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<document id="9E62C88508E2A6D76C90B633A488B3C6" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE325FFB4FF75FF3DBA90FBB0" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena quatriloba Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="35" updateTime="1698480684898" updateUser="ExternalLinkService">
<mods:mods id="5FCB383D8FE3990EC2A79A8E61D81143" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2AA2FD8BAC83A18669470073DB32AF3">
<mods:title id="5AD3CD79E942A9C29277490DADCF9808">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="F6BC1898CA612D6960430C2BB0F1B564" type="personal">
<mods:role id="E4F7017A8663EAEDF2F573885CDF8932">
<mods:roleTerm id="7DDAB401E390846BE6C75550D5C3136B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A60CD58191731D8E18C02275F252EF5">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="23C5B92EAF0BCCBC42C1500F66B4323B">text</mods:typeOfResource>
<mods:relatedItem id="715B9880BF09C196A11099456187B176" type="host">
<mods:titleInfo id="0C9E9926252401C1F94D900014A12006">
<mods:title id="AD8B92A6B51FDFCB312C491F69BD9188">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="23AF11FB540C37A6D62B492C41CC8695">
<mods:date id="0A87BB56EAE16324CF81B046CEA28455">2017</mods:date>
<mods:detail id="1A140E06829A42911009DACCDDAE314A" type="pubDate">
<mods:number id="AED024117EE810823365EDEC25CCD503">2017-11-03</mods:number>
</mods:detail>
<mods:detail id="A64CFC2AD89DCF0557F5D0611A89639E" type="volume">
<mods:number id="38393A026D49DBDE75400109FF5FB146">4342</mods:number>
</mods:detail>
<mods:detail id="05A80E35F5AEA080E4F19621F5C70512" type="issue">
<mods:number id="32A82444F90F61DAD296714C892D05EF">1</mods:number>
</mods:detail>
<mods:extent id="00A90A541F7FEB45CCAA5DB4CC18E78D" unit="page">
<mods:start id="0A4183BF99A92F0ABFB05EE77219D060">1</mods:start>
<mods:end id="F3359C95F0BBFE633A55F0467254AD39">264</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="19F0219B7570A9F0F7EB5627A6B50A08">journal article</mods:classification>
<mods:identifier id="99BA5581B162B2E954AE6AF18C2012CB" type="CLB-Dataset">31639</mods:identifier>
<mods:identifier id="B75F593DB21FB4AC7DA783EAC03092BF" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
<mods:identifier id="A42BC2CEC1A85550E41C2274195AA344" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
<mods:identifier id="5D474AE424595B65BAE127E23E80BF24" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4FF14F1BC05FFE1A09FC2B43734A1A21" type="Zenodo-Dep">1041066</mods:identifier>
<mods:identifier id="552A23D32734245A336B61ABE6D56FED" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
</mods:mods>
<treatment id="038587BBE325FFB4FF75FF3DBA90FBB0" ID-DOI="http://doi.org/10.5281/zenodo.5323417" ID-GBIF-Taxon="137233451" ID-Zenodo-Dep="5323417" LSID="urn:lsid:plazi:treatment:038587BBE325FFB4FF75FF3DBA90FBB0" httpUri="http://treatment.plazi.org/id/038587BBE325FFB4FF75FF3DBA90FBB0" lastPageId="37" lastPageNumber="38" pageId="34" pageNumber="35">
<subSubSection id="C3366526E325FFB3FF75FF3DBBB4FF56" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8B9336ADE325FFB3FF75FF3DBB0FFF37" blockId="34.[151,648,152,211]" box="[151,563,152,178]" pageId="34" pageNumber="35">
<heading id="D0DB81C1E325FFB3FF75FF3DBB0FFF37" bold="true" box="[151,563,152,178]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="B958EABFE325FFB3FF75FF3DBB0FFF37" bold="true" box="[151,563,152,178]" pageId="34" pageNumber="35">
<taxonomicName id="4C2C4D2EE325FFB3FF75FF3DB8AFFF37" ID-CoL="3N3M8" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,403,152,178]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="quatriloba" status="sp. nov.">
<emphasis id="B958EABFE325FFB3FF75FF3DB8AFFF37" bold="true" box="[151,403,152,178]" italics="true" pageId="34" pageNumber="35">Hydraena quatriloba</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E325FFB3FE41FF3DBB0FFF37" box="[419,563,152,178]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE325FFB3FF75FF1FBBB4FF56" blockId="34.[151,648,152,211]" box="[151,648,186,211]" pageId="34" pageNumber="35">
<figureCitation id="13172A28E325FFB3FF75FF1FB9CDFF56" box="[151,241,186,211]" captionStart="FIGURE 24" captionStartId="34.[151,250,2011,2033]" captionTargetBox="[264,1323,684,1990]" captionTargetId="figure@34.[264,1323,684,1990]" captionTargetPageId="34" captionText="FIGURE 24. Hydraena quatriloba, holotype, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041090/files/figure.png" pageId="34" pageNumber="35">Figs. 24</figureCitation>
(habitus), 28 (aedeagus), 239 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E325FFB3FF75FEA7BA34FD92" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8B9336ADE325FFB3FF75FEA7BA34FD92" blockId="34.[151,1437,258,643]" pageId="34" pageNumber="35">
<materialsCitation id="3B443CF0E325FFB3FF75FEA7B8F2FEBA" ID-GBIF-Occurrence="1697157514" collectingDate="2007-11-15" collectingDateMax="2007-11-16" collectingDateMin="2007-11-15" collectionCode="NMW" collectorName="J. Stastny" latitude="-24.95" location="Tsimelahy" longLatPrecision="1249" longitude="46.616665" municipality="Andohahela" pageId="34" pageNumber="35" specimenCount="2" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="B958EABFE325FFB3FF75FEA7B880FE9E" bold="true" box="[151,444,258,283]" pageId="34" pageNumber="35">
<typeStatus id="5497880FE325FFB3FF75FEA7B9EEFE9E" box="[151,210,258,283]" pageId="34" pageNumber="35">Type</typeStatus>
Material.
<typeStatus id="5497880FE325FFB3FEADFEA7B880FE9E" box="[335,444,258,283]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE325FFB3FDFEFEA7BB4EFE9E" box="[540,626,258,283]" country="Madagascar" name="Toliara" pageId="34" pageNumber="35">
<emphasis id="B958EABFE325FFB3FDFEFEA7BB4EFE9E" bold="true" box="[540,626,258,283]" pageId="34" pageNumber="35">Toliara</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BF7ACD7E325FFB3FD62FEA7BA3BFE9E" box="[640,775,258,283]" pageId="34" pageNumber="35">Andohahela</collectingMunicipality>
NP,
<location id="8EF36076E325FFB3FCDCFEA7BA8FFE9E" LSID="urn:lsid:plazi:treatment:038587BBE325FFB4FF75FF3DBA90FBB0:8EF36076E325FFB3FCDCFEA7BA8FFE9E" box="[830,947,258,283]" latitude="-24.95" longLatPrecision="1249" longitude="46.616665" municipality="Andohahela" name="Tsimelahy" pageId="34" pageNumber="35" stateProvince="Toliara">Tsimelahy</location>
, pools in creek,
<geoCoordinate id="EE18506AE325FFB3FB88FEA6BDEEFE9E" box="[1130,1234,258,283]" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="-24.95">24° 57' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE325FFB3FB3DFEA7BC75FE9F" box="[1247,1353,258,283]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="46.616665">46° 37' E</geoCoordinate>
,
<collectingDate id="EFD6E985E325FFB3FAB4FEA7B9CCFEBA" pageId="34" pageNumber="35" value="2007-11-15" valueMax="2007-11-16" valueMin="2007-11-15">
<date id="FF92106DE325FFB3FAB4FEA7B9CCFEBA" pageId="34" pageNumber="35" value="2007-11-15" valueMax="2007-11-16" valueMin="2007-11-15">15–16 xi 2007</date>
</collectingDate>
,
<collectorName id="26D9537BE325FFB3FF1EFE82B854FEBA" box="[252,360,295,319]" pageId="34" pageNumber="35">J. Stastny</collectorName>
(
<collectionCode id="ED3DAE68E325FFB3FE9AFE82B8FAFEBA" box="[376,454,295,319]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="34" pageNumber="35">NMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B443CF0E325FFB3FE38FE83BA7CFE02" ID-GBIF-Occurrence="1697157530" collectingDate="2006-05-18" collectionCode="NHRS" collectorName="Bergsten" county="Same" elevation="219" latitude="-24.516666" location="Manakaravavy" longLatPrecision="1251" longitude="44.616665" municipality="Manakaravavy" pageId="34" pageNumber="35" specimenCount="2" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="5497880FE325FFB3FE38FE83BB6EFEBA" box="[474,594,294,319]" pageId="34" pageNumber="35" type="paratype">
<emphasis id="B958EABFE325FFB3FE38FE83BB6EFEBA" bold="true" box="[474,594,294,319]" pageId="34" pageNumber="35">Paratypes</emphasis>
</typeStatus>
(15):
<collectingCounty id="62F24E21E325FFB3FD7AFE82BBEAFEBA" box="[664,726,295,319]" pageId="34" pageNumber="35">Same</collectingCounty>
data as
<typeStatus id="5497880FE325FFB3FCD7FE83BAABFEBA" box="[821,919,294,319]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
(11);
<collectingRegion id="49E8F84FE325FFB3FC3EFE83BD0CFEBA" box="[988,1072,294,319]" country="Madagascar" name="Toliara" pageId="34" pageNumber="35">
<emphasis id="B958EABFE325FFB3FC3EFE83BD0CFEBA" bold="true" box="[988,1072,294,319]" pageId="34" pageNumber="35">Toliara</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BF7ACD7E325FFB3FBA0FE83BDD6FEBA" box="[1090,1258,294,319]" pageId="34" pageNumber="35">Manakaravavy</collectingMunicipality>
,
<location id="8EF36076E325FFB3FB14FE83BCA0FEBA" LSID="urn:lsid:plazi:treatment:038587BBE325FFB4FF75FF3DBA90FBB0:8EF36076E325FFB3FB14FE83BCA0FEBA" box="[1270,1436,294,319]" county="Same" latitude="-24.516666" longLatPrecision="1251" longitude="44.616665" municipality="Manakaravavy" name="Manakaravavy" pageId="34" pageNumber="35" stateProvince="Toliara">Manakaravavy</location>
R. dried out river with waterpools on sandy bottom with algal mats, N:-24° 31.222, E:44° 37.388, elev.
<elevation id="0001D19EE325FFB3FAC2FEEEBC55FEE7" box="[1312,1385,331,355]" metricMagnitude="2" metricUnit="m" metricValue="2.19" pageId="34" pageNumber="35" unit="m" value="219.0">
<quantity id="4CD49B48E325FFB3FAC2FEEEBC55FEE7" box="[1312,1385,331,355]" metricMagnitude="2" metricUnit="m" metricValue="2.19" pageId="34" pageNumber="35" unit="m" value="219.0">219 m</quantity>
</elevation>
,
<geoCoordinate id="EE18506AE325FFB3FA97FEEEB9EDFE02" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="-24.516666">24° 31' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE325FFB3FF3CFECAB875FE03" box="[222,329,367,391]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="44.616665">44° 37' E</geoCoordinate>
,
<collectingDate id="EFD6E985E325FFB3FEB4FECAB8F7FE02" box="[342,459,366,391]" pageId="34" pageNumber="35" value="2006-05-18">
<date id="FF92106DE325FFB3FEB4FECAB8F7FE02" box="[342,459,366,391]" pageId="34" pageNumber="35" value="2006-05-18">18 v 2006</date>
</collectingDate>
,
<collectorName id="26D9537BE325FFB3FE35FECABB06FE02" box="[471,570,367,391]" pageId="34" pageNumber="35">Bergsten</collectorName>
<emphasis id="B958EABFE325FFB3FDA0FECABB40FE02" box="[578,636,366,391]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(P49) (1
<collectionCode id="ED3DAE68E325FFB3FD0BFECABA05FE02" box="[745,825,367,391]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="34" pageNumber="35">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B443CF0E325FFB3FCAEFECBBC25FE4A" ID-GBIF-Occurrence="1697157522" collectingDate="2006-05-18" collectionCode="NHRS" collectorName="Bergsten" county="Menarandra" elevation="227" latitude="-24.716667" location="Ampanihy" longLatPrecision="1249" longitude="45.05" municipality="Menarandra" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<collectingCounty id="62F24E21E325FFB3FCAEFECBBAEAFE02" box="[844,982,366,391]" pageId="34" pageNumber="35">Menarandra</collectingCounty>
,
<collectingMunicipality id="6BF7ACD7E325FFB3FC01FECBBD56FE02" box="[995,1130,366,391]" pageId="34" pageNumber="35">Menarandra</collectingMunicipality>
R.,
<quantity id="4CD49B48E325FFB3FB78FECBBDE6FE02" box="[1178,1242,366,391]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="35" unit="km" value="40.0">40km</quantity>
from
<location id="8EF36076E325FFB3FAC3FECBBCA4FE02" LSID="urn:lsid:plazi:treatment:038587BBE325FFB4FF75FF3DBA90FBB0:8EF36076E325FFB3FAC3FECBBCA4FE02" box="[1313,1432,366,391]" county="Menarandra" latitude="-24.716667" longLatPrecision="1249" longitude="45.05" municipality="Menarandra" name="Ampanihy" pageId="34" pageNumber="35" stateProvince="Toliara">Ampanihy</location>
, pools beside a river close to village, algae in pools and sandy bottom with some wood,
<collectingDate id="EFD6E985E325FFB3FB77FE37BC39FE2E" box="[1173,1285,402,427]" pageId="34" pageNumber="35" value="2006-05-18">
<date id="FF92106DE325FFB3FB77FE37BC39FE2E" box="[1173,1285,402,427]" pageId="34" pageNumber="35" value="2006-05-18">18.v.2006</date>
</collectingDate>
, P50, N:-24° 43.104, E:45° 2.859, elev.
<quantity id="4CD49B48E325FFB3FE2DFE12BB27FE4B" box="[463,539,439,463]" metricMagnitude="2" metricUnit="m" metricValue="2.27" pageId="34" pageNumber="35" unit="m" value="227.0">
<elevation id="0001D19EE325FFB3FE2DFE12BB27FE4B" box="[463,539,439,463]" metricMagnitude="2" metricUnit="m" metricValue="2.27" pageId="34" pageNumber="35" unit="m" value="227.0">227 m</elevation>
</quantity>
,
<geoCoordinate id="EE18506AE325FFB3FDC9FE12BBA5FE4A" box="[555,665,439,463]" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="-24.716667">24° 43' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE325FFB3FD4BFE13BA30FE4B" box="[681,780,438,463]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="45.05">45° 3' E</geoCoordinate>
,
<collectingDate id="EFD6E985E325FFB3FCF9FE12BAA9FE4A" box="[795,917,438,463]" pageId="34" pageNumber="35" value="2006-05-18">
<date id="FF92106DE325FFB3FCF9FE12BAA9FE4A" box="[795,917,438,463]" pageId="34" pageNumber="35" value="2006-05-18">18 v 2006</date>
</collectingDate>
,
<collectorName id="26D9537BE325FFB3FC41FE12BD3AFE4A" box="[931,1030,439,463]" pageId="34" pageNumber="35">Bergsten</collectorName>
<emphasis id="B958EABFE325FFB3FBF3FE12BD71FE4A" box="[1041,1101,438,463]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(P50) (1
<collectionCode id="ED3DAE68E325FFB3FB23FE12BC2DFE4A" box="[1217,1297,439,463]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="34" pageNumber="35">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B443CF0E325FFB3FACAFE13BA38FD92" ID-GBIF-Occurrence="1697157526" collectingDate="2001-09-06" collectionCode="NMW" county="Tsimelahy" elevation="100" latitude="-24.933332" location="Analamatsaky" longLatPrecision="1249" longitude="46.633335" municipality="Tulear" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<collectingCounty id="62F24E21E325FFB3FACAFE13BCA0FE4A" box="[1320,1436,438,463]" pageId="34" pageNumber="35">Tsimelahy</collectingCounty>
(
<collectingMunicipality id="6BF7ACD7E325FFB3FF7DFE7FB9D1FE76" box="[159,237,474,499]" pageId="34" pageNumber="35">Tulear</collectingMunicipality>
),
<location id="8EF36076E325FFB3FEE2FE7FB89DFE76" LSID="urn:lsid:plazi:treatment:038587BBE325FFB4FF75FF3DBA90FBB0:8EF36076E325FFB3FEE2FE7FB89DFE76" box="[256,417,474,499]" county="Tsimelahy" latitude="-24.933332" longLatPrecision="1249" longitude="46.633335" municipality="Tulear" name="Analamatsaky" pageId="34" pageNumber="35" stateProvince="Toliara">Analamatsaky</location>
, ditches near the village (dead branches of small stream), elev.
<elevation id="0001D19EE325FFB3FB87FE7EBD92FE77" box="[1125,1198,475,499]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="m" value="100.0">
<quantity id="4CD49B48E325FFB3FB87FE7EBD92FE77" box="[1125,1198,475,499]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="m" value="100.0">100 m</quantity>
</elevation>
,
<geoCoordinate id="EE18506AE325FFB3FB58FE7EBC1DFE76" box="[1210,1313,474,499]" direction="south" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="-24.933332">24° 56' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE325FFB3FACCFE7FBCABFE77" box="[1326,1431,474,499]" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="46.633335">46° 38' E</geoCoordinate>
,
<collectingDate id="EFD6E985E325FFB3FF75FE5BB838FD92" box="[151,260,510,535]" pageId="34" pageNumber="35" value="2001-09-06">
<date id="FF92106DE325FFB3FF75FE5BB838FD92" box="[151,260,510,535]" pageId="34" pageNumber="35" value="2001-09-06">6 ix 2001</date>
</collectingDate>
, Gerecke &amp; Goldschmidt (MD 62) (2
<collectionCode id="ED3DAE68E325FFB3FD4DFE5ABBC0FD92" box="[687,764,511,535]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="34" pageNumber="35">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E325FFB3FF25FD87B8AEFD06" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8B9336ADE325FFB3FF25FD87B8AEFD06" blockId="34.[151,1437,258,643]" pageId="34" pageNumber="35">
<emphasis id="B958EABFE325FFB3FF25FD87B8EFFDBE" bold="true" box="[199,467,546,571]" pageId="34" pageNumber="35">Differential Diagnosis.</emphasis>
Very similar in size and habitus to
<taxonomicName id="4C2C4D2EE325FFB3FC87FD81BAC3FDBE" authorityName="Perkins" authorityYear="2017" box="[869,1023,546,571]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="bisinuloba">
<emphasis id="B958EABFE325FFB3FC87FD81BAC3FDBE" box="[869,1023,546,571]" italics="true" pageId="34" pageNumber="35">H. bisinuloba</emphasis>
</taxonomicName>
; reliable determinations will require dissection of males. The distal process is shaped differently in the two species, among other differences in the aedeagi (
<figureCitation id="13172A28E325FFB3FF1EFDCFB865FD06" box="[252,345,618,643]" captionStart="FIGURES 25 – 26" captionStartId="35.[151,264,1967,1990]" captionText="FIGURES 25 – 26. Hydraena, aedeagi of holotypes." pageId="34" pageNumber="35">Figs. 26</figureCitation>
,
<figureCitation id="13172A28E325FFB3FE87FDCEB8B9FD06" box="[357,389,619,643]" captionStart="FIGURES 27 – 28" captionStartId="36.[151,264,1876,1899]" captionText="FIGURES 27 – 28. Hydraena, aedeagi of holotypes." pageId="34" pageNumber="35">28</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF536625E325FFB3FF75F87EBAF0F874" box="[151,972,2011,2033]" httpUri="https://zenodo.org/record/1041090/files/figure.png" pageId="34" pageNumber="35" targetBox="[264,1323,684,1990]" targetPageId="34">
<paragraph id="8B9336ADE325FFB3FF75F87EBAF0F874" blockId="34.[151,972,2011,2033]" box="[151,972,2011,2033]" pageId="34" pageNumber="35">
<emphasis id="B958EABFE325FFB3FF75F87EB823F875" bold="true" box="[151,287,2011,2033]" pageId="34" pageNumber="35">FIGURE 24.</emphasis>
<taxonomicName id="4C2C4D2EE325FFB3FEC7F87EB8C0F874" authorityName="Perkins" authorityYear="2017" box="[293,508,2011,2033]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="quatriloba">
<emphasis id="B958EABFE325FFB3FEC7F87EB8C0F874" box="[293,508,2011,2033]" italics="true" pageId="34" pageNumber="35">Hydraena quatriloba</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<caption id="DF536625E324FFB2FF75F80ABB98F840" box="[151,676,1967,1990]" pageId="35" pageNumber="36">
<paragraph id="8B9336ADE324FFB2FF75F80ABB98F840" blockId="35.[151,676,1967,1990]" box="[151,676,1967,1990]" pageId="35" pageNumber="36">
<emphasis id="B958EABFE324FFB2FF75F80AB86FF841" bold="true" box="[151,339,1967,1990]" pageId="35" pageNumber="36">FIGURES 25–26.</emphasis>
<taxonomicName id="4C2C4D2EE324FFB2FEB8F80AB883F840" box="[346,447,1967,1989]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B958EABFE324FFB2FEB8F80AB883F840" box="[346,447,1967,1989]" italics="true" pageId="35" pageNumber="36">Hydraena</emphasis>
</taxonomicName>
, aedeagi of holotypes.
</paragraph>
</caption>
<caption id="DF536625E323FFB5FF75F8F1BB98F8EE" box="[151,676,1876,1899]" pageId="36" pageNumber="37">
<paragraph id="8B9336ADE323FFB5FF75F8F1BB98F8EE" blockId="36.[151,676,1876,1899]" box="[151,676,1876,1899]" pageId="36" pageNumber="37">
<emphasis id="B958EABFE323FFB5FF75F8F1B868F8EF" bold="true" box="[151,340,1876,1899]" pageId="36" pageNumber="37">FIGURES 27–28.</emphasis>
<taxonomicName id="4C2C4D2EE323FFB5FEB8F8F0B883F8EE" box="[346,447,1877,1899]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B958EABFE323FFB5FEB8F8F0B883F8EE" box="[346,447,1877,1899]" italics="true" pageId="36" pageNumber="37">Hydraena</emphasis>
</taxonomicName>
, aedeagi of holotypes.
</paragraph>
</caption>
<subSubSection id="C3366526E322FFB4FF25FF32BB95FB95" pageId="37" pageNumber="38" type="description">
<paragraph id="8B9336ADE322FFB4FF25FF32BA37FF51" blockId="37.[151,1437,151,1077]" pageId="37" pageNumber="38">
<emphasis id="B958EABFE322FFB4FF25FF32B86DFF35" bold="true" box="[199,337,151,176]" pageId="37" pageNumber="38">Description</emphasis>
. Size:
<typeStatus id="5497880FE322FFB4FE41FF32BB39FF35" box="[419,517,151,176]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.69/0.72; head width 0.44; pronotum 0.41/0.54, PA 0.47, PB 0.51; elytra 1.01/0.72.
</paragraph>
<paragraph id="8B9336ADE322FFB4FF25FF45BB0AFEBA" blockId="37.[151,1437,151,1077]" pageId="37" pageNumber="38">Dorsum with head and pronotal disc dark brown to piceous; pronotum with margins light brown to brown; elytra brown; legs light brown to testaceous; maxillary palpi light brown to testaceous, distal 1/3 of last palpomere slightly darker, except tip testaceous.</paragraph>
<paragraph id="8B9336ADE322FFB4FF25FEE9BA64FE70" blockId="37.[151,1437,151,1077]" pageId="37" pageNumber="38">Head with clypeus moderately finely and moderately densely punctate, interstices shining on median area, lateral areas microreticulate, dull; frons coarsely densely punctate, interstices ca 0.5–1xpd. Pronotum coarsely densely and deeply punctate, interstices narrow walls or less than 1xpd, except slightly larger, ca. 1xpd, on center of disc, shining. Mentum moderately finely punctate, interstices shining; postmentum weakly rugose, dull. Genae raised, shining, posterior margin abrupt, posterior ridge absent.</paragraph>
<paragraph id="8B9336ADE322FFB4FF25FDA5BBBEFDB8" blockId="37.[151,1437,151,1077]" pageId="37" pageNumber="38">Pronotum sinuate laterally; anterior margin straight behind eyes, emarginate behind frons; PF1 absent; PF2 very shallow; PF3 and PF4 deep, confluent.</paragraph>
<paragraph id="8B9336ADE322FFB4FF25FDE2B8A1FD75" blockId="37.[151,1437,151,1077]" pageId="37" pageNumber="38">Elytra weakly arcuate laterally; posterior declivity steep, summit slightly past midlength; lateral explanate margins moderately wide; each elytron with very low, indistinct tumidity on posterior declivity; serial punctures round, separated by ca. 0.5–1xpd, slightly smaller than largest pronotal punctures. Intervals very weakly if at all raised, shining, ca. 0.5–1xpd. Apices in dorsal aspect conjointly truncate, in posterior aspect margins forming deep angle with one another.</paragraph>
<paragraph id="8B9336ADE322FFB4FF25FD59BB95FB95" blockId="37.[151,1437,151,1077]" pageId="37" pageNumber="38">
Ratios of P2 width and plaque shape (P2/w/l/s [anteriorly]) ca. 1/0.2/2.5/1. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt. Plaques very narrow lines, not carinate, straight or very weakly arcuate, converging anteriorly, widely separated posteriorly, ca.
<date id="FF92106DE322FFB4FC5EFCE1BAE4FCD9" box="[956,984,836,860]" pageId="37" pageNumber="38">2x</date>
as wide as anteriorly, located at sides of deep median depression. Metaventrite anteromedian carina very small, short and low, almost absent; short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca.
<date id="FF92106DE322FFB4FE65FC15B89FFC4D" box="[391,419,944,968]" pageId="37" pageNumber="38">2x</date>
P2. Protibia very weakly arcuate, distal 0.5 slightly thickened. Mesotibia and metatibia straight, slender. Metafemur rather slender, slightly arcuate on lateral margin, slightly sinuate on medial margin. Abdominal apex with deep apicomedian notch.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E322FFB4FF25FBB9BA90FBB0" box="[199,940,1052,1077]" pageId="37" pageNumber="38" type="etymology">
<paragraph id="8B9336ADE322FFB4FF25FBB9BA90FBB0" blockId="37.[151,1437,151,1077]" box="[199,940,1052,1077]" pageId="37" pageNumber="38">
<emphasis id="B958EABFE322FFB4FF25FBB9B872FBB0" bold="true" box="[199,334,1052,1077]" pageId="37" pageNumber="38">Etymology.</emphasis>
Named in reference to the four lobes of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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