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<document ID-DOI="10.3853/j.2201-4349.74.2022.1775" ID-ISSN="2201-4349" ID-Zenodo-Dep="7177922" approvalRequired="7" approvalRequired_for_textStreams="5" approvalRequired_for_treatments="2" checkinTime="1665305657841" checkinUser="felipe" docAuthor="Billingham, Zacariah D. &amp; Theischinger, Gunther" docDate="2022" docId="0392DE4AFC0CFFF0FF050A31FE8BAD0D" docLanguage="en" docName="RecAustMus.74.1.1940.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (1)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.1:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="1" docTitle="Dicranomyia (Dicranomyia) globosa Billingham &amp; Theischinger 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFABA632FC01FFFDFF990B35FFFEAA2C" masterDocTitle="New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia" masterLastPageNumber="40" masterPageNumber="19" pageNumber="32" updateTime="1665324244475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Billingham, Zacariah D.</mods:namePart>
<mods:affiliation>Water Sciences, Natural Resources and Heritage, GHD, 180 Lonsdale Street, Melbourne VIC 3000, Australia &amp; PhD candidate, Department of Ecology, Environment and Evolution, La Trobe University, Bundoora VIC 3086, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Theischinger, Gunther</mods:namePart>
<mods:affiliation>Research Associate, Australian Museum Research Institute, Australian Museum, 6 College Street, Sydney NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="series">
<mods:title>Rec. Aust. Mus.</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2022-03-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>74</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>40</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.74.2022.1775</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">7177922</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7175140" ID-Zenodo-Dep="7175140" LSID="urn:lsid:plazi:treatment:0392DE4AFC0CFFF0FF050A31FE8BAD0D" httpUri="http://treatment.plazi.org/id/0392DE4AFC0CFFF0FF050A31FE8BAD0D" lastPageNumber="32" pageId="13" pageNumber="32">
<subSubSection pageId="13" pageNumber="32" type="nomenclature">
<paragraph blockId="13.[156,713,260,287]" box="[156,713,260,287]" pageId="13" pageNumber="32">
<heading bold="true" box="[156,713,260,287]" centered="true" fontSize="12" level="2" pageId="13" pageNumber="32" reason="2">
<emphasis bold="true" box="[156,713,260,287]" pageId="13" pageNumber="32">
<taxonomicName authority="Billingham &amp; Theischinger, 2022" authorityName="Billingham &amp; Theischinger" authorityYear="2022" box="[156,609,260,287]" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="globosa" status="sp. nov." subGenus="Dicranomyia">
<emphasis bold="true" box="[156,609,260,287]" italics="true" pageId="13" pageNumber="32">Dicranomyia (Dicranomyia) globosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[617,713,260,286]" pageId="13" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[132,737,309,327]" box="[132,737,309,327]" pageId="13" pageNumber="32">
<uri box="[132,737,309,327]" pageId="13" pageNumber="32">
urn:lsid:zoobank.org:act:
<uuid box="[333,737,309,327]" pageId="13" pageNumber="32">4AC026D2-6A76-463B-AD74-A3DBE6A918DF</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[368,501,356,382]" pageId="13" pageNumber="32" type="description">
<paragraph blockId="13.[368,501,356,382]" box="[368,501,356,382]" pageId="13" pageNumber="32">
<figureCitation box="[368,501,356,382]" captionStart="Figures 2932" captionStartId="14.[166,243,1651,1673]" captionTargetBox="[200,1434,154,1645]" captionTargetId="figure-11@14.[167,1467,151,1651]" captionTargetPageId="14" captionText="Figures 2932. Dicranomyia (Dicranomyia) globosa sp. nov., male. (29) head and thoracic colouration, lateral; (30) wing; (31, 32) hypopygium, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7177952" httpUri="https://zenodo.org/record/7177952/files/figure.png" pageId="13" pageNumber="32">Figs 2932</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="32" type="materials_examined">
<paragraph blockId="13.[121,749,403,516]" pageId="13" pageNumber="32">
<materialsCitation collectingDate="2017-09-25" collectionCode="AM" collectorName="Z. Billingham &amp; G. Theischinger" country="Australia" county="Chase" latitude="-33.663" location="Upper Gledhill Falls" longLatPrecision="72" longitude="151.25" municipality="McCarrs Creek" pageId="13" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis bold="true" box="[121,226,404,428]" pageId="13" pageNumber="32">
<typeStatus box="[121,226,404,428]" pageId="13" pageNumber="32">Holotype</typeStatus>
</emphasis>
<specimenCount box="[236,258,404,427]" pageId="13" pageNumber="32" type="male"></specimenCount>
,
<collectingCountry box="[272,374,403,427]" name="Australia" pageId="13" pageNumber="32">Australia</collectingCountry>
,
<collectingRegion box="[389,591,403,427]" country="Australia" name="New South Wales" pageId="13" pageNumber="32">New South Wales</collectingRegion>
, Ku-Ring-Gai
<collectingCounty box="[121,189,433,457]" pageId="13" pageNumber="32">Chase</collectingCounty>
,
<collectingMunicipality box="[198,361,433,457]" pageId="13" pageNumber="32">McCarrs Creek</collectingMunicipality>
by
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC0CFFF0FF050A31FE8BAD0D:8EE43987FC0CFFF0FE170A84FD99ABE5" box="[398,615,433,457]" country="Australia" county="Chase" latitude="-33.663" longLatPrecision="72" longitude="151.25" municipality="McCarrs Creek" name="Upper Gledhill Falls" pageId="13" pageNumber="32" stateProvince="New South Wales">Upper Gledhill Falls</location>
off
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC0CFFF0FF050A31FE8BAD0D:8EE43987FC0CFFF0FD160A84FF25ABCA" country="Australia" county="Chase" latitude="-33.663" longLatPrecision="72" longitude="151.25" municipality="McCarrs Creek" name="McCarrs Creek Rd" pageId="13" pageNumber="32" stateProvince="New South Wales">
<location LSID="urn:lsid:plazi:treatment:0392DE4AFC0CFFF0FF050A31FE8BAD0D:8EE43987FC0CFFF0FD160A84FF46ABCA" country="Australia" county="Chase" latitude="-33.663" longLatPrecision="72" longitude="151.25" municipality="McCarrs Creek" name="McCarrs Creek" pageId="13" pageNumber="32" stateProvince="New South Wales">McCarrs Creek</location>
Rd
</location>
(
<geoCoordinate box="[231,329,462,486]" degrees="33.663" direction="south" orientation="latitude" pageId="13" pageNumber="32" precision="55" value="-33.663">33.663°S</geoCoordinate>
<geoCoordinate box="[333,449,462,486]" degrees="151.250" direction="east" orientation="longitude" pageId="13" pageNumber="32" precision="55" value="151.25">151.250°E</geoCoordinate>
),
<date box="[465,595,462,486]" pageId="13" pageNumber="32" value="2017-09-25">
<collectingDate box="[465,595,462,486]" pageId="13" pageNumber="32" value="2017-09-25">25 Sep 2017</collectingDate>
</date>
,
<collectorName box="[603,749,462,486]" pageId="13" pageNumber="32">Z. Billingham</collectorName>
&amp;
<collectorName box="[148,321,491,516]" pageId="13" pageNumber="32">G. Theischinger</collectorName>
,
<collectionCode box="[330,373,492,516]" country="Australia" name="Australian Museum" pageId="13" pageNumber="32" type="Museum">AM</collectionCode>
(K.394963).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="32" type="description">
<paragraph blockId="13.[121,750,537,1500]" box="[121,437,537,561]" pageId="13" pageNumber="32">
<emphasis bold="true" box="[121,253,537,561]" pageId="13" pageNumber="32">Description</emphasis>
<specimenCount box="[259,279,537,560]" pageId="13" pageNumber="32" type="male"></specimenCount>
(
<specimenCount box="[297,315,537,561]" pageId="13" pageNumber="32" type="female"></specimenCount>
unknown).
</paragraph>
<paragraph blockId="13.[121,750,537,1500]" pageId="13" pageNumber="32">
<emphasis box="[151,219,566,590]" italics="true" pageId="13" pageNumber="32">Head.</emphasis>
Antenna and palpus greyish brown. Antenna of moderate length, extending back to the prescutal pit, pedicel and flagellomeres 111 distinctly rounded, spherical, ultimate segment more conical.
</paragraph>
<paragraph blockId="13.[121,750,537,1500]" pageId="13" pageNumber="32">
<emphasis box="[151,229,683,707]" italics="true" pageId="13" pageNumber="32">Thorax</emphasis>
(
<figureCitation box="[247,326,683,708]" captionStart="Figures 2932" captionStartId="14.[166,243,1651,1673]" captionTargetBox="[200,1434,154,1645]" captionTargetId="figure-11@14.[167,1467,151,1651]" captionTargetPageId="14" captionText="Figures 2932. Dicranomyia (Dicranomyia) globosa sp. nov., male. (29) head and thoracic colouration, lateral; (30) wing; (31, 32) hypopygium, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7177952" httpUri="https://zenodo.org/record/7177952/files/figure.png" pageId="13" pageNumber="32">Fig 29</figureCitation>
). Thoracic segments dorsally all dark brown, posterior areas of scutal lobes and scutellum a little lighter. Pleurites mostly uniform brown, propleuron and anepisternum a little darker. Coxae brownish, trochanters yellowish brown. Femora, tibiae, and first tarsal segment yellowish brown, darkening to greyish black at the tips, remaining tarsal segments greyish black.
</paragraph>
<paragraph blockId="13.[121,750,537,1500]" pageId="13" pageNumber="32">
<emphasis box="[151,205,889,912]" italics="true" pageId="13" pageNumber="32">Wing</emphasis>
(
<figureCitation box="[217,294,889,913]" captionStart="Figures 2932" captionStartId="14.[166,243,1651,1673]" captionTargetBox="[200,1434,154,1645]" captionTargetId="figure-11@14.[167,1467,151,1651]" captionTargetPageId="14" captionText="Figures 2932. Dicranomyia (Dicranomyia) globosa sp. nov., male. (29) head and thoracic colouration, lateral; (30) wing; (31, 32) hypopygium, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7177952" httpUri="https://zenodo.org/record/7177952/files/figure.png" pageId="13" pageNumber="32">Fig. 30</figureCitation>
): brownish grey, extensively variegated with hyaline patches, veins brownish. Sc extending to nearly ⅓ length of Rs, sc-r some distance before its tip in alignment with the fork of Rs, m-cu situated a little more than half its length before the base of the discal medial cell. Halter with stem deep greyish, knob lighter grey.
</paragraph>
<paragraph blockId="13.[121,750,537,1500]" box="[151,428,1065,1089]" pageId="13" pageNumber="32">
<emphasis box="[151,261,1065,1089]" italics="true" pageId="13" pageNumber="32">Abdomen:</emphasis>
greyish brown.
</paragraph>
<paragraph blockId="13.[121,750,537,1500]" pageId="13" pageNumber="32">
<taxonomicName box="[151,267,1094,1118]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="13" pageNumber="32" phylum="Tracheophyta" rank="genus">
<emphasis box="[151,267,1094,1118]" italics="true" pageId="13" pageNumber="32">Terminalia</emphasis>
</taxonomicName>
(
<figureCitation box="[280,402,1094,1118]" captionStart="Figures 2932" captionStartId="14.[166,243,1651,1673]" captionTargetBox="[200,1434,154,1645]" captionTargetId="figure-11@14.[167,1467,151,1651]" captionTargetPageId="14" captionText="Figures 2932. Dicranomyia (Dicranomyia) globosa sp. nov., male. (29) head and thoracic colouration, lateral; (30) wing; (31, 32) hypopygium, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7177952" httpUri="https://zenodo.org/record/7177952/files/figure.png" pageId="13" pageNumber="32">Figs 31, 32</figureCitation>
). Hypopygium largely yellowish brown, lobes of inner gonostylus lighter yellowish white. T9 apically shallowly bilobed. Ventral lobe of gonocoxite rounded, about ½ the length of gonocoxite. Inner gonostylus approximately equal in length to gonocoxite, lobe rounded, globe-shaped, rostral prolongation of moderate length, lightly arched, tapering to a blunt tip, bearing two long spines at about ⅔ length of the prolongation. Outer gonostylus a strongly arched rod, narrowing gradually to a sharp tip. Aedeagus a little shorter than gonocoxite, the tip faintly bilobed, parameres about ¾ the length of the aedeagus, lateral lobe low and rounded, mesal lobe gradually tapering to a sharp curved tip.
</paragraph>
<paragraph blockId="13.[121,750,537,1500]" box="[151,745,1475,1500]" pageId="13" pageNumber="32">
<emphasis box="[151,285,1476,1499]" italics="true" pageId="13" pageNumber="32">Dimensions.</emphasis>
Body length
<quantity box="[428,512,1475,1499]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="13" pageNumber="32" unit="mm" value="6.8">6.8 mm</quantity>
, wing length
<quantity box="[656,741,1476,1500]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="32" unit="mm" value="7.3">7.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="32" type="etymology">
<paragraph blockId="13.[121,748,1521,1574]" pageId="13" pageNumber="32">
<emphasis bold="true" box="[121,245,1521,1545]" pageId="13" pageNumber="32">Etymology</emphasis>
. This species is named with reference to the globose lobe of the inner gonostylus.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="32" type="discussion">
<paragraph blockId="13.[121,749,1595,1825]" pageId="13" pageNumber="32">
<emphasis bold="true" box="[121,241,1595,1619]" pageId="13" pageNumber="32">Discussion</emphasis>
.
<taxonomicName authorityName="Billingham &amp; Theischinger" authorityYear="2022" box="[256,541,1595,1619]" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="globosa" subGenus="Dicranomyia">
<emphasis box="[256,541,1595,1619]" italics="true" pageId="13" pageNumber="32">Dicranomyia (D.) globosa</emphasis>
</taxonomicName>
is closely allied to numerous morphologically similar species around
<taxonomicName authority="Skuse, 1890" authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="saxatilis" subGenus="Dicranomyia">
<emphasis italics="true" pageId="13" pageNumber="32">D. (D.) saxatilis</emphasis>
<bibRefCitation author="Skuse, F. A. A." box="[212,339,1654,1678]" pageId="13" pageNumber="32" pagination="757 - 892" refId="ref12726" refString="Skuse, F. A. A. 1890. Diptera of Australia. Part VII. The Tipulidae Brevipalpi. Proceedings of the Linnean Society of New South Wales 4: 757 - 892. https: // doi. org / 10.5962 / bhl. part. 15067" type="journal article" year="1890">Skuse, 1890</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation author="Theischinger, G." box="[389,583,1654,1678]" pageId="13" pageNumber="32" pagination="37 - 276" refId="ref12806" refString="Theischinger, G. 1994. The Limoniinae (Diptera: Tipulidae) of Australia. V. The genera Helius Le Peletier &amp; Serville, Toxorhina Loew, Limonia Meigen (part), Tonnoiromyia Alexander and Collessophila gen. nov. (all tribe Limoniini) and Atarba Osten-Sacken, Amphineurus Skuse, Erioptera Meigen, Cheilotrichia Rossi, Gonomyia Meigen and Idiocera Dale (all tribe Eriopterini). Stapfia 36: 37 - 276." type="journal article" year="1994">Theischinger 1994</bibRefCitation>
for more details on these species).
<taxonomicName authorityName="Billingham &amp; Theischinger" authorityYear="2022" box="[319,604,1683,1707]" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="globosa" subGenus="Dicranomyia">
<emphasis box="[319,604,1683,1707]" italics="true" pageId="13" pageNumber="32">Dicranomyia (D.) globosa</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName authorityName="Skuse" authorityYear="1890" box="[230,395,1713,1737]" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="saxatilis" subGenus="Dicranomyia">
<emphasis box="[230,395,1713,1737]" italics="true" pageId="13" pageNumber="32">D. (D.) saxatilis</emphasis>
</taxonomicName>
, especially in the wing patterning, but is readily distinguished from
<taxonomicName authorityName="Skuse" authorityYear="1890" box="[472,643,1742,1766]" class="Insecta" family="Limoniidae" genus="Dicranomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="saxatilis" subGenus="Dicranomyia">
<emphasis box="[472,643,1742,1766]" italics="true" pageId="13" pageNumber="32">D. (D.) saxatilis</emphasis>
</taxonomicName>
and other similar species by the long spines on the rostral prolongation of the outer gonostylus.
</paragraph>
</subSubSection>
</treatment>
</document>