860 lines
117 KiB
XML
860 lines
117 KiB
XML
<document id="3ED808E7566643596EF0009E544492F6" ID-DOI="10.5252/geodiversitas2024v46a6" ID-ISSN="1638-9395" ID-Zenodo-Dep="11106598" ID-ZooBank="urn:lsid:zoobank.org:pub:6ACF6A79-9149-4781-808D-478668673EB6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1714640714399" checkinUser="plazi" docAuthor="Scavezzoni, Isaure, Fischer, Valentin, Johnson, Michela M. & Jouve, Stéphane" docDate="2024" docId="038A56761D0FFFC0FC8F9314FC6B54C1" docLanguage="en" docName="geodiversitas.2024.46.6.pdf" docOrigin="Geodiversitas 46 (6)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a6.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Metriorhynchus brachyrhynchus , NHMUK PV R" docType="treatment" docVersion="4" lastPageNumber="188" masterDocId="FFB32E0E1D3CFFF7FFA29243FFC5576E" masterDocTitle="Form and function of the pelvic girdle of Thalattosuchia and Dyrosauridae (Crocodyliformes)" masterLastPageNumber="326" masterPageNumber="135" pageNumber="184" updateTime="1714680973700" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="356D96B14CB077F5DAE0588071BE160F">Form and function of the pelvic girdle of Thalattosuchia and Dyrosauridae (Crocodyliformes)</mods:title>
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<mods:namePart id="E1D9043713C9E59493FD09B2CBD2C70D">Scavezzoni, Isaure</mods:namePart>
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<mods:affiliation id="851340F398079890BB51756AEFAEC0F3">Universite de Liège, Evolution and Diversity Dynamics Lab, All. du Six Août 14, 4000 Liège (Belgique) isaure. scavezzoni @ gmail. com v. fischer @ uliege. be</mods:affiliation>
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<mods:nameIdentifier id="F49EEE5E82D9DF263BE750DCDE8C49BC" type="email">isaure.scavezzoni@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3ACCCECA57257C37559AB57C5D02EA6A">Fischer, Valentin</mods:namePart>
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<mods:affiliation id="A9E1A2F67BC7197BF1D2596C2318ED75">Universite de Liège, Evolution and Diversity Dynamics Lab, All. du Six Août 14, 4000 Liège (Belgique) isaure. scavezzoni @ gmail. com v. fischer @ uliege. be</mods:affiliation>
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<mods:nameIdentifier id="E762A5EBD9C29910DB43A8FB753CCE8C" type="email">v.fischer@uliege.be</mods:nameIdentifier>
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<mods:namePart id="1CA08926601A786C3C83CBBE0BA82AF1">Johnson, Michela M.</mods:namePart>
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<mods:affiliation id="F62D454769F90382F4B012A6320F93B0">Department of Palaeontology, Staatliches Museum für Naturkunde Stuttgart, Museum am LÖwentor, Rosenstein 1, 70191 Stuttgart (Germany) michela. johnson @ smns-bw. de</mods:affiliation>
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<mods:nameIdentifier id="4CA12DF0E371F82713AC6A28F6A0DF84" type="email">michela.johnson@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="298A59EAA511418CF97960518062AE45">Jouve, Stéphane</mods:namePart>
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<mods:affiliation id="03739DB336877DACFD5D386E215162F2">Sorbonne Universite, BUPMC - Pôle Collections, Tour Zamansky, 15 étage, bureau 1513, 4 place Jussieu, 75252 Paris Cedex 05 (France) stephane. jouve @ sorbonne-universite. fr</mods:affiliation>
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<mods:nameIdentifier id="52B68E3DC5443667B7AD14B315E6D79D" type="email">stephane.jouve@sorbonne-universite.fr</mods:nameIdentifier>
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<mods:title id="8C034C804C798D6A5FC1C1754945A810">Geodiversitas</mods:title>
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<mods:part id="74C80055A83A945C72E363240E17ED24">
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<mods:date id="13CEA5FB8A42DFF23CB3C72C73737474">2024</mods:date>
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<mods:number id="8A625BB8C2DF51F1E05236F9E7862EE1">2024-05-02</mods:number>
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<mods:number id="70A7CD4EA55860CE6D2A84A1266EB811">46</mods:number>
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<mods:url id="9CC29A03E5EF925448BE5FE34F1A5BA8">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a6.pdf</mods:url>
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<mods:classification id="8303E342020F07F130A4D1DB2C7C7A0D">journal article</mods:classification>
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<mods:identifier id="8B162E37A6730FCA9D183B9D7623D55A" type="DOI">10.5252/geodiversitas2024v46a6</mods:identifier>
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<mods:identifier id="D2D90E0A39E90E8F5C0370CC9ABF5ACB" type="ISSN">1638-9395</mods:identifier>
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<treatment id="038A56761D0FFFC0FC8F9314FC6B54C1" ID-DOI="http://doi.org/10.5281/zenodo.11105976" ID-Zenodo-Dep="11105976" LSID="urn:lsid:plazi:treatment:038A56761D0FFFC0FC8F9314FC6B54C1" httpUri="http://treatment.plazi.org/id/038A56761D0FFFC0FC8F9314FC6B54C1" lastPageId="55" lastPageNumber="188" pageId="51" pageNumber="184">
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<subSubSection id="C339B4EB1D0FFFC4FC8F9314FBFF56FF" pageId="51" pageNumber="184" type="nomenclature">
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<paragraph id="8B9CE7601D0FFFC4FC8F9314FBFF56FF" blockId="51.[813,1328,343,433]" pageId="51" pageNumber="184">
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<taxonomicName id="4C239CE31D0FFFC4FC8F9314FBFF56FF" authority="(EUDES-DESLONGCHAMPS, 1867)" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="313" phylum="Chordata" rank="species" species="brachyrhynchus">
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‘
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<emphasis id="B9573B721D0FFFC4FC91931BFC38561F" box="[819,1021,343,369]" italics="true" pageId="51" pageNumber="184">METRIORHYNCHUS’</emphasis>
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<smallCapsWord id="8D7A71BC1D0FFFC4FBA1931FFB7A5601" baselines="362" box="[1027,1215,348,367]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="brachyrhynchus" pageId="51" pageNumber="184">
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<emphasis id="B9573B721D0FFFC4FBA1931FFB7A5601" box="[1027,1215,348,367]" italics="true" pageId="51" pageNumber="184">BRACHYRHYNCHUS</emphasis>
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</smallCapsWord>
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(
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<bibRefCitation id="EFB29A911D0FFFC4FB729314FBF756FF" author="EUDES-DESLONGCHAMPS J. - A." pageId="51" pageNumber="184" refId="ref110195" refString="EUDES-DESLONGCHAMPS J. - A. 1867. - Note sur les teleosauriens [posthumous work]. Le Blanc-Hardel, Caen, 34 p." type="book" year="1867">
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<smallCapsWord id="8D7A71BC1D0FFFC4FB729314FC2E56E1" baselines="362,362,362,394,394" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Eudes-Deslongchamps" pageId="51" pageNumber="184">EUDES-DESLONGCHAMPS</smallCapsWord>
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, 1867
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</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C339B4EB1D0FFFC4FBE39334FAF556FF" box="[1089,1328,375,401]" pageId="51" pageNumber="184" type="materials_examined">
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<paragraph id="8B9CE7601D0FFFC4FBE39334FAF556FF" blockId="51.[813,1328,343,433]" box="[1089,1328,375,401]" pageId="51" pageNumber="184">NHMUK PV R 3804</paragraph>
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</subSubSection>
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<subSubSection id="C339B4EB1D0FFFC0FC8F93D4FC6B54C1" lastPageId="55" lastPageNumber="188" pageId="51" pageNumber="184" type="description">
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<paragraph id="8B9CE7601D0FFFC4FC8F93D4FB5956DF" blockId="51.[813,1328,343,433]" box="[813,1180,406,433]" pageId="51" pageNumber="184">
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For measurements, see
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<tableCitation id="C6A1D2DB1D0FFFC4FB8793D5FB5056DF" box="[1061,1173,406,433]" captionStart-0="TABLE" captionStart-1="TABLE" captionStart-2="TABLE" captionStartId-0="19.[61,72,462,479]" captionStartId-1="25.[61,72,457,474]" captionStartId-2="29.[61,72,511,528]" captionTargetPageId-0="19" captionTargetPageId-1="25" captionTargetPageId-2="29" captionText-0="TABLE 7. — List of measurements in mm of crocodyliformes ilia. Abbreviations: Ha, anterior height; Hv, ventral length; Hd, dorsal length; PST.l, postacetabular process anteroposterior length; PST.h, postacetabular process dorsoventral height; PRE.l, preacetabular process anteroposterior length; PRE.h, preacetabular process dorsoventral height; ACE.h, acetabulum dorsoventral height; ACE.w, acetabulum width at mid-height; GAP.w, base length of acetabular perforation; GAP.h, max height of acetabular perforation; PUB.l, pubic peduncle anteroposterior length; PUB.h, pubic peduncle dorsoventral height; ISC.l, ischial peduncle anteroposterior length; ISC.h, Ischial peduncle dorsoventral height." captionText-1="TABLE 8. — List of measurements in mm of crocodyliformes ischia. When measurements are missing on the first ischium, it takes measurements from next ischium. NHMUK PV R 4763 shaft reconstructed.Abbreviations: Hd, total height from posterior peduncle to tip of posterior process; PP.h, height of posterior process tip; AP.h, base height of anterior process; AP.w, total length of anterior process; BL.l, total length of ischial blade; GAP.w, base length of acetabular perforation; GAP.h, max height of acetabular perforation;PPed.Ll, anteroposterior length of posterior peduncle taken laterally;PPed.Wa, mediolateral width of posterior peduncle taken anteriorly; BR.l, total length of peduncle bridge; APed.w, mediolateral width of anterior peduncle; APed.h, dorsoventral height of anterior peduncle; Arch.l, anterior arch length from anterior peduncle to anterior process; Arch.h, maximal depth between base arch and bone margin; S, length from proximal extremity to maximal constriction; C, anteroposterior width of constriction." captionText-2="TABLE 9. — List of measurements in mm of crocodyliformes pubes. When measurements are missing on the first bone, it takes measurements from the next one. SMNS 81608 has been reconstructed. Abbreviations: H, total proximodistal height; Wh, mediolateral width at mid length; Ldia, length of diaphysis; Lbl, freehand length of distal blade up to diaphysis; Dbl, distance between both medial and lateral extremities of distal blade; Wprox, anteroposterior width of proximal peduncle; Lprox, mediolateral length of proximal peduncle; Shaft, proximodistal length of shaft; Neck, proximodistal length of neck; C, anteroposterior width of constriction; Arch.l, anterior arch length from peduncle to pubic diaphysis; Arch.h, maximal depth between base arch and bone margin; Arch.d, distance between proximal base of arch and position of max depth along the arch." httpUri-0="http://table.plazi.org/id/DF5CB7E81D2FFFE4FF9F938DFDDF554D" httpUri-1="http://table.plazi.org/id/DF5CB7E81D25FFEEFF9F938AFAB1555A" httpUri-2="http://table.plazi.org/id/DF5CB7E81D21FFEAFF9F93BCF8FE553A" pageId="51" pageNumber="184" tableUuid-0="DF5CB7E81D2FFFE4FF9F938DFDDF554D" tableUuid-1="DF5CB7E81D25FFEEFF9F938AFAB1555A" tableUuid-2="DF5CB7E81D21FFEAFF9F93BCF8FE553A">Tables 7-9</tableCitation>
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.
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</paragraph>
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<paragraph id="8B9CE7601D0FFFC4FC8F9395FCA2569E" blockId="51.[813,871,470,496]" box="[813,871,470,496]" pageId="51" pageNumber="184">
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<heading id="D0D4500C1D0FFFC4FC8F9395FCA2569E" box="[813,871,470,496]" fontSize="11" level="2" pageId="51" pageNumber="184" reason="8">
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<emphasis id="B9573B721D0FFFC4FC8F9395FCA2569E" box="[813,871,470,496]" italics="true" pageId="51" pageNumber="184">Ilium</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9CE7601D0FFFC4FC8F93B5FB1E53C0" blockId="51.[811,1457,502,2027]" pageId="51" pageNumber="184">
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The ilium of ‘
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<taxonomicName id="4C239CE31D0FFFC4FC6093B5FCFA555F" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="brachyrhynchus">
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<emphasis id="B9573B721D0FFFC4FC6093B5FBA6557E" box="[962,1123,502,528]" italics="true" pageId="51" pageNumber="184">Metriorhynchus</emphasis>
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’
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<emphasis id="B9573B721D0FFFC4FBD093B5FAD5557E" box="[1138,1296,502,528]" italics="true" pageId="51" pageNumber="184">brachyrhynchus</emphasis>
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NHMUK PV R
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</taxonomicName>
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3804 (
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<figureCitation id="1318FBE51D0FFFC4FC289055FC11555E" box="[906,980,534,560]" captionStart="FIG" captionStartId="52.[132,143,1963,1980]" captionTargetBox="[129,1457,198,1873]" captionTargetId="figure-17@52.[129,1486,205,1971]" captionTargetPageId="52" captionText="FIG. 24. — Pelvic girdle elements of ‘Metriorhynchus’ brachyrhynchus (Eudes-Deslongchamps,1867), NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens Andrews, 1913): A, right ilium in lateral view; B, right ilium in medial view; C, right pubis in anteroposterior;D, left ischium in lateral view. Target indicates anterior. Arrow points anteriorly. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106650" httpUri="https://zenodo.org/record/11106650/files/figure.png" pageId="51" pageNumber="184">Fig. 24</figureCitation>
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) bear the typical metriorhynchoid characteristics, which are an overall small size in relation to the other pelvic bones, and the absence of a postacetabular process. The preacetabular process of ‘
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<taxonomicName id="4C239CE31D0FFFC4FBC69035FC1955DE" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="brachyrhynchus">
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<emphasis id="B9573B721D0FFFC4FBC69035FAC055FE" box="[1124,1285,630,656]" italics="true" pageId="51" pageNumber="184">Metriorhynchus</emphasis>
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’
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<emphasis id="B9573B721D0FFFC4FAB39035FA6A55FE" box="[1297,1455,630,656]" italics="true" pageId="51" pageNumber="184">brachyrhynchus</emphasis>
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NHMUK PV R
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</taxonomicName>
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3804 is short but lateromedially thick and points anterodorsally (forms an angle of approximately 36° with the ventral margin of the ischial peduncle) rather than strictly anteriorly like extant crocodylians (e.g.
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<taxonomicName id="4C239CE31D0FFFC4FA9C90B6FC1C545E" authority="NMW" authorityName="NMW" class="Reptilia" family="Crocodylidae" genus="Crocodylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="niloticus">
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<emphasis id="B9573B721D0FFFC4FA9C90B6FC425441" italics="true" pageId="51" pageNumber="184">Crocodylus niloticus</emphasis>
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NMW
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</taxonomicName>
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31137) and dyrosaurids (e.g.
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<taxonomicName id="4C239CE31D0FFFC4FA899155FBE35421" authority="MRAC 1806" authorityName="MRAC" authorityYear="1806" class="Reptilia" family="Dyrosauridae" genus="Congosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="bequaerti">
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<emphasis id="B9573B721D0FFFC4FA899155FC4A5421" italics="true" pageId="51" pageNumber="184">Congosaurus bequaerti</emphasis>
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<collectionCode id="ED327FA51D0FFFC4FC369176FC215421" box="[916,996,821,847]" pageId="51" pageNumber="184">MRAC</collectionCode>
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1806
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</taxonomicName>
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). This difference in inclination of the preacetabular process can be attributed to the different way the ilium is borne by the sacrals: the ilium of thalattosuchians is rather tilted in relation to the coronal plane (e.g.
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<bibRefCitation id="EFB29A911D0FFFC4FAFE91D6FC6054A1" author="HERRERA Y. & FERNANDEZ M. & LAMAS S. G. & CAMPOS L. & TALEVI M. & GASPARINI Z." pageId="51" pageNumber="184" pagination="247 - 255" refId="ref111760" refString="HERRERA Y., FERNANDEZ M., LAMAS S. G., CAMPOS L., TALEVI M. & GASPARINI Z. 2017. - Morphology of the sacral region and reproductive strategies of Metriorhynchidae: A counterinductive approach. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 106 (4): 247 - 255. https: // doi. org / 10.1017 / S 1755691016000165" type="journal article" year="2017">
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Herrera
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<emphasis id="B9573B721D0FFFC4FC8F91F6FCA654A1" box="[813,867,949,975]" italics="true" pageId="51" pageNumber="184">et al.</emphasis>
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2017
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</bibRefCitation>
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||
) compared to extant crocodylians (e.g.
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<emphasis id="B9573B721D0FFFC4FAC091F6FC2A5480" italics="true" pageId="51" pageNumber="184">Palaeosuchus palpebrosus</emphasis>
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[
|
||
<figureCitation id="1318FBE51D0FFFC4FBA09197FB875481" box="[1026,1090,980,1007]" captionStart="FIG" captionStartId="18.[132,143,1963,1980]" captionTargetBox="[169,1453,184,1914]" captionTargetId="figure-17@18.[296,1243,221,1867]" captionTargetPageId="18" captionText="FIG. 7. — Snapshot of the pelvic girdle of the alligatoroid Palaeosuchus palpebrosus (Cuvier, 1807), RVC-JRH-PP1: A, in anterior view; B, in lateral view. Note the orientation of the pubes. Cross indicates posterior view. Arrow points anteriorly. CT scan courtesy of Professor John Hutchinson (see https://skfb.ly/6ByyV). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11106614" httpUri="https://zenodo.org/record/11106614/files/figure.png" pageId="51" pageNumber="184">Fig. 7</figureCitation>
|
||
],
|
||
<taxonomicName id="4C239CE31D0FFFC4FBFE9196FA895480" box="[1116,1356,980,1006]" class="Crocodylia" family="Crocodylidae" genus="Mecistops" kingdom="Animalia" order="Eusuchia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="cataphractus">
|
||
<emphasis id="B9573B721D0FFFC4FBFE9196FA895480" box="[1116,1356,980,1006]" italics="true" pageId="51" pageNumber="184">Mecistops cataphractus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="1318FBE51D0FFFC4FAFC9197FA5A5481" box="[1374,1439,980,1007]" captionStart="FIG" captionStartId="20.[132,143,1986,2003]" captionTargetBox="[149,1433,219,1874]" captionTargetId="figure-17@20.[226,1462,231,1900]" captionTargetPageId="20" captionText="FIG. 8. — Right pelvic girdle of Mecistops cataphractus Cuvier, 1825, RBINS 18374: A, ilium in lateral view; B, ischium in lateral view; C, pubis in ventral view; D, ilium in medial view. Cross indicates posterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106616" httpUri="https://zenodo.org/record/11106616/files/figure.png" pageId="51" pageNumber="184">Fig. 8</figureCitation>
|
||
],
|
||
<taxonomicName id="4C239CE31D0FFFC4FC8F91B6FC295360" box="[813,1004,1012,1039]" class="Crocodylia" family="Alligatoridae" genus="Caiman" kingdom="Animalia" order="Eusuchia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="crocodilus">
|
||
<emphasis id="B9573B721D0FFFC4FC8F91B6FC295360" box="[813,1004,1012,1039]" italics="true" pageId="51" pageNumber="184">Caiman crocodilus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="1318FBE51D0FFFC4FC5E91B7FBFC5361" box="[1020,1081,1012,1039]" captionStart="FIG" captionStartId="21.[132,143,1986,2003]" captionTargetBox="[132,1441,166,1905]" captionTargetId="figure-15@21.[-252,1479,167,1865]" captionTargetPageId="21" captionText="FIG. 9. — Left pelvic girdle of Caiman crocodilus (Linnaeus, 1758), NHMW 30900: A, ilium in medial view; B, ilium in lateral view; C, ischium in medial view; D, ischium in lateral view; E, pubis in posterior view. Target indicates anterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106620" httpUri="https://zenodo.org/record/11106620/files/figure.png" pageId="51" pageNumber="184">Fig. 9</figureCitation>
|
||
]) or dyrosaurids (
|
||
<taxonomicName id="4C239CE31D0FFFC4FB5891B6FCAE5340" authorityName="MRAC" authorityYear="1806" class="Reptilia" family="Dyrosauridae" genus="Congosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="bequaerti">
|
||
<emphasis id="B9573B721D0FFFC4FB5891B6FCAE5340" italics="true" pageId="51" pageNumber="184">Congosaurus bequaerti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0FFFC4FCD59656FB845340" box="[887,1089,1045,1070]" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="natator">
|
||
<emphasis id="B9573B721D0FFFC4FCD59656FB845340" box="[887,1089,1045,1070]" italics="true" pageId="51" pageNumber="184">Hyposaurus natator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0FFFC4FBEC9657FAB55340" authority="Hastings, Bloch & Jaramillo, 2011" authorityName="Hastings, Bloch & Jaramillo" authorityYear="2011" box="[1102,1392,1044,1070]" family="Dyrosauridae" genus="Acherontisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="guajiraensis">
|
||
<emphasis id="B9573B721D0FFFC4FBEC9657FAB55340" box="[1102,1392,1044,1070]" italics="true" pageId="51" pageNumber="184">Acherontisuchus guajiraensis</emphasis>
|
||
</taxonomicName>
|
||
). The preacetabular process of NMH PV R 3804 strongly differs from that of NHMUK PV R 4763, and resembles those of
|
||
<taxonomicName id="4C239CE31D0FFFC4FC8F9637FBDF53E0" authorityName="NHMUK PV R" authorityYear="2618" box="[813,1050,1140,1166]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D0FFFC4FC8F9637FBDF53E0" box="[813,1050,1140,1166]" italics="true" pageId="51" pageNumber="184">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0FFFC4FB8A9637FA8653E0" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" box="[1064,1347,1140,1166]" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0FFFC4FB8A9637FA8653E0" box="[1064,1347,1140,1166]" italics="true" pageId="51" pageNumber="184">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
and more specifically
|
||
<taxonomicName id="4C239CE31D0FFFC4FC0A96D6FB1053C0" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[936,1237,1172,1198]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D0FFFC4FC0A96D6FB1053C0" box="[936,1237,1172,1198]" italics="true" pageId="51" pageNumber="184">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D0FFFC2FCE196F7FD195082" blockId="51.[811,1457,502,2027]" lastBlockId="53.[125,778,215,2028]" lastPageId="53" lastPageNumber="186" pageId="51" pageNumber="184">
|
||
The anterior margin of the ilium of ‘
|
||
<taxonomicName id="4C239CE31D0FFFC4FB1896F7FBFD5380" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0FFFC4FB1896F7FA9C53A0" box="[1210,1369,1204,1230]" italics="true" pageId="51" pageNumber="184">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0FFFC4FAC696F7FC425380" italics="true" pageId="51" pageNumber="184">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 underneath the preacetabular process is convex, and meets distally with the pubic peduncle of the ilium. The dorsal margin is globally reduced as the anterior and posterior margins almost directly connect with the large base of the preacetabular process (
|
||
<figureCitation id="1318FBE51D0FFFC4FB6A9710FAD45203" box="[1224,1297,1363,1389]" captionStart="FIG" captionStartId="52.[132,143,1963,1980]" captionTargetBox="[129,1457,198,1873]" captionTargetId="figure-17@52.[129,1486,205,1971]" captionTargetPageId="52" captionText="FIG. 24. — Pelvic girdle elements of ‘Metriorhynchus’ brachyrhynchus (Eudes-Deslongchamps,1867), NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens Andrews, 1913): A, right ilium in lateral view; B, right ilium in medial view; C, right pubis in anteroposterior;D, left ischium in lateral view. Target indicates anterior. Arrow points anteriorly. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106650" httpUri="https://zenodo.org/record/11106650/files/figure.png" pageId="51" pageNumber="184">Fig. 24</figureCitation>
|
||
). The junction between the dorsal and posterior margins is achieved through an obtuse and rounded corner, similar to those of
|
||
<taxonomicName id="4C239CE31D0FFFC4FAF297D0FC7152A3" authorityName="NHMUK PV R" authorityYear="2618" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D0FFFC4FAF297D0FC7152A3" italics="true" pageId="51" pageNumber="184">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0FFFC4FC1D97F7FB2252A3" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[959,1255,1459,1485]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D0FFFC4FC1D97F7FB2252A3" box="[959,1255,1459,1485]" italics="true" pageId="51" pageNumber="184">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C239CE31D0FFFC4FABB97F0FB655280" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0FFFC4FABB97F0FC6F5283" italics="true" pageId="51" pageNumber="184">Thalattosuchus superciliosus</emphasis>
|
||
NHMUK PV R 2054
|
||
</taxonomicName>
|
||
. The posterior margin of the ilium is concave and ends distally to form the posterior corner of the ischial peduncle of the ischium. The ventral margin of the ilium is slightly undulated: it bears the pubic peduncle of the ilium anteriorly, and the ischial peduncle of the ischium posteriorly. The undulation of the ventral margin of ‘
|
||
<taxonomicName id="4C239CE31D0FFFC4FCD494D1FAB551C3" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[886,1392,1682,1709]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0FFFC4FCD494D1FBD251C2" box="[886,1047,1682,1708]" italics="true" pageId="51" pageNumber="184">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0FFFC4FB8094D1FB0551C2" box="[1058,1216,1682,1708]" italics="true" pageId="51" pageNumber="184">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 is more subtle than that of ‘
|
||
<taxonomicName id="4C239CE31D0FFFC4FBFD94F1FC205182" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0FFFC4FBFD94F1FAC451A2" box="[1119,1281,1714,1740]" italics="true" pageId="51" pageNumber="184">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0FFFC4FAB294F1FA6A51A2" box="[1296,1455,1714,1740]" italics="true" pageId="51" pageNumber="184">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763 or
|
||
<taxonomicName id="4C239CE31D0FFFC4FBF09491FAB05182" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" box="[1106,1397,1746,1772]" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0FFFC4FBF09491FAB05182" box="[1106,1397,1746,1772]" italics="true" pageId="51" pageNumber="184">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
, and closely resembles that of
|
||
<taxonomicName id="4C239CE31D0FFFC4FBEA94B0FABA5062" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[1096,1407,1778,1804]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D0FFFC4FBEA94B0FABA5062" box="[1096,1407,1778,1804]" italics="true" pageId="51" pageNumber="184">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C239CE31D0FFFC4FC8F9551FBDB5042" authorityName="NHMUK PV R" authorityYear="2618" box="[813,1054,1810,1836]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="51" pageNumber="184" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D0FFFC4FC8F9551FBDB5042" box="[813,1054,1810,1836]" italics="true" pageId="51" pageNumber="184">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
. The ischial peduncle represents the region where the ilium is the thickest mediolaterally. Comparatively, the ischial peduncle is subtlety longer dorsoventrally than the pubic peduncle, so that both peduncles appear to have the same height. Like in other metriorhynchoids, the ischial peduncle takes the shape of an isosceles triangle whose orientation is both lateral and anterior. The outline of the pubic peduncle diplays three successive triangular mounds, with the supraacetabular crest originating from the hollow between the two first summits. On the lateral side of the ilium, the pubic peduncle gradually transitions to the ischial peduncle through a recess but without rupturing the articular surface (as in other metriorhynchoids). There is a subtle opening along the ventral margin of the ilium of ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FDFC93D5FE1856BF" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FDFC93D5FCC556DE" box="[606,768,406,432]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FF2693F5FEE656BE" box="[132,291,438,464]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 corresponding to the acetabular perforation. The acetabular perforation almost indiscernible in ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FE9C93B5FF06555F" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FE9C93B5FE24557E" box="[318,481,502,528]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FE5293B5FD4A557E" box="[496,655,502,528]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804, as in
|
||
<taxonomicName id="4C239CE31D09FFC2FEFB9055FD96555E" authorityName="NHMUK PV R" authorityYear="2618" box="[345,595,534,560]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D09FFC2FEFB9055FD96555E" box="[345,595,534,560]" italics="true" pageId="53" pageNumber="186">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FDC69054FED4553E" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FDC69054FED4553E" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C239CE31D09FFC2FEF79075FDED553E" box="[341,552,566,592]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9573B721D09FFC2FEF79075FE0A553E" box="[341,463,566,592]" italics="true" pageId="53" pageNumber="186">Cricosaurus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
The acetabular perforation of the ilium of ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FE079015FEF155FE" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FE079015FD8A551E" box="[421,591,598,624]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FDC29015FCC3551E" box="[608,774,598,624]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 does not separate the peduncles of the ilium but is rather indicating the transition between both. Indeed, the acetabular perforation on the ilium is slightly shifted anteriorly, thus ending up within the posterior-most portion of the pubic peduncle. Furthermore, the peduncles of the ilium meet each other on the lateral surface of the ilium, well above the ventral margin of the bone, meaning that both peduncles where brought together presumably at the expense of the acetabular perforation. This highly contrasts with dyrosaurids (
|
||
<taxonomicName id="4C239CE31D09FFC2FE2D91D6FDBB54C1" authorityName="MRAC" authorityYear="1806" box="[399,638,917,943]" class="Reptilia" family="Dyrosauridae" genus="Congosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="bequaerti">
|
||
<emphasis id="B9573B721D09FFC2FE2D91D6FDBB54C1" box="[399,638,917,943]" italics="true" pageId="53" pageNumber="186">Congosaurus bequaerti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FD2C91D5FF1554A0" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="natator">
|
||
<emphasis id="B9573B721D09FFC2FD2C91D5FF1554A0" italics="true" pageId="53" pageNumber="186">Hyposaurus natator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FF7E91F6FE3854A0" authority="Hastings, Bloch & Jaramillo, 2011" authorityName="Hastings, Bloch & Jaramillo" authorityYear="2011" box="[220,509,949,975]" family="Dyrosauridae" genus="Acherontisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="guajiraensis">
|
||
<emphasis id="B9573B721D09FFC2FF7E91F6FE3854A0" box="[220,509,949,975]" italics="true" pageId="53" pageNumber="186">Acherontisuchus guajiraensis</emphasis>
|
||
</taxonomicName>
|
||
) and extant crocodylians (e.g.
|
||
<emphasis id="B9573B721D09FFC2FF189197FE7F5480" box="[186,442,980,1006]" italics="true" pageId="53" pageNumber="186">Palaeosuchus palpebrosus</emphasis>
|
||
[
|
||
<figureCitation id="1318FBE51D09FFC2FE689197FDC25481" box="[458,519,980,1007]" captionStart="FIG" captionStartId="18.[132,143,1963,1980]" captionTargetBox="[169,1453,184,1914]" captionTargetId="figure-17@18.[296,1243,221,1867]" captionTargetPageId="18" captionText="FIG. 7. — Snapshot of the pelvic girdle of the alligatoroid Palaeosuchus palpebrosus (Cuvier, 1807), RVC-JRH-PP1: A, in anterior view; B, in lateral view. Note the orientation of the pubes. Cross indicates posterior view. Arrow points anteriorly. CT scan courtesy of Professor John Hutchinson (see https://skfb.ly/6ByyV). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11106614" httpUri="https://zenodo.org/record/11106614/files/figure.png" pageId="53" pageNumber="186">Fig. 7</figureCitation>
|
||
],
|
||
<taxonomicName id="4C239CE31D09FFC2FDBC9196FCC35480" box="[542,774,980,1006]" class="Crocodylia" family="Crocodylidae" genus="Mecistops" kingdom="Animalia" order="Eusuchia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="cataphractus">
|
||
<emphasis id="B9573B721D09FFC2FDBC9196FCC35480" box="[542,774,980,1006]" italics="true" pageId="53" pageNumber="186">Mecistops cataphractus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="1318FBE51D09FFC2FF2F91B7FF0C5361" box="[141,201,1012,1039]" captionStart="FIG" captionStartId="20.[132,143,1986,2003]" captionTargetBox="[149,1433,219,1874]" captionTargetId="figure-17@20.[226,1462,231,1900]" captionTargetPageId="20" captionText="FIG. 8. — Right pelvic girdle of Mecistops cataphractus Cuvier, 1825, RBINS 18374: A, ilium in lateral view; B, ischium in lateral view; C, pubis in ventral view; D, ilium in medial view. Cross indicates posterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106616" httpUri="https://zenodo.org/record/11106616/files/figure.png" pageId="53" pageNumber="186">Fig. 8</figureCitation>
|
||
],
|
||
<taxonomicName id="4C239CE31D09FFC2FF7C91B6FE595360" box="[222,412,1012,1039]" class="Crocodylia" family="Alligatoridae" genus="Caiman" kingdom="Animalia" order="Eusuchia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="crocodilus">
|
||
<emphasis id="B9573B721D09FFC2FF7C91B6FE595360" box="[222,412,1012,1039]" italics="true" pageId="53" pageNumber="186">Caiman crocodilus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="1318FBE51D09FFC2FE0B91B7FE205361" box="[425,485,1012,1039]" captionStart="FIG" captionStartId="21.[132,143,1986,2003]" captionTargetBox="[132,1441,166,1905]" captionTargetId="figure-15@21.[-252,1479,167,1865]" captionTargetPageId="21" captionText="FIG. 9. — Left pelvic girdle of Caiman crocodilus (Linnaeus, 1758), NHMW 30900: A, ilium in medial view; B, ilium in lateral view; C, ischium in medial view; D, ischium in lateral view; E, pubis in posterior view. Target indicates anterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106620" httpUri="https://zenodo.org/record/11106620/files/figure.png" pageId="53" pageNumber="186">Fig. 9</figureCitation>
|
||
]) where both peduncles are entirely separated by a large acetabular perforation. In ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FD1B9656FD975321" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FD1B9656FF245320" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FF539677FE575320" box="[241,402,1076,1102]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 and other metriorhynchoids (e.g.
|
||
<taxonomicName id="4C239CE31D09FFC2FED99617FDA05300" authorityName="NHMUK PV R" authorityYear="2618" box="[379,613,1108,1134]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D09FFC2FED99617FDA05300" box="[379,613,1108,1134]" italics="true" pageId="53" pageNumber="186">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FDD29617FF3F53E0" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D09FFC2FDD29617FF3F53E0" italics="true" pageId="53" pageNumber="186">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FEB29637FE2A53E0" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[272,495,1140,1166]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B9573B721D09FFC2FEB29637FE2A53E0" box="[272,495,1140,1166]" italics="true" pageId="53" pageNumber="186">Dakosaurus maximus</emphasis>
|
||
</taxonomicName>
|
||
, etc.), the ilium is greatly reduced while still conserving the bony acetabulum and the connections to the other hip bones (i.e. the peduncles), and this reduction presumably happened at the expense of the acetabular perforation and the postacetabular process. The bony acetabulum is limited dorsally by the supraacetabular crest, and runs all the way down to the peduncles ventrally. The supraacetabular crest is a parabolic-shaped ridge that is prominent anteriorly and fades posteriorly. The supraacetabular crest is not centered on the ilium, and its posterior border is slightly closer to the posterior margin of the bone than its anterior border is to the anterior margin of the ilium unlike in ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FF5597B0FCCC5163" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[247,777,1523,1549]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FF5597B0FE5F5163" box="[247,410,1523,1549]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FE0897B0FD8F5163" box="[426,586,1523,1549]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763. The area comprising the anterodorsal surface of the acetabulum up to the anterior margin of the ilium, across the supraacetabular crest, is rugged thus reflecting the anchoring of cartilage. Like in other metriorhynchoids, this coarse area extends over the anterior portion of the acetabulum as the rest of the surface is smoother (e.g.
|
||
<taxonomicName id="4C239CE31D09FFC2FE5D94F0FF765182" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FE5D94F0FF765182" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FF1D9491FE6F5182" authorityName="NHMUK PV R" authorityYear="2618" box="[191,426,1746,1772]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D09FFC2FF1D9491FE6F5182" box="[191,426,1746,1772]" italics="true" pageId="53" pageNumber="186">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
, ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FE1F9491FEF55062" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FE1F9491FD985182" box="[445,605,1746,1772]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FDCB9491FCC35182" box="[617,774,1746,1772]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763,
|
||
<taxonomicName id="4C239CE31D09FFC2FED994B1FD535062" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" box="[379,662,1778,1804]" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D09FFC2FED994B1FD535062" box="[379,662,1778,1804]" italics="true" pageId="53" pageNumber="186">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
, etc.). The anterior margin of the ilium and supraacetabular crests are, however, strongly pitted and presumably hosted a structure similar to the acetabular labrum of extant crocodylians.
|
||
<taxonomicName id="4C239CE31D09FFC2FD509511FF6A50E5" baseAuthorityName="Tsai & Holliday" baseAuthorityYear="2015" family="Teleosauridae" genus="Indosinosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="vivo">
|
||
<emphasis id="B9573B721D09FFC2FD509511FF6A50E5" italics="true" pageId="53" pageNumber="186">In vivo</emphasis>
|
||
</taxonomicName>
|
||
, the bony acetabulum was probably covered in cartilage which likely extended over the ischium and pubis, forming a puboischial synchondrosis like that of extant crocodylians (
|
||
<bibRefCitation id="EFB29A911D09FFC2FF2F9592FE145085" author="CLAESSENS L. & VICKARYOUS M. K." box="[141,465,2001,2027]" pageId="53" pageNumber="186" pagination="1185 - 1198" refId="ref109599" refString="CLAESSENS L. & VICKARYOUS M. K. 2012. - The evolution, development and skeletal identity of the crocodylian pelvis: Revisiting a forgotten scientific debate. Journal of Morphology 273 (10): 1185 - 1198. https: // doi. org / 10.1002 / jmor. 20059" type="journal article" year="2012">Claessens & Vickaryous 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB29A911D09FFC2FE7C9592FD085082" author="TSAI H. P. & HOLLIDAY C. M." box="[478,717,2001,2028]" pageId="53" pageNumber="186" pagination="1 - 30" refId="ref116719" refString="TSAI H. P. & HOLLIDAY C. M. 2015. - Articular soft tissue anatomy of the archosaur hip joint: Structural homology and functional implications. Journal of Morphology 276 (6): 1 - 30. https: // doi. org / 10.1002 / jmor. 20360" type="journal article" year="2015">Tsai & Holliday 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5CB7E81D08FFC3FF2695E8FE1B5087" ID-DOI="http://doi.org/10.5281/zenodo.11106650" ID-Zenodo-Dep="11106650" httpUri="https://zenodo.org/record/11106650/files/figure.png" pageId="52" pageNumber="185" startId="52.[132,143,1963,1980]" targetBox="[129,1457,198,1873]" targetPageId="52" targetType="figure">
|
||
<paragraph id="8B9CE7601D08FFC3FF2695E8FE1B5087" blockId="52.[131,1457,1963,2025]" pageId="52" pageNumber="185">
|
||
<smallCapsWord id="8D7A71BC1D08FFC3FF2695E8FF5850D5" baselines="1976,1976" box="[132,157,1963,1980]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="52" pageNumber="185">FIG</smallCapsWord>
|
||
. 24. — Pelvic girdle elements of ‘
|
||
<taxonomicName id="4C239CE31D08FFC3FE0F95E8FC0550D2" authority="(Eudes-Deslongchamps, 1867)" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[429,960,1963,1980]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="52" pageNumber="185" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D08FFC3FE0F95E8FDEB50D2" box="[429,558,1963,1980]" italics="true" pageId="52" pageNumber="185">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D08FFC3FD9595E8FD7E50D2" box="[567,699,1963,1980]" italics="true" pageId="52" pageNumber="185">brachyrhynchus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB29A911D08FFC3FD6095E8FC7E50D2" author="EUDES-DESLONGCHAMPS J. - A." box="[706,955,1963,1980]" pageId="52" pageNumber="185" refId="ref110195" refString="EUDES-DESLONGCHAMPS J. - A. 1867. - Note sur les teleosauriens [posthumous work]. Le Blanc-Hardel, Caen, 34 p." type="book" year="1867">Eudes-Deslongchamps,1867</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMUK PV R 3804 (holotype of ‘
|
||
<taxonomicName id="4C239CE31D08FFC3FB7695E8FA9050D2" box="[1236,1365,1963,1980]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="52" pageNumber="185" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9573B721D08FFC3FB7695E8FA9050D2" box="[1236,1365,1963,1980]" italics="true" pageId="52" pageNumber="185">Metriorhynchus</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B9573B721D08FFC3FAFC95E8FA6A50D2" box="[1374,1455,1963,1980]" italics="true" pageId="52" pageNumber="185">cultridens</emphasis>
|
||
<bibRefCitation id="EFB29A911D08FFC3FF219581FEC650BD" author="ANDREWS C. W." box="[131,259,1986,2003]" pageId="52" pageNumber="185" refId="ref108381" refString="ANDREWS C. W. 1913. - A Descriptive Catalogue of the Marine Reptiles of the Oxford Clay. Part II. Order the the Trustees of the British Museum, London, 292 p. https: // doi. org / 10.5962 / bhl. title. 61785" type="book" year="1913">Andrews, 1913</bibRefCitation>
|
||
):
|
||
<emphasis id="B9573B721D08FFC3FEAD9581FED950BD" bold="true" box="[271,284,1986,2003]" pageId="52" pageNumber="185">A</emphasis>
|
||
, right ilium in lateral view;
|
||
<emphasis id="B9573B721D08FFC3FE569581FDC450BD" bold="true" box="[500,513,1986,2003]" pageId="52" pageNumber="185">B</emphasis>
|
||
, right ilium in medial view;
|
||
<emphasis id="B9573B721D08FFC3FD7C9581FD2950BD" bold="true" box="[734,748,1986,2003]" pageId="52" pageNumber="185">C</emphasis>
|
||
, right pubis in anteroposterior;
|
||
<emphasis id="B9573B721D08FFC3FC4F9581FC3E50BD" bold="true" box="[1005,1019,1986,2003]" pageId="52" pageNumber="185">D</emphasis>
|
||
, left ischium in lateral view. Target indicates anterior. Arrow points anteriorly. Scale bars: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9CE7601D09FFC2FCE19294FBD553A0" blockId="53.[811,1458,215,1677]" pageId="53" pageNumber="186">
|
||
The attachment sites for the two sacral ribs are visible on the medial side of the ilium. They are recognizable by their overall bilobate shape, with the biggest lobe positioned ventrally as in
|
||
<taxonomicName id="4C239CE31D09FFC2FC0A937BFB10563F" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[936,1237,311,337]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FC0A937BFB10563F" box="[936,1237,311,337]" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FB469374FC9E561F" authorityName="NHMUK PV R" authorityYear="2618" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D09FFC2FB469374FC9E561F" italics="true" pageId="53" pageNumber="186">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
, and ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FC009314FA74561F" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[930,1457,343,369]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FC009314FB81561F" box="[930,1092,343,369]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FBF69314FB36561F" box="[1108,1267,343,369]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763. The anterior attachment site is slightly larger than the posterior one, and both show a relatively smooth ventral lobe along with a rougher dorsal lobe indicating the existence of cartilage joining the ribs and the ilium
|
||
<taxonomicName id="4C239CE31D09FFC2FB7F9394FAE1569E" baseAuthorityName="Tsai & Holliday" baseAuthorityYear="2015" box="[1245,1316,471,496]" family="Teleosauridae" genus="Indosinosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="264" phylum="Chordata" rank="species" species="vivo">
|
||
<emphasis id="B9573B721D09FFC2FB7F9394FAE1569E" box="[1245,1316,471,496]" italics="true" pageId="53" pageNumber="186">in vivo</emphasis>
|
||
</taxonomicName>
|
||
. The attachments sites do not form depressions on the ilium, but are rather in relief or raised, unlike in
|
||
<taxonomicName id="4C239CE31D09FFC2FB2C9055FA62555E" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" box="[1166,1447,534,560]" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D09FFC2FB2C9055FA62555E" box="[1166,1447,534,560]" italics="true" pageId="53" pageNumber="186">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FC8F9075FBF0553E" baseAuthorityName="Gasparini & Dellape" baseAuthorityYear="1976" box="[813,1077,566,592]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="araucanensis">
|
||
<emphasis id="B9573B721D09FFC2FC8F9075FBF0553E" box="[813,1077,566,592]" italics="true" pageId="53" pageNumber="186">Cricosaurus araucanensis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C239CE31D09FFC2FBFC9075FA86553E" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[1118,1347,566,592]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B9573B721D09FFC2FBFC9075FA86553E" box="[1118,1347,566,592]" italics="true" pageId="53" pageNumber="186">Dakosaurus maximus</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, the sacral rib attachment sites are adjacent, and both situated relatively high on the ilium near the preacetabular process, which is similar to other derived metriorhynchoids (e.g.
|
||
<taxonomicName id="4C239CE31D09FFC2FCC390F6FB8A55A1" authorityName="NHMUK PV R" authorityYear="2618" box="[865,1103,693,719]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D09FFC2FCC390F6FB8A55A1" box="[865,1103,693,719]" italics="true" pageId="53" pageNumber="186">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
, ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FBC190F6FC19559E" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FBC190F6FAC155A1" box="[1123,1284,693,719]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FAB390F6FA6A55A1" box="[1297,1455,693,719]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763,
|
||
<taxonomicName id="4C239CE31D09FFC2FB899095FAE95581" baseAuthorityName="Gasparini & Dellape" baseAuthorityYear="1976" box="[1067,1324,726,752]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="araucanensis">
|
||
<emphasis id="B9573B721D09FFC2FB899095FAE95581" box="[1067,1324,726,752]" italics="true" pageId="53" pageNumber="186">Cricosaurus araucanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FA989096FC185461" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D09FFC2FA989096FC185461" italics="true" pageId="53" pageNumber="186">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FC4E90B5FADF5461" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[1004,1306,757,783]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FC4E90B5FADF5461" box="[1004,1306,757,783]" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
, etc.). Yet, all these traits differ from dyrosaurids (e.g.
|
||
<taxonomicName id="4C239CE31D09FFC2FB679155FA6C5441" authorityName="MRAC" authorityYear="1806" box="[1221,1449,789,816]" class="Reptilia" family="Dyrosauridae" genus="Congosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="bequaerti">
|
||
<emphasis id="B9573B721D09FFC2FB679155FA6C5441" box="[1221,1449,789,816]" italics="true" pageId="53" pageNumber="186">Congosaurus bequaerti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FC8F9175FC305421" box="[813,1013,822,847]" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="natator">
|
||
<emphasis id="B9573B721D09FFC2FC8F9175FC305421" box="[813,1013,822,847]" italics="true" pageId="53" pageNumber="186">Hyposaurus natator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D09FFC2FBA39176FAE75421" authority="Hastings, Bloch & Jaramillo, 2011" authorityName="Hastings, Bloch & Jaramillo" authorityYear="2011" box="[1025,1314,821,847]" family="Dyrosauridae" genus="Acherontisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="guajiraensis">
|
||
<emphasis id="B9573B721D09FFC2FBA39176FAE75421" box="[1025,1314,821,847]" italics="true" pageId="53" pageNumber="186">Acherontisuchus guajiraensis</emphasis>
|
||
</taxonomicName>
|
||
) which show distinct elliptic attachments sites for each sacral process. In addition, in dyrosaurids the attachments sites for each sacral are separated by the acetabular perforation of the ilium, and are borne near the ventral margin of the bone. The ventral deflection of the pelvic girdle of metriorhynchoids, which presumably happened concurrently with its reduction, can be held responsible for the dorsal position of the sacral rib attachment sites near the preacetabular process. In comparison, dyrosaurids possess short and upright sacral processes which are anchored along the anterior and posterior margins of the ilium, leading to pelvic girdles positioned higher dorsally than in metriorhynchoids.
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D09FFC2FCE19697FAB651E3" blockId="53.[811,1458,215,1677]" pageId="53" pageNumber="186">
|
||
The shape of the ilium – both overall and in detail – of ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FC9696B0FA965260" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[820,1363,1267,1294]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FC9696B0FC1E5263" box="[820,987,1267,1293]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FC4996B0FB4A5263" box="[1003,1167,1267,1293]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 is strongly similar to that of
|
||
<taxonomicName id="4C239CE31D09FFC2FB9B9757FAA75243" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[1081,1378,1299,1325]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FB9B9757FAA75243" box="[1081,1378,1299,1325]" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
so that both putatively belong to the same taxon. At the present stage ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FC919710FBBB5203" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[819,1150,1363,1389]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FC919710FC115203" box="[819,980,1363,1389]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FC429710FBBB5203" box="[992,1150,1363,1389]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
is a polyphyletic wastebasket taxon (
|
||
<bibRefCitation id="EFB29A911D09FFC2FCDB9730FB8B52E3" author="WASKOW K. & GRZEGORCZYK D. & SANDER P. M." box="[889,1102,1395,1421]" pageId="53" pageNumber="186" pagination="457 - 480" refId="ref117472" refString="WASKOW K., GRZEGORCZYK D. & SANDER P. M. 2018. - The first record of Tyrannoneustes (Thalattosuchia: Metriorhynchidae): A complete skull from the Callovian (late Middle Jurassic) of Germany. PalZ 92 (3): 457 - 480. https: // doi. org / 10.1007 / s 12542 - 017 - 0395 - z" type="journal article" year="2018">
|
||
Waskow
|
||
<emphasis id="B9573B721D09FFC2FC799737FBCA52E3" box="[987,1039,1395,1421]" italics="true" pageId="53" pageNumber="186">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) as it is filled with markedly differing specimens on both cranium and postcranium levels (see ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FCC697F0FAB552A3" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[868,1392,1459,1485]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FCC697F0FBC052A3" box="[868,1029,1459,1485]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FBB697F0FB7752A3" box="[1044,1202,1459,1485]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 and ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FCC69790FAB55283" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[868,1392,1491,1517]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FCC69790FBC05283" box="[868,1029,1491,1517]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FBB69790FB775283" box="[1044,1202,1491,1517]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763 for a striking example). In parallel, the cranium (skull and mandible) of
|
||
<taxonomicName id="4C239CE31D09FFC2FC179450FB1D5142" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[949,1240,1554,1580]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FC179450FB1D5142" box="[949,1240,1554,1580]" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
has been introduced with some characters that appear sometimes approximate and dubious (see
|
||
<bibRefCitation id="EFB29A911D09FFC2FC1E9411FB5A5102" author="FOFFA D. & YOUNG M. T." box="[956,1183,1618,1645]" pageId="53" pageNumber="186" pagination="608" refId="ref110745" refString="FOFFA D. & YOUNG M. T. 2014. - The cranial osteology of Tyrannoneustes lythrodectikos (Crocodylomorpha: Metriorhynchidae) from the Middle Jurassic of Europe. PeerJ 2: e 608. https: // doi. org / 10.7717 / peerj. 608" type="journal article" year="2014">Foffa & Young 2014</bibRefCitation>
|
||
), especially pertaining to ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FC919431FAA851E2" authority="(Waskow et al. 2018)" baseAuthorityName="Waskow" baseAuthorityYear="2018" box="[819,1389,1650,1676]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FC919431FC1151E2" box="[819,980,1650,1676]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FC409431FB4551E2" box="[994,1152,1650,1676]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB29A911D09FFC2FB339431FAA051E2" author="WASKOW K. & GRZEGORCZYK D. & SANDER P. M." box="[1169,1381,1650,1676]" pageId="53" pageNumber="186" pagination="457 - 480" refId="ref117472" refString="WASKOW K., GRZEGORCZYK D. & SANDER P. M. 2018. - The first record of Tyrannoneustes (Thalattosuchia: Metriorhynchidae): A complete skull from the Callovian (late Middle Jurassic) of Germany. PalZ 92 (3): 457 - 480. https: // doi. org / 10.1007 / s 12542 - 017 - 0395 - z" type="journal article" year="2018">
|
||
Waskow
|
||
<emphasis id="B9573B721D09FFC2FB509430FAE051E2" box="[1266,1317,1650,1676]" italics="true" pageId="53" pageNumber="186">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D09FFC2FC8F94F1FCBB51A2" blockId="53.[813,894,1714,1740]" box="[813,894,1714,1740]" pageId="53" pageNumber="186">
|
||
<heading id="D0D4500C1D09FFC2FC8F94F1FCBB51A2" box="[813,894,1714,1740]" fontSize="11" level="2" pageId="53" pageNumber="186" reason="8">
|
||
<emphasis id="B9573B721D09FFC2FC8F94F1FCBB51A2" box="[813,894,1714,1740]" italics="true" pageId="53" pageNumber="186">Ischium</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D09FFC0FC8F9491FEC25441" blockId="53.[813,1457,1746,2027]" lastBlockId="55.[130,777,215,815]" lastPageId="55" lastPageNumber="188" pageId="53" pageNumber="186">
|
||
The ischium of ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FC469491FCAC5062" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FC469491FB425182" box="[996,1159,1746,1772]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FB359491FAF25182" box="[1175,1335,1746,1772]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 (
|
||
<figureCitation id="1318FBE51D09FFC2FC1B94B1FBC35062" box="[953,1030,1778,1804]" captionStart="FIG" captionStartId="52.[132,143,1963,1980]" captionTargetBox="[129,1457,198,1873]" captionTargetId="figure-17@52.[129,1486,205,1971]" captionTargetPageId="52" captionText="FIG. 24. — Pelvic girdle elements of ‘Metriorhynchus’ brachyrhynchus (Eudes-Deslongchamps,1867), NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens Andrews, 1913): A, right ilium in lateral view; B, right ilium in medial view; C, right pubis in anteroposterior;D, left ischium in lateral view. Target indicates anterior. Arrow points anteriorly. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106650" httpUri="https://zenodo.org/record/11106650/files/figure.png" pageId="53" pageNumber="186">Fig. 24</figureCitation>
|
||
) possesses a short and thick shaft similar to
|
||
<taxonomicName id="4C239CE31D09FFC2FCCF9551FB5F5042" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[877,1178,1810,1836]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FCCF9551FB5F5042" box="[877,1178,1810,1836]" italics="true" pageId="53" pageNumber="186">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C239CE31D09FFC2FB729551FA6A5045" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[1232,1455,1810,1836]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B9573B721D09FFC2FB729551FA6A5045" box="[1232,1455,1810,1836]" italics="true" pageId="53" pageNumber="186">Dakosaurus maximus</emphasis>
|
||
</taxonomicName>
|
||
among metriorhynchoids. Indeed, the shaft of the ischium of ‘
|
||
<taxonomicName id="4C239CE31D09FFC2FCF39512FA9D5002" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[849,1368,1873,1900]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D09FFC2FCF39512FC375005" box="[849,1010,1873,1899]" italics="true" pageId="53" pageNumber="186">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D09FFC2FBA29512FB5B5005" box="[1024,1182,1873,1899]" italics="true" pageId="53" pageNumber="186">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 is 1/3 longer anteroposteriorly as it is tall proximodistally. Still, the minimum thickness of the shaft (defining the extension of the shaft) is not drastically greater than the largest anteroposterior length of the posterior peduncle like
|
||
<taxonomicName id="4C239CE31D09FFC0FAB79591FECE579F" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="55" lastPageNumber="188" order="Crocodylia" pageId="53" pageNumber="186" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D09FFC2FAB79591FA685085" box="[1301,1453,2002,2027]" italics="true" pageId="53" pageNumber="186">Tyrannoneustes</emphasis>
|
||
<emphasis id="B9573B721D0BFFC0FF269294FECE579F" box="[132,267,215,241]" italics="true" pageId="55" pageNumber="188">lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C239CE31D0BFFC0FEE39294FDDC579F" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" box="[321,537,215,241]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B9573B721D0BFFC0FEE39294FDDC579F" box="[321,537,215,241]" italics="true" pageId="55" pageNumber="188">Dakosaurus maximus</emphasis>
|
||
</taxonomicName>
|
||
to a lesser extent. As in other metriorhynchoids, the acetabular perforation is strongly reduced on the ischium of ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FE159354FEFD563F" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FE159354FD9C565F" box="[439,601,279,305]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FDCA9354FCC2565F" box="[616,775,279,305]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804, and corresponds to a shallow notch between both peduncle which then transitions to a shallow groove on the medial margin of the bone (e.g.
|
||
<taxonomicName id="4C239CE31D0BFFC0FDCD9334FEC556DE" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0BFFC0FDCD9334FEC556DE" italics="true" pageId="55" pageNumber="188">Thalattosuchus superciliosus</emphasis>
|
||
</taxonomicName>
|
||
, ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FEB593D5FF5D56BF" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FEB593D5FE7D56DE" box="[279,440,406,432]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FE6593D5FDA056DE" box="[455,613,406,432]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763,
|
||
<taxonomicName id="4C239CE31D0BFFC0FF4993F4FDED56BE" authority="YOUNG, DE ANDRADE, BRUSATTE, SAKAMOTO & LISTON, 2012" authorityName="Young, Andrade, Brusatte, Sakamoto & Liston" authorityYear="2012" box="[235,552,438,464]" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="B9573B721D0BFFC0FF4993F4FDED56BE" box="[235,552,438,464]" italics="true" pageId="55" pageNumber="188">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
, etc.). The anterior margin of the ischium underneath the peduncles is strongly concave even semicircular (greater than in ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FDFD93B5FE1C555F" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FDFD93B5FCC5557E" box="[607,768,502,528]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FF269055FEE7555E" box="[132,290,534,560]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763). The anterior margin culminates ventrally to form the anterior process of the distal blade, which is sharp as in several metriorhynchoids (i.e. ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FF299035FD5255FE" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[139,663,630,656]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FF299035FEE955FE" box="[139,300,630,656]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FE999035FE1C55FE" box="[315,473,630,656]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
4763 and LEICT G.418.1956.13.5,
|
||
<taxonomicName id="4C239CE31D0BFFC0FE1290D5FDBA55C1" authorityName="NHMUK PV R" authorityYear="1230" box="[432,639,662,688]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis id="B9573B721D0BFFC0FE1290D5FDBA55C1" box="[432,639,662,688]" italics="true" pageId="55" pageNumber="188">Geosaurus giganteus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FD2F90D5FF2755A1" authorityName="SMNS" baseAuthorityName="Plieninger" baseAuthorityYear="1846" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="maximus">
|
||
<emphasis id="B9573B721D0BFFC0FD2F90D5FF2755A1" italics="true" pageId="55" pageNumber="188">Dakosaurus maximus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C239CE31D0BFFC0FEBD90F5FDC855A1" baseAuthorityName="Wilkinson, Young & Benton" baseAuthorityYear="2008" box="[287,525,694,719]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="B9573B721D0BFFC0FEBD90F5FDC855A1" box="[287,525,694,719]" italics="true" pageId="55" pageNumber="188">Torvoneustes carpenteri</emphasis>
|
||
</taxonomicName>
|
||
). The posterior margin of the ischium of ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FEE59096FF05547E" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FEE59096FE2D5581" box="[327,488,725,751]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FE579096FD565581" box="[501,659,725,751]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 appears slightly concave as opposed to its anterior margin.
|
||
</paragraph>
|
||
<caption id="DF5CB7E81D0AFFC1FF26953EFED25087" ID-DOI="http://doi.org/10.5281/zenodo.11106652" ID-Zenodo-Dep="11106652" httpUri="https://zenodo.org/record/11106652/files/figure.png" pageId="54" pageNumber="187" startId="54.[132,143,1917,1934]" targetBox="[147,1456,184,1888]" targetPageId="54" targetType="figure">
|
||
<paragraph id="8B9CE7601D0AFFC1FF26953EFED25087" blockId="54.[132,1457,1917,2025]" pageId="54" pageNumber="187">
|
||
<smallCapsWord id="8D7A71BC1D0AFFC1FF26953EFF5850E0" baselines="1930,1931" box="[132,157,1917,1934]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="54" pageNumber="187">FIG</smallCapsWord>
|
||
. 25. — Reconstruction of the pelvic girdle of ‘
|
||
<taxonomicName id="4C239CE31D0AFFC1FDBC953EFBF250E0" authority="(Eudes-Deslongchamps, 1867)" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[542,1079,1917,1934]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="54" pageNumber="187" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0AFFC1FDBC953EFD5A50E0" box="[542,671,1917,1934]" italics="true" pageId="54" pageNumber="187">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0AFFC1FD0B953EFCE850E0" box="[681,813,1917,1934]" italics="true" pageId="54" pageNumber="187">brachyrhynchus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB29A911D0AFFC1FC95953EFBF750E0" author="EUDES-DESLONGCHAMPS J. - A." box="[823,1074,1917,1934]" pageId="54" pageNumber="187" refId="ref110195" refString="EUDES-DESLONGCHAMPS J. - A. 1867. - Note sur les teleosauriens [posthumous work]. Le Blanc-Hardel, Caen, 34 p." type="book" year="1867">Eudes-Deslongchamps, 1867</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMUK PV R 4763 and NHMUK PV R 3804 (holotype of ‘
|
||
<taxonomicName id="4C239CE31D0AFFC1FF5295D7FEB450CB" box="[240,369,1940,1957]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="54" pageNumber="187" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9573B721D0AFFC1FF5295D7FEB450CB" box="[240,369,1940,1957]" italics="true" pageId="54" pageNumber="187">Metriorhynchus</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B9573B721D0AFFC1FED995D7FE0950CB" box="[379,460,1940,1957]" italics="true" pageId="54" pageNumber="187">cultridens</emphasis>
|
||
):
|
||
<emphasis id="B9573B721D0AFFC1FE7995D7FE2D50CB" bold="true" box="[475,488,1940,1957]" pageId="54" pageNumber="187">A</emphasis>
|
||
, NHMUK PV R 3804, right lateral view;
|
||
<emphasis id="B9573B721D0AFFC1FC8E95D7FCFC50CB" bold="true" box="[812,825,1940,1957]" pageId="54" pageNumber="187">B</emphasis>
|
||
, NHMUK PV R 4763, right lateral view;
|
||
<emphasis id="B9573B721D0AFFC1FBDC95D7FB4950CB" bold="true" box="[1150,1164,1940,1957]" pageId="54" pageNumber="187">C</emphasis>
|
||
, NHMUK PV R 3804, anterior view;
|
||
<emphasis id="B9573B721D0AFFC1FF2695E8FF5750D2" bold="true" box="[132,146,1963,1980]" pageId="54" pageNumber="187">D</emphasis>
|
||
, NHMUK PV R 4763, anterior view;
|
||
<emphasis id="B9573B721D0AFFC1FE1F95E8FE0C50D2" bold="true" box="[445,457,1963,1980]" pageId="54" pageNumber="187">E</emphasis>
|
||
, NHMUK PV R 3804, ventral view;
|
||
<emphasis id="B9573B721D0AFFC1FD4E95E8FD3250D2" bold="true" box="[748,759,1963,1980]" pageId="54" pageNumber="187">F</emphasis>
|
||
, NHMUK PV R 4763, ventral view. Target indicates anterior. Arrow points anteriorly. The right ischium and left ilium and pubis of NHMUK PV R 3804 are mirrored. The sacral ribs of NHMUK PV R 4763 have been modified to contact the ilium. Scale bars: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9CE7601D0BFFC0FF269116FF785401" blockId="55.[132,189,853,879]" box="[132,189,853,879]" pageId="55" pageNumber="188">
|
||
<heading id="D0D4500C1D0BFFC0FF269116FF785401" box="[132,189,853,879]" fontSize="11" level="2" pageId="55" pageNumber="188" reason="8">
|
||
<emphasis id="B9573B721D0BFFC0FF269116FF785401" box="[132,189,853,879]" italics="true" pageId="55" pageNumber="188">Pubis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D0BFFC0FF269136FDB45102" blockId="55.[130,777,885,2028]" pageId="55" pageNumber="188">
|
||
The pubis of ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FEB09136FCC054E1" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[274,773,885,911]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FEB09136FE7454E1" box="[274,433,885,911]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FE1E9136FD9D54E1" box="[444,600,885,911]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 (
|
||
<figureCitation id="1318FBE51D0BFFC0FF6891D6FED654C1" box="[202,275,917,943]" captionStart="FIG" captionStartId="52.[132,143,1963,1980]" captionTargetBox="[129,1457,198,1873]" captionTargetId="figure-17@52.[129,1486,205,1971]" captionTargetPageId="52" captionText="FIG. 24. — Pelvic girdle elements of ‘Metriorhynchus’ brachyrhynchus (Eudes-Deslongchamps,1867), NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens Andrews, 1913): A, right ilium in lateral view; B, right ilium in medial view; C, right pubis in anteroposterior;D, left ischium in lateral view. Target indicates anterior. Arrow points anteriorly. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106650" httpUri="https://zenodo.org/record/11106650/files/figure.png" pageId="55" pageNumber="188">Fig. 24</figureCitation>
|
||
) displays a well-developed pubic symphysis like other thalattosuchians. The proximal peduncle of the bone is oval in proximal view, but its anterior margin shows a deep extension towards the shaft (vs
|
||
<taxonomicName id="4C239CE31D0BFFC0FE6591B7FD6A5360" authorityName="NHMUK PV R" authorityYear="2618" box="[455,687,1012,1038]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D0BFFC0FE6591B7FD6A5360" box="[455,687,1012,1038]" italics="true" pageId="55" pageNumber="188">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FD1991B7FD905341" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0BFFC0FD1991B7FE985340" italics="true" pageId="55" pageNumber="188">Thalattosuchus superciliosus</emphasis>
|
||
NHMUK PV R 2054
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FDC19657FF1D5320" authority="(Jager, 1828)" authorityName="(Jager)" baseAuthorityName="Jager" baseAuthorityYear="1828" class="Reptilia" family="Teleosauridae" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="bollensis">
|
||
<emphasis id="B9573B721D0BFFC0FDC19657FF1D5320" italics="true" pageId="55" pageNumber="188">Macrospondylus bollensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FF469677FE225320" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" box="[228,487,1076,1102]" genus="Charitomenosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="leedsi">
|
||
<emphasis id="B9573B721D0BFFC0FF469677FE225320" box="[228,487,1076,1102]" italics="true" pageId="55" pageNumber="188">Charitomenosuchus leedsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FE519676FD325320" authority="(Eudes-Deslongchamps, 1868)" authorityName="(Eudes-Deslongchamps)" authorityYear="1868" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1868" box="[499,759,1076,1102]" genus="Neosteneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="edwardsi">
|
||
<emphasis id="B9573B721D0BFFC0FE519676FD325320" box="[499,759,1076,1102]" italics="true" pageId="55" pageNumber="188">Neosteneosaurus edwardsi</emphasis>
|
||
</taxonomicName>
|
||
). Also, the greatest axis of the pubic peduncle is not in line with that of the section of the shaft but is tilted at about 30° (
|
||
<figureCitation id="1318FBE51D0BFFC0FF2F96D7FF1C53C1" box="[141,217,1172,1199]" captionStart="FIG" captionStartId="54.[132,143,1917,1934]" captionTargetBox="[147,1456,184,1888]" captionTargetId="figure-17@54.[165,1419,200,1875]" captionTargetPageId="54" captionText="FIG. 25. — Reconstruction of the pelvic girdle of ‘Metriorhynchus’ brachyrhynchus (Eudes-Deslongchamps, 1867), NHMUK PV R 4763 and NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens): A, NHMUK PV R 3804, right lateral view; B, NHMUK PV R 4763, right lateral view; C, NHMUK PV R 3804, anterior view; D, NHMUK PV R 4763, anterior view; E, NHMUK PV R 3804, ventral view; F, NHMUK PV R 4763, ventral view. Target indicates anterior. Arrow points anteriorly. The right ischium and left ilium and pubis of NHMUK PV R 3804 are mirrored. The sacral ribs of NHMUK PV R 4763 have been modified to contact the ilium. Scale bars: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.11106652" httpUri="https://zenodo.org/record/11106652/files/figure.png" pageId="55" pageNumber="188">Fig. 25</figureCitation>
|
||
). Overall, the pubis of ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FE7F96D7FEA353A0" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FE7F96D7FDBB53C0" box="[477,638,1172,1198]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FD2E96D7FF7553A0" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 takes the shape of a palette knife with a rod-like shaft and a distal blade bearing a straight surface medially (i.e. the large pubic symphysis;
|
||
<figureCitation id="1318FBE51D0BFFC0FDCD96B0FD7E5260" box="[623,699,1267,1294]" captionStart="FIG" captionStartId="60.[132,143,1963,1980]" captionTargetBox="[136,1456,193,1896]" captionTargetId="figure-17@60.[136,1371,245,1867]" captionTargetPageId="60" captionText="FIG. 28. — Pelvic girdle elements of Suchodus durobrivensis Lydekker, 1890, NHMUK PV R 2618: A, right ilium in lateral view; B, right ilium in medial view; C, right pubis in anterodorsal view; D, right pubis in lateral view; E, left pubis in posteroventral view. Target indicates anterior. Cross indicates posterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106658" httpUri="https://zenodo.org/record/11106658/files/figure.png" pageId="55" pageNumber="188">Fig. 28</figureCitation>
|
||
) and a rounded one laterally. Thereby, it differs from that of extant crocodylians which show an almost isosceles triangular distal blade. Indeed, in
|
||
<taxonomicName id="4C239CE31D0BFFC0FE989717FE3D5203" box="[314,504,1363,1390]" class="Crocodylia" family="Alligatoridae" genus="Caiman" kingdom="Animalia" order="Eusuchia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="crocodilus">
|
||
<emphasis id="B9573B721D0BFFC0FE989717FE3D5203" box="[314,504,1363,1390]" italics="true" pageId="55" pageNumber="188">Caiman crocodilus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1318FBE51D0BFFC0FDA59710FD865203" box="[519,579,1363,1389]" captionStart="FIG" captionStartId="21.[132,143,1986,2003]" captionTargetBox="[132,1441,166,1905]" captionTargetId="figure-15@21.[-252,1479,167,1865]" captionTargetPageId="21" captionText="FIG. 9. — Left pelvic girdle of Caiman crocodilus (Linnaeus, 1758), NHMW 30900: A, ilium in medial view; B, ilium in lateral view; C, ischium in medial view; D, ischium in lateral view; E, pubis in posterior view. Target indicates anterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106620" httpUri="https://zenodo.org/record/11106620/files/figure.png" pageId="55" pageNumber="188">Fig. 9</figureCitation>
|
||
) or
|
||
<taxonomicName id="4C239CE31D0BFFC0FDCE9717FF1F52E3" class="Crocodylia" family="Crocodylidae" genus="Mecistops" kingdom="Animalia" order="Eusuchia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="cataphractus">
|
||
<emphasis id="B9573B721D0BFFC0FDCE9717FF1F52E3" italics="true" pageId="55" pageNumber="188">Mecistops cataphractus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1318FBE51D0BFFC0FF499730FEEF52E3" box="[235,298,1395,1421]" captionStart="FIG" captionStartId="20.[132,143,1986,2003]" captionTargetBox="[149,1433,219,1874]" captionTargetId="figure-17@20.[226,1462,231,1900]" captionTargetPageId="20" captionText="FIG. 8. — Right pelvic girdle of Mecistops cataphractus Cuvier, 1825, RBINS 18374: A, ilium in lateral view; B, ischium in lateral view; C, pubis in ventral view; D, ilium in medial view. Cross indicates posterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106616" httpUri="https://zenodo.org/record/11106616/files/figure.png" pageId="55" pageNumber="188">Fig. 8</figureCitation>
|
||
), the lateral and medial margins are almost symmetrical thus displaying a reduced pubic symphysis. The dyrosaurids
|
||
<taxonomicName id="4C239CE31D0BFFC0FEA597F7FE1452A3" box="[263,465,1460,1485]" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="natator">
|
||
<emphasis id="B9573B721D0BFFC0FEA597F7FE1452A3" box="[263,465,1460,1485]" italics="true" pageId="55" pageNumber="188">Hyposaurus natator</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C239CE31D0BFFC0FDA497F7FCC352A3" box="[518,774,1459,1485]" class="Reptilia" family="Dyrosauridae" genus="Dyrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="maghribensis">
|
||
<emphasis id="B9573B721D0BFFC0FDA497F7FCC352A3" box="[518,774,1459,1485]" italics="true" pageId="55" pageNumber="188">Dyrosaurus maghribensis</emphasis>
|
||
</taxonomicName>
|
||
(OCP DEK-GE 252 and OCP DEK-GE 255) further differ from ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FF6197B0FD745163" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[195,689,1523,1549]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FF6197B0FEA55163" box="[195,352,1523,1549]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FEC997B0FDC05163" box="[363,517,1523,1549]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 as they also possess a rather symmetrical isosceles triangular distal blade, but this is not the case for
|
||
<taxonomicName id="4C239CE31D0BFFC0FE4D9471FCCD5122" box="[495,776,1586,1612]" family="Dyrosauridae" genus="Cerrejonisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="improcerus">
|
||
<emphasis id="B9573B721D0BFFC0FE4D9471FCCD5122" box="[495,776,1586,1612]" italics="true" pageId="55" pageNumber="188">Cerrejonisuchus improcerus</emphasis>
|
||
</taxonomicName>
|
||
which displays a strongly asymmetrical pubis.
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D0BFFC0FF399431FA6A555E" blockId="55.[130,777,885,2028]" lastBlockId="55.[810,1457,215,943]" pageId="55" pageNumber="188">
|
||
Comparatively, the pubes of ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FE7E9431FEAB51C3" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FE7E9431FDB851E2" box="[476,637,1650,1676]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FD2E9431FF7551C2" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 and
|
||
<taxonomicName id="4C239CE31D0BFFC0FE4E94D1FCCC51C2" box="[492,777,1682,1708]" family="Dyrosauridae" genus="Cerrejonisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="improcerus">
|
||
<emphasis id="B9573B721D0BFFC0FE4E94D1FCCC51C2" box="[492,777,1682,1708]" italics="true" pageId="55" pageNumber="188">Cerrejonisuchus improcerus</emphasis>
|
||
</taxonomicName>
|
||
may appear similar with a short and concave medial margin of the shaft leading to a straight portion of the distal blade, and a more elongated lateral margin of the shaft ending with a rounded portion of the distal blade (the posterior protuberance). In both cases, the medial margin of the shaft and the distal blade meet at an angle slightly greater than 90°. Yet, the main difference resides in the junction between the lateral margins of the shaft and the distal blade: while those are aligned in ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FF5D95F2FCCD50A2" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[255,776,1969,1996]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FF5D95F2FE6550A5" box="[255,416,1969,1995]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FE0D95F2FD8850A5" box="[431,589,1969,1995]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804, in
|
||
<taxonomicName id="4C239CE31D0BFFC0FF419592FE3E5085" box="[227,507,2001,2027]" family="Dyrosauridae" genus="Cerrejonisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="improcerus">
|
||
<emphasis id="B9573B721D0BFFC0FF419592FE3E5085" box="[227,507,2001,2027]" italics="true" pageId="55" pageNumber="188">Cerrejonisuchus improcerus</emphasis>
|
||
</taxonomicName>
|
||
they form an angle lesser than 180°. The overall flat lateral margin of ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FAA59294FB4C567C" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FAA59294FA6D579F" box="[1287,1448,215,241]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FC8F92B4FC09567F" box="[813,972,247,273]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804 along with its larger distal blade, creates a smooth and gradual transition between the shaft and the distal blade. On the contrary, in extant crocodylians (e.g.
|
||
<taxonomicName id="4C239CE31D0BFFC0FC48931BFB0A561F" box="[1002,1231,343,369]" class="Crocodylia" family="Crocodylidae" genus="Mecistops" kingdom="Animalia" order="Eusuchia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="cataphractus">
|
||
<emphasis id="B9573B721D0BFFC0FC48931BFB0A561F" box="[1002,1231,343,369]" italics="true" pageId="55" pageNumber="188">Mecistops cataphractus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="1318FBE51D0BFFC0FB7E9314FADD561F" box="[1244,1304,343,369]" captionStart="FIG" captionStartId="20.[132,143,1986,2003]" captionTargetBox="[149,1433,219,1874]" captionTargetId="figure-17@20.[226,1462,231,1900]" captionTargetPageId="20" captionText="FIG. 8. — Right pelvic girdle of Mecistops cataphractus Cuvier, 1825, RBINS 18374: A, ilium in lateral view; B, ischium in lateral view; C, pubis in ventral view; D, ilium in medial view. Cross indicates posterior. Arrow points anteriorly. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.11106616" httpUri="https://zenodo.org/record/11106616/files/figure.png" pageId="55" pageNumber="188">Fig. 8</figureCitation>
|
||
]) and in other thalattosuchians the shaft of the pubis flares out both laterally and medially to form the distal blade (e.g.
|
||
<taxonomicName id="4C239CE31D0BFFC0FB4F93D5FA6D56DE" authority="Bronn, 1841" box="[1261,1448,406,432]" class="Reptilia" family="Metriorhynchidae" genus="Pelagosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species">
|
||
<emphasis id="B9573B721D0BFFC0FB4F93D5FA6D56DE" box="[1261,1448,406,432]" italics="true" pageId="55" pageNumber="188">Pelagosaurus typus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FC8F93F5FBDD56BE" authorityName="NHMUK PV R" authorityYear="2618" box="[813,1048,438,464]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D0BFFC0FC8F93F5FBDD56BE" box="[813,1048,438,464]" italics="true" pageId="55" pageNumber="188">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FB8693F5FC6C569F" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0BFFC0FB8693F5FAF856BE" box="[1060,1341,438,464]" italics="true" pageId="55" pageNumber="188">Thalattosuchus superciliosus</emphasis>
|
||
NHMUK PV R 2054
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FC1B9394FB4F569E" baseAuthorityName="Fraas" baseAuthorityYear="1901" box="[953,1162,471,497]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="suevicus">
|
||
<emphasis id="B9573B721D0BFFC0FC1B9394FB4F569E" box="[953,1162,471,497]" italics="true" pageId="55" pageNumber="188">Cricosaurus suevicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FB389394FAAC569E" authorityName="NHMUK PV R" authorityYear="1230" box="[1178,1385,471,497]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis id="B9573B721D0BFFC0FB389394FAAC569E" box="[1178,1385,471,497]" italics="true" pageId="55" pageNumber="188">Geosaurus giganteus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FADB9394FC3D557E" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" class="Reptilia" family="Teleosauridae" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="obtusidens">
|
||
<emphasis id="B9573B721D0BFFC0FADB9394FC3D557E" italics="true" pageId="55" pageNumber="188">Lemmysuchus obtusidens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FBA793B5FB3A557E" authority="(Jager, 1828)" authorityName="(Jager)" baseAuthorityName="Jager" baseAuthorityYear="1828" box="[1029,1279,502,528]" class="Reptilia" family="Teleosauridae" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="bollensis">
|
||
<emphasis id="B9573B721D0BFFC0FBA793B5FB3A557E" box="[1029,1279,502,528]" italics="true" pageId="55" pageNumber="188">Macrospondylus bollensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FAA993B5FC53555E" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" genus="Charitomenosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="leedsi">
|
||
<emphasis id="B9573B721D0BFFC0FAA993B5FC53555E" italics="true" pageId="55" pageNumber="188">Charitomenosuchus leedsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FC069054FB6E555E" authority="(Eudes-Deslongchamps, 1868)" authorityName="(Eudes-Deslongchamps)" authorityYear="1868" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1868" box="[932,1195,534,560]" genus="Neosteneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="edwardsi">
|
||
<emphasis id="B9573B721D0BFFC0FC069054FB6E555E" box="[932,1195,534,560]" italics="true" pageId="55" pageNumber="188">Neosteneosaurus edwardsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FB1B9055FA5A555E" authority="(Andrews, 1909)" authorityName="(Andrews)" authorityYear="1909" baseAuthorityName="Andrews" baseAuthorityYear="1909" box="[1209,1439,534,560]" family="Teleosauridae" genus="Mycterosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9573B721D0BFFC0FB1B9055FA5A555E" box="[1209,1439,534,560]" italics="true" pageId="55" pageNumber="188">Mycterosuchus nasutus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9CE7601D0BFFC0FCE19074FC6B54C1" blockId="55.[810,1457,215,943]" pageId="55" pageNumber="188">
|
||
In ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FCCA9075FAAC553F" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" box="[872,1385,566,593]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FCCA9075FBCC553E" box="[872,1033,566,592]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FBB49075FB71553E" box="[1046,1204,566,592]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804, the pubic blade displays a posterior protuberance similar to what is observed in dyrosaurids (i.e.
|
||
<taxonomicName id="4C239CE31D0BFFC0FB129034FABF55FE" box="[1200,1402,631,656]" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="natator">
|
||
<emphasis id="B9573B721D0BFFC0FB129034FABF55FE" box="[1200,1402,631,656]" italics="true" pageId="55" pageNumber="188">Hyposaurus natator</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C239CE31D0BFFC0FC8E90D5FB0455DE" authority="OCP DEK-GE" authorityName="OCP DEK-GE" box="[812,1217,662,688]" class="Reptilia" family="Dyrosauridae" genus="Dyrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="maghribensis">
|
||
<emphasis id="B9573B721D0BFFC0FC8E90D5FBDB55DE" box="[812,1054,662,688]" italics="true" pageId="55" pageNumber="188">Dyrosaurus maghribensis</emphasis>
|
||
OCP DEK-GE
|
||
</taxonomicName>
|
||
252 & OCP DEK-GE 255), but unlike most thalattosuchians (e.g.
|
||
<taxonomicName id="4C239CE31D0BFFC0FB5190F6FA6D55A1" authority="Bronn, 1841" box="[1267,1448,693,719]" class="Reptilia" family="Metriorhynchidae" genus="Pelagosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species">
|
||
<emphasis id="B9573B721D0BFFC0FB5190F6FA6D55A1" box="[1267,1448,693,719]" italics="true" pageId="55" pageNumber="188">Pelagosaurus typus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FC8F9096FBDD5581" authorityName="NHMUK PV R" authorityYear="2618" box="[813,1048,725,751]" class="Reptilia" family="Metriorhynchidae" genus="Suchodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="durobrivensis">
|
||
<emphasis id="B9573B721D0BFFC0FC8F9096FBDD5581" box="[813,1048,725,751]" italics="true" pageId="55" pageNumber="188">Suchodus durobrivensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FB869096FC67547E" authority="(Blainville, 1853)" authorityName="(Blainville)" authorityYear="1852" baseAuthorityName="Blainville" baseAuthorityYear="1853" class="Reptilia" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="superciliosus">
|
||
<emphasis id="B9573B721D0BFFC0FB869096FAF85581" box="[1060,1341,725,751]" italics="true" pageId="55" pageNumber="188">Thalattosuchus superciliosus</emphasis>
|
||
NHMUK PV R 2054
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FC0C90B5FBBF5461" baseAuthorityName="Fraas" baseAuthorityYear="1901" box="[942,1146,758,784]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="suevicus">
|
||
<emphasis id="B9573B721D0BFFC0FC0C90B5FBBF5461" box="[942,1146,758,784]" italics="true" pageId="55" pageNumber="188">Cricosaurus suevicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FB2490B5FA8A5461" authorityName="NHMUK PV R" authorityYear="1230" box="[1158,1359,758,784]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis id="B9573B721D0BFFC0FB2490B5FA8A5461" box="[1158,1359,758,784]" italics="true" pageId="55" pageNumber="188">Geosaurus giganteus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FAF990B5FC1F5441" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" class="Reptilia" family="Teleosauridae" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="obtusidens">
|
||
<emphasis id="B9573B721D0BFFC0FAF990B5FC1F5441" italics="true" pageId="55" pageNumber="188">Lemmysuchus obtusidens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FC479156FB185441" authority="(Jager, 1828)" authorityName="(Jager)" baseAuthorityName="Jager" baseAuthorityYear="1828" box="[997,1245,789,815]" class="Reptilia" family="Teleosauridae" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="bollensis">
|
||
<emphasis id="B9573B721D0BFFC0FC479156FB185441" box="[997,1245,789,815]" italics="true" pageId="55" pageNumber="188">Macrospondylus bollensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FB4B9156FCA45421" authority="(Andrews, 1909)" authorityName="(Andrews)" baseAuthorityName="Andrews" baseAuthorityYear="1909" genus="Charitomenosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="leedsi">
|
||
<emphasis id="B9573B721D0BFFC0FB4B9156FCA45421" italics="true" pageId="55" pageNumber="188">Charitomenosuchus leedsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C239CE31D0BFFC0FCCF9175FBAC5421" authority="(Eudes-Deslongchamps, 1868)" authorityName="(Eudes-Deslongchamps)" authorityYear="1868" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1868" box="[877,1129,821,847]" genus="Neosteneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="edwardsi">
|
||
<emphasis id="B9573B721D0BFFC0FCCF9175FBAC5421" box="[877,1129,821,847]" italics="true" pageId="55" pageNumber="188">Neosteneosaurus edwardsi</emphasis>
|
||
</taxonomicName>
|
||
). Comparatively,
|
||
<taxonomicName id="4C239CE31D0BFFC0FABD9176FCBF5401" authority="(Andrews, 1909)" authorityName="(Andrews)" authorityYear="1909" baseAuthorityName="Andrews" baseAuthorityYear="1909" family="Teleosauridae" genus="Mycterosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B9573B721D0BFFC0FABD9176FCBF5401" italics="true" pageId="55" pageNumber="188">Mycterosuchus nasutus</emphasis>
|
||
</taxonomicName>
|
||
also possesses a posterior protuberance but of greater intensity than in ‘
|
||
<taxonomicName id="4C239CE31D0BFFC0FC4F9136FCAD54C1" authority="NHMUK PV R" authorityName=", NHMUK PV R" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1867" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="188" phylum="Chordata" rank="species" species="brachyrhynchus">
|
||
<emphasis id="B9573B721D0BFFC0FC4F9136FB4B54E1" box="[1005,1166,885,911]" italics="true" pageId="55" pageNumber="188">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="B9573B721D0BFFC0FB3E9136FAFF54E1" box="[1180,1338,885,911]" italics="true" pageId="55" pageNumber="188">brachyrhynchus</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3804.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |