psammodesmus fasciolatus Silvestri. Right gonopod of holotype, mesal aspect (left gonopod missing)." httpUri="https://zenodo.org/record/206006/files/figure.png" pageId="4" pageNumber="56">Fig. 11</figureCitation>
|
||
), extended forward to sternum of segment 6, coxa black on anterior surface, white on ventral, with a single long, simple seta on the dorsal surface; telopodite nearly black except for clear testaceous apical lamina. Shape of apical lamina (
|
||
<figureCitation id="130AEACBFFB8821BFC9BFF25FC7787B9" box="[825,908,188,212]" captionStart="FIGURES 11 – 12" captionStartId="4.[151,264,1690,1713]" captionTargetBox="[218,1356,188,1620]" captionTargetId="figure@4.[151,1436,162,1669]" captionTargetPageId="4" captionText="FIGURES 11 – 12. Psammodesmus bryophorus, n. sp. 11. Gonopods in situ, view view. 12. Left gonopod, mesal aspect. Fig. 13. Psammodesmus fasciolatus Silvestri. Right gonopod of holotype, mesal aspect (left gonopod missing)." httpUri="https://zenodo.org/record/206006/files/figure.png" pageId="5" pageNumber="57">Fig. 12</figureCitation>
|
||
) similar to that of
|
||
<taxonomicName id="4C318DCDFFB8821BFBC7FF24FB0787B9" box="[1125,1276,188,212]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="57" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFB8821BFBC7FF24FB0787B9" box="[1125,1276,188,212]" italics="true" pageId="5" pageNumber="57">P. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 13), but slightly smaller as seen in mesal aspect..
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFB8821BFF65FE9DFEC8857A" blockId="5.[151,1436,151,535]" pageId="5" pageNumber="57">
|
||
Female
|
||
<typeStatus id="548A48ECFFB8821BFE80FE9CFE7C8670" box="[290,391,261,285]" pageId="5" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
: Length ca.
|
||
<quantity id="4CC95BABFFB8821BFDB6FE9DFD9C8671" box="[532,615,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="57" unit="mm" value="64.0">64 mm</quantity>
|
||
, maximum width (near segment 14),
|
||
<quantity id="4CC95BABFFB8821BFBB4FE9DFB8B8671" box="[1046,1136,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="5" pageNumber="57" unit="mm" value="9.2">9.2 mm</quantity>
|
||
, but body almost parallelsided over most of length, W/L ratio ca. 15%. Agreeing very closely with male in color and peripheral structure, but differing in that the paranota are less elevated, and metazona with a series of evident small tubercles along posterior margin. Cyphopods (Figs. 14, 15) reduced in size and sclerotization, largely concealed in intersegmental membrane and visible only with dissection. Receptacle prolonged and narrowed mesad, the surface strio-annulate, merging into thin clear membrane proximad, subglobosely enlarged laterad, with smooth surface, valves much reduced in size and indistinct, apparently contained within receptacle; operculum large, flattened, entire cyphopod without setae.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFB8821BFF35F8F2FBD480D2" blockId="5.[151,1436,1897,1983]" pageId="5" pageNumber="57">
|
||
<emphasis id="B9452A5CFFB8821BFF35F8F2FEAF80ED" bold="true" box="[151,340,1899,1922]" pageId="5" pageNumber="57">FIGURES 14–15.</emphasis>
|
||
<emphasis id="B9452A5CFFB8821BFEF8F8F2FD8580EC" box="[346,638,1899,1921]" italics="true" pageId="5" pageNumber="57">
|
||
<taxonomicName id="4C318DCDFFB8821BFEF8F8F2FD8180EC" box="[346,634,1899,1921]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="57" phylum="Arthropoda" rank="species" species="bryophorus" status="sp. nov.">Psammodesmus bryophorus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B9452A5CFFB8821BFD26F8F5FD4580EC" bold="true" box="[644,702,1900,1921]" pageId="5" pageNumber="57">
|
||
<taxonomicNameLabel id="A2769727FFB8821BFD26F8F5FD4580EC" box="[644,702,1900,1921]" pageId="5" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
14. Left side of sternum of 2nd pair of legs of female
|
||
<typeStatus id="548A48ECFFB8821BFB7CF8F5FAC380EC" box="[1246,1336,1900,1921]" pageId="5" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
, posterior aspect, showing cyphopod
|
||
<emphasis id="B9452A5CFFB8821BFE08F812FE1480CD" box="[426,495,1931,1952]" italics="true" pageId="5" pageNumber="57">in situ.</emphasis>
|
||
15. Left cyphopod dissected free from membrane, posterolateral aspect. M: retractor muscle from tracheal apodeme to receptacle. O: operculum; OV: oviduct; R: receptacle; V: valves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4EA6C6FFBB8218FF35FCF4FDEF84CF" httpUri="https://zenodo.org/record/206007/files/figure.png" pageId="6" pageNumber="58" targetBox="[157,1424,202,845]" targetPageId="6">
|
||
<paragraph id="8B8EF64EFFBB8218FF35FCF4FDEF84CF" blockId="6.[151,1436,877,930]" pageId="6" pageNumber="58">
|
||
<emphasis id="B9452A5CFFBB8218FF35FCF4FEDB84EE" bold="true" box="[151,288,877,900]" pageId="6" pageNumber="58">FIGURE 16.</emphasis>
|
||
Southwestern Colombia, showing location of type locality (on left), boxed area in south central Nariño Department shown on right for closer detail.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32BA5C5FFBB8218FF65FC57FEC581FE" pageId="6" pageNumber="58" type="discussion">
|
||
<paragraph id="8B8EF64EFFBB8218FF65FC57FAE28246" blockId="6.[151,1437,974,1539]" pageId="6" pageNumber="58">
|
||
<emphasis id="B9452A5CFFBB8218FF65FC57FEB3848A" bold="true" box="[199,328,974,999]" pageId="6" pageNumber="58">Comments</emphasis>
|
||
. The
|
||
<typeStatus id="548A48ECFFBB8218FE29FC49FE408485" box="[395,443,976,1000]" pageId="6" pageNumber="58">type</typeStatus>
|
||
locality, Altaquer, is shown on large scale maps (e.g.,
|
||
<collectingCountry id="F326B6DEFFBB8218FB8BFC56FB6C848A" box="[1065,1175,975,999]" name="Colombia" pageId="6" pageNumber="58">Colombia</collectingCountry>
|
||
1950), also on Google Earth satellite imagery. The reserve is located in southwestern
|
||
<collectingCountry id="F326B6DEFFBB8218FCCBFC6AFC218366" box="[873,986,1011,1035]" name="Colombia" pageId="6" pageNumber="58">Colombia</collectingCountry>
|
||
, in a transitional Andean-Pacific forest between
|
||
<quantity id="4CC95BABFFBB8218FF5FFB8EFEB28342" box="[253,329,1047,1071]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="58" unit="m" value="1100.0">1100m</quantity>
|
||
and
|
||
<quantity id="4CC95BABFFBB8218FE21FB8EFE278342" box="[387,476,1047,1071]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="58" unit="m" value="1500.0">1500 m</quantity>
|
||
, with annual rainfall of
|
||
<quantity id="4CC95BABFFBB8218FD4DFB8EFCA78342" box="[751,860,1047,1071]" metricMagnitude="0" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="58" unit="mm" value="8500.0">8500 mm</quantity>
|
||
and average temperature 19° C. It is in the Choco phytogeographic region (
|
||
<bibRefCitation id="EFA08BBFFFBB8218FE10FBA2FDB5833E" author="Rangel" box="[434,590,1083,1107]" pageId="6" pageNumber="58" refString="Rangel, O. (2004) Colombia diversidad biotica IV. El chocho biogeografico / Costa Pacifica. Universidad Nacional de Colombia, 41, 977 - 985." type="journal article" year="2004">Rangel, 2004</bibRefCitation>
|
||
), the most outstanding characteristic of which, according
|
||
<bibRefCitation id="EFA08BBFFFBB8218FB78FBA2FA87833E" author="Gentry" box="[1242,1404,1083,1107]" pageId="6" pageNumber="58" refString="Gentry, A. H. (1982) Phytogeographic patterns as evidence for Choco refuge. In G. T. Prance (ed.), Biological diversification in the tropics. Columbia Univ. Press, New York, 112 - 136." type="book chapter" year="1982">Gentry (1982)</bibRefCitation>
|
||
, is the extremely high average annual precipitation, reaching
|
||
<quantity id="4CC95BABFFBB8218FC92FBC6FC51831A" box="[816,938,1119,1143]" metricMagnitude="1" metricUnit="m" metricValue="1.177" pageId="6" pageNumber="58" unit="mm" value="11770.0">11770 mm</quantity>
|
||
at Tutunendo in central Choco department. Choco is the only place in the neotropics in which pluvial forest (Tropical rain forest) occurs, a region with high endemicity and the most biodiverse ecosystems on earth (
|
||
<bibRefCitation id="EFA08BBFFFBB8218FCB5FB3EFC3683D2" author="Lourenco" box="[791,973,1191,1215]" pageId="6" pageNumber="58" refString="Lourenco, W (1994) Diversity and endemism in tropical versus temperate scorpion communities. Biogeographica, 70 (3), 155 - 160." type="journal article" year="1994">Lourenço, 1994</bibRefCitation>
|
||
). This reserve is also characterized by the forest sharing, according to
|
||
<bibRefCitation id="EFA08BBFFFBB8218FE79FB52FCE0838E" box="[475,795,1227,1251]" pageId="6" pageNumber="58" refString="Franco-Roselli, P., Betancur, J. & Fernandez-Alonso, J. (1997) Diversidad floristica en dos bosques subandinos del sur de Colombia. Caldasia, 19 (1 - 2), 205 - 234." type="journal article">Franco-Roselli et al. (1997)</bibRefCitation>
|
||
, many of its features, such as floristic composition and stratification, with the rain forest (<
|
||
<quantity id="4CC95BABFFBB8218FD90FB76FD79826A" box="[562,642,1263,1287]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="58" unit="m" value="1000.0">1000m</quantity>
|
||
) and montane forest (> 1800). It also shares many species with sub- Andean and Pacific forests, so the Ñambí Reserve is considered an ecotone between these two regions.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBB8218FF65FAAEFCDB82FA" blockId="6.[151,1437,974,1539]" pageId="6" pageNumber="58">
|
||
Field notes specify collections made by hand; the millipeds being active both diurnally and nocturnally, on the surface and ascending trees.Most individuals were active at night, on mossy trunks about
|
||
<quantity id="4CC95BABFFBB8218FB2FFAC2FB19821E" box="[1165,1250,1371,1395]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="58" unit="m" value="1.0">1 meter</quantity>
|
||
height, some on the petioles of living leaves, others in litter on the ground.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBB8218FF65FA3AFC77816E" blockId="6.[151,1437,974,1539]" pageId="6" pageNumber="58">
|
||
Eleven of the 18 individuals carried more than 400 individuals of bryophytes, chiefly on the dorsal side of the paranota. Families of bryophytes found include
|
||
<taxonomicName id="4C318DCDFFBB8218FD0CFA5EFCBD82B2" box="[686,838,1479,1503]" class="Jungermanniopsida" family="Lejeuneaceae" kingdom="Plantae" order="Porellales" pageId="6" pageNumber="58" phylum="Bryophyta" rank="family">Lejeuneaceae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C318DCDFFBB8218FCDEFA5EFBE582B2" box="[892,1054,1479,1503]" class="Bryopsida" family="Pilotrichaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hookeriales" pageId="6" pageNumber="58" phylum="Bryophyta" rank="family">Pilotrichaceae</taxonomicName>
|
||
, which principally grow on leaves of shrubs and trees, and
|
||
<taxonomicName id="4C318DCDFFBB8218FE05FA72FDB1816E" box="[423,586,1515,1539]" class="Bryopsida" family="Fissidentaceae" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="58" phylum="Bryophyta" rank="family">Fissidentaceae</taxonomicName>
|
||
species associated with soil.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBB8218FF35F9CFFEC581FE" blockId="6.[151,568,1622,1683]" pageId="6" pageNumber="58">
|
||
<taxonomicName id="4C318DCDFFBB8218FF35F9CFFDC38102" authority="Silvestri" authorityName="Silvestri" box="[151,568,1622,1647]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBB8218FF35F9CFFE288102" bold="true" box="[151,467,1622,1647]" italics="true" pageId="6" pageNumber="58">Psammodesmus fasciolatus</emphasis>
|
||
Silvestri
|
||
</taxonomicName>
|
||
Figs. 7–10, 13.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32BA5C5FFBB8218FF35F95AFCC08058" pageId="6" pageNumber="58" type="reference_group">
|
||
<paragraph id="8B8EF64EFFBB8218FF35F95AFEEF819A" blockId="6.[151,1436,1730,1845]" pageId="6" pageNumber="58">
|
||
<treatmentCitationGroup id="AB21D160FFBB8218FF35F95AFEEF819A" pageId="6" pageNumber="58">
|
||
<taxonomicName id="4C318DCDFFBB8218FF35F95AFD9A81B5" ID-CoL="4NCG7" authority="Silvestri, 1898: 4" authorityName="Silvestri" authorityPageNumber="4" authorityYear="1898" box="[151,609,1730,1753]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBB8218FF35F95AFE5781B4" box="[151,428,1731,1753]" italics="true" pageId="6" pageNumber="58">Psammodesmus fasciolatus</emphasis>
|
||
<treatmentCitation id="0A90D05FFFBB8218FE10F95BFD9A81B5" author="Silvestri" box="[434,609,1730,1752]" page="4" pageId="6" pageNumber="58" year="1898">
|
||
<bibRefCitation id="EFA08BBFFFBB8218FE10F95BFDB381B5" author="Silvestri" box="[434,584,1730,1752]" pageId="6" pageNumber="58" refString="Silvestri, F. (1898) Viaggio del Dr. Enrico Festa nell'Ecuador e regionali vicine. XI. Diplopodi. Bolletino dei Musei di Zoologia ed Anatomia comparata della Reale Universita di Torino, 13 (324), 1 - 11." type="journal article" year="1898">Silvestri, 1898</bibRefCitation>
|
||
: 4
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, fig. 6. Male
|
||
<typeStatus id="548A48ECFFBB8218FD44F95BFCC481B5" box="[742,831,1730,1752]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
(MCSNG) from “Foreste del Rio Peripa” [see “Comments” below].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBB8218FF35F899FCC08058" blockId="6.[151,1436,1730,1845]" pageId="6" pageNumber="58">
|
||
<treatmentCitationGroup id="AB21D160FFBB8218FF35F899FCC08058" pageId="6" pageNumber="58">
|
||
<taxonomicName id="4C318DCDFFBB8218FF35F899FD79807B" authority="Brolemann, 1919: 254" authorityName="Brolemann" authorityPageNumber="254" authorityYear="1919" box="[151,642,1792,1814]" class="Diplopoda" family="Platyrhacidae" genus="Platyrrhacus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBB8218FF35F899FE75807B" box="[151,398,1792,1814]" italics="true" pageId="6" pageNumber="58">Platyrrhacus fasciolatus</emphasis>
|
||
:
|
||
<treatmentCitation id="0A90D05FFFBB8218FE3EF899FD79807B" author="Brolemann" box="[412,642,1792,1814]" page="254" pageId="6" pageNumber="58" year="1919">
|
||
<bibRefCitation id="EFA08BBFFFBB8218FE3EF899FDB4807B" author="Brolemann" box="[412,591,1792,1814]" pageId="6" pageNumber="58" refString="Brolemann, H. - W. (1919) Myriapodes, in: Mission du Service geographique de l'Armee pour la Mesure d'un Arc de Meridien equatorial en Amerique du Sud 1899 - 1906, 10 (2), 235 - 275." type="journal article" year="1919">Brolemann, 1919</bibRefCitation>
|
||
: 254
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, fig, figs. 11, 21. [Santo
|
||
<collectingCountry id="F326B6DEFFBB8218FCD9F899FC21807B" box="[891,986,1792,1814]" name="Dominican Republic" pageId="6" pageNumber="58">Domingo</collectingCountry>
|
||
de los Colorado,
|
||
<collectingCountry id="F326B6DEFFBB8218FB2FF899FB1C807B" box="[1165,1255,1792,1814]" name="Ecuador" pageId="6" pageNumber="58">Ecuador</collectingCountry>
|
||
].
|
||
<taxonomicName id="4C318DCDFFBB8218FF35F886FD208059" authority="Attems, 1938: 238" authorityName="Attems" authorityPageNumber="238" authorityYear="1938" box="[151,731,1823,1845]" class="Diplopoda" family="Platyrhacidae" genus="Platyrhacus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="fasciolatus" subGenus="Tirodesmus">
|
||
<emphasis id="B9452A5CFFBB8218FF35F886FDEB8058" box="[151,528,1823,1845]" italics="true" pageId="6" pageNumber="58">Platyrhacus (Tirodesmus) fasciolatus</emphasis>
|
||
:
|
||
<treatmentCitation id="0A90D05FFFBB8218FDBFF886FD208059" author="Attems" box="[541,731,1823,1844]" page="238" pageId="6" pageNumber="58" year="1938">
|
||
<bibRefCitation id="EFA08BBFFFBB8218FDBFF886FD528059" author="Attems" box="[541,681,1823,1844]" pageId="6" pageNumber="58" refString="Attems, C. (1938) Polydesmoidea II. Fam. Leptodesmidae, Platyrrhacidae, Oxydesmidae, Gomphodesmidae, in Das Tierreich, 69, 1 - 487." type="journal article" year="1938">Attems, 1938</bibRefCitation>
|
||
: 238
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, fig. 263.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32BA5C5FFBB8219FF35F8FBFC8F8412" lastPageId="7" lastPageNumber="59" pageId="6" pageNumber="58" type="materials_examined">
|
||
<paragraph id="8B8EF64EFFBB8218FF35F8FBFC7180F0" blockId="6.[151,1436,1889,2022]" pageId="6" pageNumber="58">
|
||
<emphasis id="B9452A5CFFBB8218FF35F8FBFEFF8017" bold="true" box="[151,260,1890,1914]" pageId="6" pageNumber="58">
|
||
<typeStatus id="548A48ECFFBB8218FF35F8FBFEFF8017" box="[151,260,1890,1914]" pageId="6" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
: Male, fragmented, length ca.
|
||
<quantity id="4CC95BABFFBB8218FDC6F8F8FD438014" box="[612,696,1889,1913]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="58" unit="mm" value="55.0">55 mm</quantity>
|
||
. Segment 2,
|
||
<quantity id="4CC95BABFFBB8218FCECF8F8FC5C8014" box="[846,935,1889,1913]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="58" unit="mm" value="8.5">8.5 mm</quantity>
|
||
, segment 6,
|
||
<quantity id="4CC95BABFFBB8218FB9BF8F8FB688014" box="[1081,1171,1889,1913]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="6" pageNumber="58" unit="mm" value="9.2">9.2 mm</quantity>
|
||
, segments 10 – 14,
|
||
<quantity id="4CC95BABFFBB8218FADBF8F8FF3E80F0" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="6" pageNumber="58" unit="mm" value="9.4">9.4 mm</quantity>
|
||
, segment 16, 9.0 mm, segment 18,
|
||
<quantity id="4CC95BABFFBB8218FDF3F81CFD5180F0" box="[593,682,1925,1949]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="58" unit="mm" value="6.6">6.6 mm</quantity>
|
||
. W/L ratio ca. 17%.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBB8218FF65F830FAE88088" blockId="6.[151,1436,1889,2022]" pageId="6" pageNumber="58">After a century in alcohol, the color pattern is still evident although obviously bleached: dorsum light brown with light paramedian stripes, oblique and widely spaced on metaterga, closer and parallel on prozona.</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FF0EFCE0879A" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
Epicranial furrow deep and prominent, antennal sockets with high, polished edges medially and dorsally; dorsal half of genae coarsely granular, forming a cristate edge subtending antennal socket; labroclypeal median area smooth. Antennae ca.
|
||
<quantity id="4CC95BABFFBA8219FE32FF79FE28879A" box="[400,467,224,247]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="59" unit="mm" value="7.0">7 mm</quantity>
|
||
, interantennal space
|
||
<quantity id="4CC95BABFFBA8219FD1FFF46FCED879A" box="[701,790,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="59" unit="mm" value="0.5">0.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FE9DFD1A86BD" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">Collum appearing strongly elevated medially owing to abrupt depression of its paranota, middorsal area flat to slightly concave, the entire surface invested in flattened oval tubercles, anterior margin with a row of 6-6 enlarged oval polished tubercles set off by a transverse depression. Subsequent metaterga more or less uniformly ornamented with irregular flattened tubercles, smallest near stricture, becoming much larger posteriad. A series of 12 to 14 small tubercles across posterior margin of metaterga, another series of much smaller tubercles across midlength, both series most distinct on posteriormost segments.</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FE44FAC985E9" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
Paranota of segments 2–4 directed anteriad, laterally distinctly decurved, paranota of segments 4 to about 14 transverse, nearly horizontal and set high on sides, the middorsum only slightly convex (Fig. 10). Outlines of representative paranota shown in Figs. 7–9. Anterior edges not notably elevated and modified as in
|
||
<emphasis id="B9452A5CFFBA8219FB56FDBCFA678551" box="[1268,1436,548,572]" italics="true" pageId="7" pageNumber="59">
|
||
<taxonomicName id="4C318DCDFFBA8219FB56FDBCFA628551" box="[1268,1433,548,572]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="bryophorus">P. bryophorus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Epiproct similar to that of
|
||
<taxonomicName id="4C318DCDFFBA8219FE1FFDD1FDC5850D" box="[445,574,584,608]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="bryophorus">
|
||
<emphasis id="B9452A5CFFBA8219FE1FFDD1FDC5850D" box="[445,574,584,608]" italics="true" pageId="7" pageNumber="59">bryophorus</emphasis>
|
||
</taxonomicName>
|
||
. Paraprocts finely granular, the discal and marginal setiferous tubercles notably large and polished. Paramedian tubercles of hypoproct prominent, about twice as long as basal width.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FD16FF188479" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">Sterna elevated, granular, spiniform at coxal condyles on a few midbody segments only. Sides of metazona uniformly and evenly granular. Anterior stigma located on anterior face of procoxal condyle, extending down into stricture; posterior stigma located somewhat anteriad to metacoxal condyle, nearly straight. Legs stout, sparsely setose.</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FC86FC8F8412" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
Gonopod aperture small, oval, similar to that of
|
||
<emphasis id="B9452A5CFFBA8219FD7FFCB9FC9E8455" box="[733,869,800,824]" italics="true" pageId="7" pageNumber="59">
|
||
<taxonomicName id="4C318DCDFFBA8219FD7FFCB9FC998455" box="[733,866,800,824]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="bryophorus">bryophorus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Gonopods straight, massive, setose on ventral side up to level of solenomere, apical lamina appearing somewhat narrower than in that species (cf.
|
||
<figureCitation id="130AEACBFFBA8219FB42FCDDFAC78431" box="[1248,1340,836,860]" captionStart="FIGURES 11 – 12" captionStartId="4.[151,264,1690,1713]" captionTargetBox="[218,1356,188,1620]" captionTargetId="figure@4.[151,1436,162,1669]" captionTargetPageId="4" captionText="FIGURES 11 – 12. Psammodesmus bryophorus, n. sp. 11. Gonopods in situ, view view. 12. Left gonopod, mesal aspect. Fig. 13. Psammodesmus fasciolatus Silvestri. Right gonopod of holotype, mesal aspect (left gonopod missing)." httpUri="https://zenodo.org/record/206006/files/figure.png" pageId="7" pageNumber="59">Figs. 12</figureCitation>
|
||
and 13) owing to a different positional orientation (see comments below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32BA5C5FFBA8219FF65FC12FC8E816A" pageId="7" pageNumber="59" type="discussion">
|
||
<paragraph id="8B8EF64EFFBA8219FF65FC12FE3E8359" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
<emphasis id="B9452A5CFFBA8219FF65FC12FEB484C9" bold="true" box="[199,335,907,932]" pageId="7" pageNumber="59">Comments.</emphasis>
|
||
“Rio Peripa” is taken to be the stream forming the boundary between Manabi and Guaymas provinces, ca.
|
||
<quantity id="4CC95BABFFBA8219FEA6FC36FEB084AA" box="[260,331,943,967]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="59" unit="km" value="30.0">30 km</quantity>
|
||
northwest of Quevedo, at coordinates
|
||
<geoCoordinate id="EE059089FFBA8219FD5BFC36FC7F84AA" box="[761,900,943,967]" direction="west" orientation="longitude" pageId="7" pageNumber="59" precision="1" value="-79.71667">79.71667 W</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="EE059089FFBA8219FC30FC36FC0584AA" box="[914,1022,943,967]" direction="south" orientation="latitude" pageId="7" pageNumber="59" precision="1" value="-0.88333">0.88333S</geoCoordinate>
|
||
. Festa collected there during December of 1897; his localities may now be inundated. by an impoundment.. Information on the Internet associating the river with Pichincha do not refer to the Ecuadorian province by that name, but to a small settlement on Rt. 40 about
|
||
<quantity id="4CC95BABFFBA8219FF35FB85FF258359" box="[151,222,1052,1076]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="59" unit="km" value="37.0">37 km</quantity>
|
||
south of Rio Peripa.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FBD9FAAA8385" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
A specimen in the British Museum (1903.7.2.50) labeled “Rio Durango,
|
||
<collectingCountry id="F326B6DEFFBA8219FC5FFBD9FBA68335" box="[1021,1117,1088,1112]" name="Ecuador" pageId="7" pageNumber="59">Ecuador</collectingCountry>
|
||
” was examined in 1960 and identified as
|
||
<taxonomicName id="4C318DCDFFBA8219FE88FBFCFE448311" box="[298,447,1124,1148]" class="Diplopoda" family="Platyrhacidae" genus="Platyrhacus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBA8219FE88FBFCFE448311" box="[298,447,1124,1148]" italics="true" pageId="7" pageNumber="59">P. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
on the basis of Silvestri’s description. Although the gonopod structure closely matches that of the
|
||
<typeStatus id="548A48ECFFBA8219FEB2FB10FEBB83CC" box="[272,320,1161,1185]" pageId="7" pageNumber="59">type</typeStatus>
|
||
specimen, no notes were made about peripheral structure, so this specimen must be re-examined to establish its possible identity with
|
||
<taxonomicName id="4C318DCDFFBA8219FDBEFB34FD4383A9" box="[540,696,1196,1220]" class="Diplopoda" family="Platyrhacidae" genus="Psammodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="bryophorus">
|
||
<emphasis id="B9452A5CFFBA8219FDBEFB34FD4383A9" box="[540,696,1196,1220]" italics="true" pageId="7" pageNumber="59">P. bryophorus</emphasis>
|
||
</taxonomicName>
|
||
, especially since the locality presumably is the Rio Durango near San Lorenzo in Esmeraldas Province, only about
|
||
<quantity id="4CC95BABFFBA8219FD62FB49FCEE8385" box="[704,789,1232,1256]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="59" unit="km" value="100.0">100 km</quantity>
|
||
west and south of the Reserva Natural Río Ńembí.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FB6DFC7B8242" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
Brolemann’s 1919 detailed description of his material from Santo
|
||
<collectingCountry id="F326B6DEFFBA8219FC04FB6DFBEB8261" box="[934,1040,1268,1292]" name="Dominican Republic" pageId="7" pageNumber="59">Domingo</collectingCountry>
|
||
de los Colorados is entirely consistent with the
|
||
<typeStatus id="548A48ECFFBA8219FE88FA8EFE778242" box="[298,396,1303,1327]" pageId="7" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C318DCDFFBA8219FE13FA81FDD1825D" box="[433,554,1304,1328]" class="Diplopoda" family="Platyrhacidae" genus="Platyrhacus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBA8219FE13FA81FDD1825D" box="[433,554,1304,1328]" italics="true" pageId="7" pageNumber="59">fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
, confirming his identification.
|
||
</paragraph>
|
||
<paragraph id="8B8EF64EFFBA8219FF65FAA5FC8E816A" blockId="7.[151,1436,151,1543]" pageId="7" pageNumber="59">
|
||
Observations of the gonopods of the two species under discussion demonstrated that the least difference in position of the appendage resulted in substantial alteration in the appearance of the apical laminla in particular. In
|
||
<taxonomicName id="4C318DCDFFBA8219FF35FA1CFED082F1" box="[151,299,1412,1436]" class="Diplopoda" family="Platyrhacidae" genus="Platyrhacus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="59" phylum="Arthropoda" rank="species" species="fasciolatus">
|
||
<emphasis id="B9452A5CFFBA8219FF35FA1CFED082F1" box="[151,299,1412,1436]" italics="true" pageId="7" pageNumber="59">P. fasciolatus</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 13) this blade is so positioned that in mesal aspect more of the thin edge is visible than in
|
||
<emphasis id="B9452A5CFFBA8219FAF0FA1CFF0882AD" italics="true" pageId="7" pageNumber="59">P. b r y - ophorus</emphasis>
|
||
and therefore seems smaller and more slender. Perhaps a precaution to be considered in future studies would be to illustrate the gonopod (in addition to the standard mesal aspect) in a position that shows the maximum surface/shape of the lamina whatever its actual position might be.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |