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<document id="A44D513B6DEA1E900E9C234D55B75563" ID-CLB-Dataset="7176" ID-DOI="10.5252/zoosystema2021v43a10" ID-GBIF-Dataset="120db7ca-e885-4d5e-bc3d-83af0671f8d1" ID-ISSN="1638-9387" ID-Zenodo-Dep="4720674" ID-ZooBank="urn:lsid:zoobank.org:pub:2763DA07-4D8F-4CA2-BB63-E5BC26470296" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445312570" checkinUser="felipe" docAuthor="Drolshagen, Bastian &amp; Bäckstam, Christian M." docDate="2021" docId="0386CD35FF81FFBD7580FA4D401090B9" docLanguage="en" docName="Zoosystema.43.10.163-196.pdf.imf" docOrigin="Zoosystema 43 (10)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Diplura gymnognatha Bertkau 1880" docType="treatment" docVersion="8" lastPageNumber="190" masterDocId="FFBFB54DFF9CFFA0757CFF9F44099755" masterDocTitle="A taxonomic review of the mygalomorph spider genus Linothele Karsch, 1879 (Araneae, Dipluridae)" masterLastPageNumber="196" masterPageNumber="163" pageNumber="190" updateTime="1698937603326" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F94443534D17FC7E1F66471FCAD414AF">A taxonomic review of the mygalomorph spider genus Linothele Karsch, 1879 (Araneae, Dipluridae)</mods:title>
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<mods:namePart id="5849D5BE117FB2332196975976A69AAF">Drolshagen, Bastian</mods:namePart>
<mods:affiliation id="5AB8EF0E11B8EF7B9FA57C6D9C993D88">Kolberger Strasse 12 F, 76139 Karlsruhe (Germany) bdrolshagen @ gmail. com</mods:affiliation>
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<mods:namePart id="8F91D507E172CC11D0005A06DDCF7E0C">Bäckstam, Christian M.</mods:namePart>
<mods:affiliation id="D3DCF73941512C389DD5C065EAD9C0FC">Swedish Museum of Natural History, Department of Zoology, Box 50007, 104 05 Stockholm (Sweden) christian. backstam @ nrm. se</mods:affiliation>
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<taxonomicName id="4C2F07A0FF81FFBD7580FA4D468692B8" authority="Bertkau, 1880" authorityName="Bertkau" authorityYear="1880" box="[252,655,1490,1517]" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD7580FA4D45E592B8" bold="true" box="[252,492,1490,1517]" italics="true" pageId="29" pageNumber="190">Diplura gymnognatha</emphasis>
<bibRefCitation id="EFBE01D2FF81FFBD7488FA4C468692B8" author="Bertkau" box="[500,655,1491,1517]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau, 1880</bibRefCitation>
</taxonomicName>
comb. rev.
<emphasis id="B95BA031FF81FFBD74CDFA6C46599158" box="[433,592,1523,1549]" italics="true" pageId="29" pageNumber="190">nomen dubium</emphasis>
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<paragraph id="8B907C23FF81FFBD75F8F9AA46FF9119" blockId="29.[132,758,1589,1613]" box="[132,758,1589,1613]" pageId="29" pageNumber="190">
<treatmentCitationGroup id="AB3F5B0DFF81FFBD75F8F9AA46FF9119" box="[132,758,1589,1613]" pageId="29" pageNumber="190">
<treatmentCitation id="0A8E5A32FF81FFBD75F8F9AA46079118" author="BERTKAU P." box="[132,526,1589,1613]" page="21" pageId="29" pageNumber="190" year="1880">
<taxonomicName id="4C2F07A0FF81FFBD75F8F9AA46079118" authority="Bertkau, 1880: 21" authorityName="Bertkau" authorityPageNumber="21" authorityYear="1880" box="[132,526,1589,1613]" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD75F8F9AA455D9118" box="[132,340,1589,1613]" italics="true" pageId="29" pageNumber="190">Diplura gymnognatha</emphasis>
<bibRefCitation id="EFBE01D2FF81FFBD7426F9AA46079118" author="Bertkau" box="[346,526,1589,1613]" firstAuthor="Bertkau" page="21" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau, 1880: 21</bibRefCitation>
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, pl. 1, fig. 5. comb. rev.
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<paragraph id="8B907C23FF81FFBD75F8F9F2461291D3" blockId="29.[132,539,1645,1670]" box="[132,539,1645,1670]" pageId="29" pageNumber="190">
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<taxonomicName id="4C2F07A0FF81FFBD75F8F9F2455691D0" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[132,351,1645,1669]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD75F8F9F2455691D0" box="[132,351,1645,1669]" italics="true" pageId="29" pageNumber="190">Linothele gymnognatha</emphasis>
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<treatmentCitation id="0A8E5A32FF81FFBD7404F9F1461E91D3" author="RAVEN R. J." box="[376,535,1645,1670]" page="74" pageId="29" pageNumber="190" year="1985">
<bibRefCitation id="EFBE01D2FF81FFBD7404F9F1461E91D3" author="Raven" box="[376,535,1645,1670]" firstAuthor="Raven" page="74" pageId="29" pageNumber="190" pagination="1 - 180" refId="ref26139" refString="RAVEN R. J. 1985. - The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of natural History 182: 1 - 180." type="journal article" year="1985">Raven 1985: 74</bibRefCitation>
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.
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<paragraph id="8B907C23FF81FFBD75F8F938462D918C" blockId="29.[132,775,1702,1753]" pageId="29" pageNumber="190">
TYPE MATERIAL. —
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<typeStatus id="5494C281FF81FFBD7437F93945A491EB" box="[331,429,1702,1726]" pageId="29" pageNumber="190" type="holotype">Holotype</typeStatus>
.
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;
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;
<collectorName id="26DA19F5FF81FFBD77BFF93844D2918C" pageId="29" pageNumber="190">E. Van Beneden</collectorName>
leg; not located, presumably lost.
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</paragraph>
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<subSubSection id="C3352FA8FF81FFBD75F8F88D401090B9" pageId="29" pageNumber="190" type="discussion">
<paragraph id="8B907C23FF81FFBD75F8F88D44E1907F" blockId="29.[130,777,1810,2028]" box="[132,232,1810,1836]" pageId="29" pageNumber="190">REMARKS</paragraph>
<paragraph id="8B907C23FF81FFBD75F8F8AE405D9564" blockId="29.[130,777,1810,2028]" lastBlockId="29.[811,1457,215,2028]" pageId="29" pageNumber="190">
The original description (
<bibRefCitation id="EFBE01D2FF81FFBD74D3F8AE46469019" author="Bertkau" box="[431,591,1841,1868]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau 1880</bibRefCitation>
) mentioned 11 cheliceral teeth and 20 maxillary cuspules. The description is confusing with regards to the legs:
<bibRefCitation id="EFBE01D2FF81FFBD7496F8EE468590D9" author="Bertkau" box="[490,652,1905,1932]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
mentioned the presence of ventral spines on tarsi, pseudo-segmented metatarsi, presence of a dense scopula on anterior tarsi and metatarsi, but divided scopula on posterior metatarsi. It seems like the terms metatarsi and tarsi were mixed in the original description. Bertkau further mentioned short spinnerets. The female genitalia are illustrated (
<bibRefCitation id="EFBE01D2FF81FFBD7073FE8841A19664" author="Bertkau" box="[1295,1448,279,305]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau 1880</bibRefCitation>
: Pl. 1, fig. 5) and resemble the shape observed in
<taxonomicName id="4C2F07A0FF81FFBD7032FEA847759624" baseAuthorityName="Simon" baseAuthorityYear="1889" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="B95BA031FF81FFBD7032FEA847759624" italics="true" pageId="29" pageNumber="190">Linothele gaujoni</emphasis>
</taxonomicName>
, with long, tube-shaped stalks, as also reported for
<taxonomicName id="4C2F07A0FF81FFBD7651FEE8402896E5" authority="Guadanucci, Fonseca-Ferreira, Baptista &amp; Pedroso, 2016" authorityName="Guadanucci, Fonseca-Ferreira, Baptista &amp; Pedroso" authorityYear="2016" class="Arachnida" family="Dipluridae" genus="Trechona" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="diamantina">
<emphasis id="B95BA031FF81FFBD7651FEE8401C96C4" box="[813,1045,375,401]" italics="true" pageId="29" pageNumber="190">Trechona diamantina</emphasis>
Guadanucci, Fonseca-Ferreira, Baptista &amp;
<bibRefCitation id="EFBE01D2FF81FFBD7603FE09402896E5" author="Pedroso" box="[895,1057,406,432]" firstAuthor="Pedroso" pageId="29" pageNumber="190" pagination="1 - 21" refId="ref25712" refString="PEDROSO D. R., DE SOUZA CASTANHEIRA P. &amp; CERQUEIRA BAP- TISTA R. L. 2016. - Redescription and synonymies of Diplura macrura (C. L. Koch, 1841) and D. lineata (Lucas, 1857), with notes on the genus (Araneae, Dipluridae). European Journal of Taxonomy (210): 1 - 21. https: // doi. org / 10.5852 / ejt. 2016.210" type="journal article" year="2016">Pedroso, 2016</bibRefCitation>
</taxonomicName>
. The spermathecae of
<taxonomicName id="4C2F07A0FF81FFBD706FFE08417696E5" baseAuthorityName="Simon" baseAuthorityYear="1889" box="[1299,1407,407,432]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="B95BA031FF81FFBD706FFE08417696E5" box="[1299,1407,407,432]" italics="true" pageId="29" pageNumber="190">L. gaujoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F07A0FF81FFBD7651FE2847C39685" authorityName="Guadanucci, Fonseca-Ferreira, Baptista &amp; Pedroso" authorityYear="2016" box="[813,970,438,464]" class="Arachnida" family="Dipluridae" genus="Trechona" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="diamantina">
<emphasis id="B95BA031FF81FFBD7651FE2847C39685" box="[813,970,438,464]" italics="true" pageId="29" pageNumber="190">T. diamantina</emphasis>
</taxonomicName>
differ from the latter by the lack of lateral vesicles. It might have been hard to identify such details for Bertkau, but the
<typeStatus id="5494C281FF81FFBD7173FE6840379544" box="[1039,1086,503,529]" pageId="29" pageNumber="190">type</typeStatus>
locality is also noticeably disjunct from the one of
<taxonomicName id="4C2F07A0FF81FFBD769CFD8840449565" baseAuthorityName="Simon" baseAuthorityYear="1889" box="[992,1101,535,560]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gaujoni">
<emphasis id="B95BA031FF81FFBD769CFD8840449565" box="[992,1101,535,560]" italics="true" pageId="29" pageNumber="190">L. gaujoni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B907C23FF81FFBD763FFDA9403C95A5" blockId="29.[811,1457,215,2028]" pageId="29" pageNumber="190">
According to
<bibRefCitation id="EFBE01D2FF81FFBD76B1FDA940969504" author="Pedroso" box="[973,1183,566,593]" etAl="et al." firstAuthor="Pedroso" pageId="29" pageNumber="190" pagination="360 - 367" refId="ref25645" refString="PEDROSO D. R., BAPTISTA R. L. C. &amp; FERREIRA P. S. F. 2008. - Trechona rufa (Araneae, Dipluridae): new status, redescription and neotype designation with notes on the genus. Journal of Arachnology 36: 360 - 367. https: // doi. org / 10.1636 / CA 07 - 109.1" type="journal article" year="2008">
Pedroso
<emphasis id="B95BA031FF81FFBD715FFDA8405B9505" box="[1059,1106,566,592]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
(2008)
</bibRefCitation>
, who refer to
<bibRefCitation id="EFBE01D2FF81FFBD7054FDA841A39504" author="Kury" box="[1320,1450,566,593]" firstAuthor="Kury" pageId="29" pageNumber="190" pagination="1 - 337" refId="ref25009" refString="KURY A. B. 2003. - Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revtista Iberica de Aracnologia, vol. especial monografico 1: 1 - 337." type="journal article" year="2003">Kury (2003)</bibRefCitation>
, most of the arachnids collected during the Belgian mission to
<collectingCountry id="F3383CB3FF81FFBD7635FDE9478E95C5" box="[841,903,630,656]" name="Brazil" pageId="29" pageNumber="190">Brazil</collectingCountry>
and described by
<bibRefCitation id="EFBE01D2FF81FFBD7135FDE940E695C5" author="Bertkau" box="[1097,1263,630,656]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
were deposited at IRSNB. Unfortunately, the
<typeStatus id="5494C281FF81FFBD712AFD0940BC95E5" box="[1110,1205,662,688]" pageId="29" pageNumber="190" type="holotype">holotype</typeStatus>
could not be located in IRSNB collection (L. Baert, pers. comm.) and might have been lost during WWII.
</paragraph>
<paragraph id="8B907C23FF81FFBD763FFD6A404692D8" blockId="29.[811,1457,215,2028]" pageId="29" pageNumber="190">
<bibRefCitation id="EFBE01D2FF81FFBD763FFD6A403B9445" author="Bucherl" box="[835,1074,757,784]" etAl="et al." firstAuthor="Bucherl" pageId="29" pageNumber="190" pagination="117 - 138" refId="ref24105" refString="BUCHERL W., TIMOTHEO DA COSTA A. &amp; LUCAS S. M. 1971. - Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan 35: 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="B95BA031FF81FFBD76D8FD6947D5945A" box="[932,988,757,783]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
(1971)
</bibRefCitation>
synonymized
<taxonomicName id="4C2F07A0FF81FFBD71A4FD69417A945A" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1924" box="[1240,1395,757,783]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="borgmeyeri">
<emphasis id="B95BA031FF81FFBD71A4FD69417A945A" box="[1240,1395,757,783]" italics="true" pageId="29" pageNumber="190">D. borgmeyeri</emphasis>
</taxonomicName>
with
<taxonomicName id="4C2F07A0FF81FFBD7653FC8947D5947A" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[815,988,789,815]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD7653FC8947D5947A" box="[815,988,789,815]" italics="true" pageId="29" pageNumber="190">D. gymnognatha</emphasis>
</taxonomicName>
and provided variation data for
<taxonomicName id="4C2F07A0FF81FFBD704DFC894741941A" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1924" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="borgmeyeri">
<emphasis id="B95BA031FF81FFBD704DFC894741941A" italics="true" pageId="29" pageNumber="190">D. borgmeyeri</emphasis>
</taxonomicName>
(CT = 12-14). They further added measurements for the opisthosoma (10.5) and a PLS: 8.9 (2.5, 2.6, 3.8). It becomes clear that these measurements do not match the ones provided for
<taxonomicName id="4C2F07A0FF81FFBD76E8FC09404B94FA" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[916,1090,917,943]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD76E8FC09404B94FA" box="[916,1090,917,943]" italics="true" pageId="29" pageNumber="190">D. gymnognatha</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFBE01D2FF81FFBD7115FC0A411894FA" author="Bertkau" box="[1129,1297,917,943]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
, who reported the spinnerets (6.0) to be a lot shorter than the opisthosoma (10.0).
<bibRefCitation id="EFBE01D2FF81FFBD7603FC4B406A94BA" author="Bucherl" box="[895,1123,980,1007]" etAl="et al." firstAuthor="Bucherl" pageId="29" pageNumber="190" pagination="117 - 138" refId="ref24105" refString="BUCHERL W., TIMOTHEO DA COSTA A. &amp; LUCAS S. M. 1971. - Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan 35: 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="B95BA031FF81FFBD76A0FC4A401894BB" box="[988,1041,980,1006]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
(1971)
</bibRefCitation>
stated they examined the epigastrium of
<specimenCount id="9D29B7AAFF81FFBD76CDFC6A4015935B" box="[945,1052,1012,1039]" pageId="29" pageNumber="190" type="juvenile">3 juvenile</specimenCount>
females, of which two lacked a spermatheca (= imm. male) and the only spermatheca found was rudimentary. Yet, they considered it to equal the illustration of the spermatheca of
<taxonomicName id="4C2F07A0FF81FFBD7160FBCA40C3933B" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[1052,1226,1108,1134]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD7160FBCA40C3933B" box="[1052,1226,1108,1134]" italics="true" pageId="29" pageNumber="190">D. gymnognatha</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFBE01D2FF81FFBD7025FBCB477093DA" author="Bertkau" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
. Contradictory to
<bibRefCitation id="EFBE01D2FF81FFBD7135FBEB40FA93DA" author="Bertkau" box="[1097,1267,1140,1167]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
, who mentioned the presence of 20 maxillary cuspules,
<bibRefCitation id="EFBE01D2FF81FFBD71B2FB0B41A693FB" author="Bucherl" box="[1230,1455,1172,1198]" etAl="et al." firstAuthor="Bucherl" pageId="29" pageNumber="190" pagination="117 - 138" refId="ref24105" refString="BUCHERL W., TIMOTHEO DA COSTA A. &amp; LUCAS S. M. 1971. - Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan 35: 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="B95BA031FF81FFBD7056FB0A415793FB" box="[1322,1374,1172,1198]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
(1971)
</bibRefCitation>
noted the absence of maxillary cuspules in
<taxonomicName id="4C2F07A0FF81FFBD706DFB2A41A1939B" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1924" box="[1297,1448,1204,1230]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="borgmeyeri">
<emphasis id="B95BA031FF81FFBD706DFB2A41A1939B" box="[1297,1448,1204,1230]" italics="true" pageId="29" pageNumber="190">D. borgmeyeri</emphasis>
</taxonomicName>
. Considering the possible lapsus in the first description of
<taxonomicName id="4C2F07A0FF81FFBD7652FB6B47D49258" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[814,989,1267,1293]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD7652FB6B47D49258" box="[814,989,1267,1293]" italics="true" pageId="29" pageNumber="190">D. gymnognatha</emphasis>
</taxonomicName>
, the scopula on anterior tarsi and metatarsi was dense and possibly undivided. The fact that
<bibRefCitation id="EFBE01D2FF81FFBD702BFA8C47BC921B" author="Bucherl" etAl="et al." firstAuthor="Bucherl" pageId="29" pageNumber="190" pagination="117 - 138" refId="ref24105" refString="BUCHERL W., TIMOTHEO DA COSTA A. &amp; LUCAS S. M. 1971. - Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan 35: 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="B95BA031FF81FFBD7651FAAB476B9218" box="[813,866,1331,1357]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
(1971)
</bibRefCitation>
did not transfer
<taxonomicName id="4C2F07A0FF81FFBD7108FAAB41039218" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1924" box="[1140,1290,1331,1357]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="borgmeyeri">
<emphasis id="B95BA031FF81FFBD7108FAAB41039218" box="[1140,1290,1331,1357]" italics="true" pageId="29" pageNumber="190">D. borgmeyeri</emphasis>
</taxonomicName>
to
<taxonomicName id="4C2F07A0FF81FFBD704EFAAC41839218" authorityName="Simon" authorityYear="1889" box="[1330,1418,1331,1357]" class="Arachnida" family="Dipluridae" genus="Uruchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF81FFBD704EFAAC41839218" box="[1330,1418,1331,1357]" italics="true" pageId="29" pageNumber="190">Uruchus</emphasis>
</taxonomicName>
, as they did with
<taxonomicName id="4C2F07A0FF81FFBD76B5FACB402E9238" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1926" box="[969,1063,1363,1389]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B95BA031FF81FFBD76B5FACB402E9238" box="[969,1063,1363,1389]" italics="true" pageId="29" pageNumber="190">D. fallax</emphasis>
</taxonomicName>
, indicates the scopula on all tarsi of
<taxonomicName id="4C2F07A0FF81FFBD7651FAEB47C892D8" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1924" box="[813,961,1395,1421]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="borgmeyeri">
<emphasis id="B95BA031FF81FFBD7651FAEB47C892D8" box="[813,961,1395,1421]" italics="true" pageId="29" pageNumber="190">D. borgmeyeri</emphasis>
</taxonomicName>
was divided.
</paragraph>
<paragraph id="8B907C23FF81FFBD763FFA0C47819179" blockId="29.[811,1457,215,2028]" pageId="29" pageNumber="190">
As the
<typeStatus id="5494C281FF81FFBD76F6FA0C47E192F8" box="[906,1000,1427,1453]" pageId="29" pageNumber="190" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2F07A0FF81FFBD7174FA0B40BA92F8" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[1032,1203,1427,1453]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD7174FA0B40BA92F8" box="[1032,1203,1427,1453]" italics="true" pageId="29" pageNumber="190">D. gymnognatha</emphasis>
</taxonomicName>
is presumably lost since WWII, Bücherl
<emphasis id="B95BA031FF81FFBD769EFA2B40119298" box="[994,1048,1459,1485]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
certainly did not examine it and the specimens they examined were not conspecific. As a result, the synonymy established by
<bibRefCitation id="EFBE01D2FF81FFBD7117FA6C41459158" author="Bucherl" box="[1131,1356,1523,1549]" etAl="et al." firstAuthor="Bucherl" pageId="29" pageNumber="190" pagination="117 - 138" refId="ref24105" refString="BUCHERL W., TIMOTHEO DA COSTA A. &amp; LUCAS S. M. 1971. - Revisao de alguns tipos de aranhas caranguejeiras (Orthognatha) estabelecidos por Candido de Mello-Leitao e depositados no Museu Nacional do Rio. Memorias do Instituto Butantan 35: 117 - 138." type="journal article" year="1971">
Bücherl
<emphasis id="B95BA031FF81FFBD71BBFA6C40F29158" box="[1223,1275,1523,1549]" italics="true" pageId="29" pageNumber="190">et al.</emphasis>
(1971)
</bibRefCitation>
is herein rejected.
</paragraph>
<paragraph id="8B907C23FF81FFBD763FF9AD400F9079" blockId="29.[811,1457,215,2028]" pageId="29" pageNumber="190">
<bibRefCitation id="EFBE01D2FF81FFBD763FF9AD47EF9118" author="Bertkau" box="[835,998,1586,1613]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
neither mentioned the presence, or absence of a maxillary lyra for
<taxonomicName id="4C2F07A0FF81FFBD715AF9CC40DE9139" baseAuthorityName="Bertkau" baseAuthorityYear="1880" box="[1062,1239,1618,1644]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD715AF9CC40DE9139" box="[1062,1239,1618,1644]" italics="true" pageId="29" pageNumber="190">D. gymnognatha</emphasis>
</taxonomicName>
. In the same work,
<bibRefCitation id="EFBE01D2FF81FFBD7652F9ED47DA91D8" author="Bertkau" box="[814,979,1650,1677]" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau (1880)</bibRefCitation>
proposed
<taxonomicName id="4C2F07A0FF81FFBD713DF9ED40B791D9" authorityName="Bertkau" authorityYear="1880" box="[1089,1214,1650,1676]" class="Arachnida" family="Dipluridae" genus="Thalerothele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF81FFBD713DF9ED40B791D9" box="[1089,1214,1650,1676]" italics="true" pageId="29" pageNumber="190">Thalerothele</emphasis>
</taxonomicName>
for
<taxonomicName id="4C2F07A0FF81FFBD7194F9EC476C91F9" authority="Bertkau, 1880" authorityName="Bertkau" authorityYear="1880" class="Arachnida" family="Dipluridae" genus="Thalerothele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95BA031FF81FFBD7194F9EC414691D9" box="[1256,1359,1650,1676]" italics="true" pageId="29" pageNumber="190">T. fasciata</emphasis>
<bibRefCitation id="EFBE01D2FF81FFBD7029F9ED476C91F9" author="Bertkau" firstAuthor="Bertkau" pageId="29" pageNumber="190" pagination="1 - 120" refId="ref23904" refString="BERTKAU P. 1880. - Verzeichniss der von Prof. Ed. van Beneden auf seiner im Auftrage der Belgischen Regierung unternommen wissenschaftlichen Reise nach Brasilien und La Plata im Jahren 1872 - 73 gensammelten Arachniden. Memoires couronnes et memoires des savants etrangers / publies par l'Academie royale des sciences, des lettres et des beaux-arts de Belgique 43: 1 - 120." type="journal article" year="1880">Bertkau, 1880</bibRefCitation>
</taxonomicName>
without mentioning a lyra.
<taxonomicName id="4C2F07A0FF81FFBD71E7F90D411191F9" authorityName="Bertkau" authorityYear="1880" box="[1179,1304,1682,1708]" class="Arachnida" family="Dipluridae" genus="Thalerothele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF81FFBD71E7F90D411191F9" box="[1179,1304,1682,1708]" italics="true" pageId="29" pageNumber="190">Thalerothele</emphasis>
</taxonomicName>
was found to bear a lyra by
<bibRefCitation id="EFBE01D2FF81FFBD76B6F92D406C9198" author="Simon" box="[970,1125,1714,1741]" firstAuthor="Simon" pageId="29" pageNumber="190" pagination="314" refId="ref26556" refString="SIMON E. 1903. - Etudes arachnologiques. 34 e Memoire. LVI. Descriptions de deux especes nouvelles de la famille des Avicularides recueillis dans l'Ecuador par M. le Dr Rivet et faisant partie des collections du Museum de Paris. Annales de la Societe Entomologique de France 72: 314. https: // www. biodiversitylibrary. org / page / 8248189" type="journal article" year="1903">Simon (1903)</bibRefCitation>
, clearly indicating Berkau did not check for the presence of such a structure. Considering the possible lapsus, the description might match either
<taxonomicName id="4C2F07A0FF81FFBD70EDF96C477F9079" authorityName="Karsch" authorityYear="1879" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF81FFBD70EDF96C477F9079" italics="true" pageId="29" pageNumber="190">Linothele</emphasis>
</taxonomicName>
, or
<taxonomicName id="4C2F07A0FF81FFBD76DDF88D47F69079" authorityName="C. L. Koch" authorityYear="1850" box="[929,1023,1810,1836]" class="Arachnida" family="Dipluridae" genus="Trechona" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF81FFBD76DDF88D47F69079" box="[929,1023,1810,1836]" italics="true" pageId="29" pageNumber="190">Trechona</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B907C23FF81FFBD763FF8AD401090B9" blockId="29.[811,1457,215,2028]" pageId="29" pageNumber="190">
As the absence of a lyra cannot be confirmed by original designation and the
<typeStatus id="5494C281FF81FFBD717AF8CD403D9039" box="[1030,1076,1874,1900]" pageId="29" pageNumber="190">type</typeStatus>
is lost, the species cannot be placed in any diplurine genus with certainty. As a result,
<taxonomicName id="4C2F07A0FF81FFBD702CF8EE47B190FE" baseAuthorityName="Bertkau" baseAuthorityYear="1880" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD702CF8EE47B190FE" italics="true" pageId="29" pageNumber="190">Linothele gymnognatha</emphasis>
</taxonomicName>
is transferred back to
<taxonomicName id="4C2F07A0FF81FFBD71CDF80E410D90FE" authorityName="C. L. Koch" authorityYear="1850" box="[1201,1284,1937,1963]" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF81FFBD71CDF80E410D90FE" box="[1201,1284,1937,1963]" italics="true" pageId="29" pageNumber="190">Diplura</emphasis>
</taxonomicName>
due to original designation and
<taxonomicName id="4C2F07A0FF81FFBD769FF82E40C0909E" authorityName="Bertkau" authorityYear="1880" box="[995,1225,1969,1995]" class="Arachnida" family="Dipluridae" genus="Diplura" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="190" phylum="Arthropoda" rank="species" species="gymnognatha">
<emphasis id="B95BA031FF81FFBD769FF82E40C0909E" box="[995,1225,1969,1995]" italics="true" pageId="29" pageNumber="190">Diplura gymnognatha</emphasis>
</taxonomicName>
comb. rev. is considered a
<emphasis id="B95BA031FF81FFBD760FF84D401B90BE" box="[883,1042,2001,2027]" italics="true" pageId="29" pageNumber="190">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>