treatments-xml/data/FA/F3/23/FAF323D416B4836684475A52F60E0B59.xml
2024-06-21 12:58:46 +02:00

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<document id="58D40D01F9877BA5E34A52DC0BB887BC" ID-CLB-Dataset="30577" ID-DOI="http://dx.doi.org/10.3897/subtbiol.25.23563" ID-GBIF-Dataset="d5461c88-fca2-4f85-96f5-c63ff3a0ca55" ID-Pensoft-Pub="1314-2615-25-9" ID-ZooBank="9F789679CD744D54A7F2B0087E154571" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2615-25-9" ModsDocOrigin="Subterranean Biology 25" ModsDocTitle="First record of subterranean rissoidean gastropod assemblages in Southeast Asia (Mollusca, Gastropoda, Pomatiopsidae)" checkinTime="1520490019267" checkinUser="pensoft" docAuthor="Grego, Jozef" docDate="2018" docId="FAF323D416B4836684475A52F60E0B59" docLanguage="en" docName="SubterBiol 25: 9-34" docOrigin="Subterranean Biology 25" docSource="http://dx.doi.org/10.3897/subtbiol.25.23563" docTitle="Pseudoiglica pseudoiglica Grego, 2018, sp. n." docType="treatment" docUuid="3D5A63D3-B516-4929-B002-DF41C10B23BA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="13" masterDocId="F504CB6AE163DD43C7208D79FF885C6D" masterDocTitle="First record of subterranean rissoidean gastropod assemblages in Southeast Asia (Mollusca, Gastropoda, Pomatiopsidae)" masterLastPageNumber="34" masterPageNumber="9" pageNumber="11" updateTime="1701378715001" updateUser="ExternalLinkService">
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<mods:title id="039CBFCCA3197D0D490AD66A9A072997">First record of subterranean rissoidean gastropod assemblages in Southeast Asia (Mollusca, Gastropoda, Pomatiopsidae)</mods:title>
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<mods:name id="080E2ACD4BE4EC2B19558543ACED4054" type="personal">
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<mods:roleTerm id="8DD633A8EE2F99D0D05B1266820587A6">Author</mods:roleTerm>
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<mods:namePart id="B59A88ED8163EEFDCFDBD5EC88B6FA93">Grego, Jozef</mods:namePart>
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<mods:title id="845F764E01A8F88A85081E9EF10D5033">Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part id="276F5DB04E0E44010644C3AFF1C12469">
<mods:date id="9CB084A29023DAE04395669463B084C1">2018</mods:date>
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<mods:number id="174F30B6A41E7E0C0BEA37D2E37EB53B">25</mods:number>
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<mods:classification id="6E4FD3823BC35D91A4E8E8802A89CE20">journal article</mods:classification>
<mods:identifier id="66FD51EA1CC41FAF647EAD07D44B2D65" type="DOI">http://dx.doi.org/10.3897/subtbiol.25.23563</mods:identifier>
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<treatment id="FAF323D416B4836684475A52F60E0B59" ID-GBIF-Taxon="141400996" LSID="urn:lsid:zoobank.org:act:3D5A63D3-B516-4929-B002-DF41C10B23BA" httpUri="http://treatment.plazi.org/id/FAF323D416B4836684475A52F60E0B59" lastPageId="4" lastPageNumber="13" pageId="2" pageNumber="11">
<subSubSection id="7FBD5FF7B4DAE960EAE8B7DCE085B9CE" pageId="2" pageNumber="11" type="nomenclature">
<paragraph id="1F75B0DB7185F4B7DD5EA82EAB445232" pageId="2" pageNumber="11">
<taxonomicName id="A0843AB0CD4F7D4A31E883EB082AD3D3" ID-CoL="4NYKR" LSID="http://zoobank.org/3D5A63D3-B516-4929-B002-DF41C10B23BA" class="Gastropoda" family="Pomatiopsidae" genus="Pseudoiglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoiglica pseudoiglica" order="Littorinimorpha" pageId="2" pageNumber="11" phylum="Mollusca" rank="species" species="pseudoiglica">Pseudoiglica pseudoiglica</taxonomicName>
<taxonomicNameLabel id="CA18B85E24C33B13AA58E5EB51619DCB" pageId="2" pageNumber="11">sp. n.</taxonomicNameLabel>
Figures 3-6
</paragraph>
</subSubSection>
<subSubSection id="C713D8EAC004E5DCA1732674523EBF09" pageId="2" pageNumber="11" type="type locality">
<paragraph id="4FEDB7518439487C5945EC7F5FE9D149" pageId="2" pageNumber="11">Type locality.</paragraph>
<paragraph id="9A9037C66291667283749A61D20373AB" pageId="2" pageNumber="11">
Laos; Khammouane Province, Ban Na village 20 km NNE of Thakhek, Tham Khon
<normalizedToken id="8AC15E892640E597A1C75C8CCD474986" originalValue="Dôn">Don</normalizedToken>
Cave
<geoCoordinate id="CDE999FDDF21C14F764336ADC245FFE4" direction="north" orientation="latitude" precision="9" value="17.563667">17°33.82'N</geoCoordinate>
;
<geoCoordinate id="2EBF349AB8457AC5C17260C13B13F549" direction="east" orientation="longitude" precision="9" value="104.871666">104°52.30'E</geoCoordinate>
, 161 m a.s.l., Earthquake Dome 3 km from the south entrance, sand sediments on bank of cave river (Fig. 2B).
</paragraph>
</subSubSection>
<subSubSection id="5DE2E7059D1B940F5802C37D9766BAC1" pageId="2" pageNumber="11" type="type material">
<paragraph id="3C4F5B5F2F0CCDBCFAEB7A6E29384FAD" pageId="2" pageNumber="11">Type material.</paragraph>
<paragraph id="9D977FEC0D56B4C3493BC6AAF7E35261" pageId="2" pageNumber="11">
Holotype: type locality: J. Grego and M.
<normalizedToken id="0131730FC00C69CDB5C75E8F691941AD" originalValue="Olšavský">Olsavsky</normalizedToken>
leg. 11-12 February 2017 (NHMUK 20180001).
</paragraph>
<paragraph id="F44C710A52AC9B8403BF959FBE5C9D9E" pageId="2" pageNumber="11">
Paratypes: type locality (NHMUK 20180016 - 1 specimen; HNHM 102769 - 1 specimen; coll. Grego F0871 - 6 specimens); Laos, Khammouane Province, Tham Nam
<normalizedToken id="995B9777E621D2A01C60B2F20F614059" originalValue="Dôn">Don</normalizedToken>
Cave
<geoCoordinate id="920C2BBDA9E89421D18F1A2D5B2089FF" direction="north" orientation="latitude" precision="9" value="17.563667">17°33.82'N</geoCoordinate>
;
<geoCoordinate id="7602B3D5659C26A5DDF293F483B64221" direction="east" orientation="longitude" precision="9" value="104.871666">104°52.30'E</geoCoordinate>
, 161 m a.s.l., temporary side rivulet sediment at entrance passage 1.5 km from the main entrance, dry sand on the cave floor; J. Grego leg. 11 February 2017 (coll. Grego F0863 - 1 specimen).
</paragraph>
</subSubSection>
<subSubSection id="6886E8C1CAA8AB822CFA49455D6AEFDF" pageId="2" pageNumber="11" type="materials_examined">
<paragraph id="086A27B0E6A81BE7BBFCBA3B020662D3" pageId="2" pageNumber="11">Other material.</paragraph>
<paragraph id="AE77678518055B72BB29AA235BC3D38D" pageId="2" pageNumber="11">
Laos, Khammouane Province, 2 km WNW of Ban Na village, Pha Soung Cave, Frog Lake, 155 m a.s.l., J. Grego leg. 09 February 2017,
<geoCoordinate id="7DB13DAD0BC4C209F29F1F9A9CA9030D" direction="north" orientation="latitude" precision="1" value="17.550867">17°33.052'N</geoCoordinate>
;
<geoCoordinate id="5F04C1CAC6FE4532FF2B71CAADCFD57D" direction="east" orientation="longitude" precision="1" value="104.8735">104°52.410'E</geoCoordinate>
(coll. Grego F0888).
</paragraph>
</subSubSection>
<subSubSection id="DA764640301FB29E920FBA340085B0E7" pageId="2" pageNumber="11" type="description">
<paragraph id="8E200ECE6C5346D36A27D3AADFEF17BD" pageId="2" pageNumber="11">Measurements.</paragraph>
<paragraph id="D87C0D5BDF8EC492A3A8047BA1A71243" pageId="2" pageNumber="11">Holotype: H 3.08 mm; W 1.32 mm; BW 0.95 mm; BH 1.50 mm; AH 0.81 mm; AW 0.75 mm; H/W 2.33; AH/AW 1.08; W/BW 1.39; H/BH 2.05; H/AH 3.80; W/AW 1.76. Paratype 1: H 3.05 mm; W 1.31 mm; BH 1.00 mm; BW 1.55 mm; AH 0.87 mm; AW 0.75 mm; H/W 2.33; AH/AW 1.16; W/BW 1.31; H/BH 1.97; H/AH 3.51; W/AW 1.76.</paragraph>
</subSubSection>
<subSubSection id="5064214FD4F889B0AA9BDCECFE7FD946" lastPageId="3" lastPageNumber="12" pageId="2" pageNumber="11" type="diagnosis">
<paragraph id="CDE2214C460554CC082B89D8FA1B4BFE" pageId="2" pageNumber="11">Diagnosis.</paragraph>
<paragraph id="4BBCD63B8EB594FAC3A1898C5BBC93A1" lastPageId="3" lastPageNumber="12" pageId="2" pageNumber="11">
This new species is similar to the syntopic
<taxonomicName id="8425202AEB353B703F0B7ED638F5056F" class="Gastropoda" family="Pomatiopsidae" genus="Pseudoiglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoiglica kameniari" order="Littorinimorpha" pageId="2" pageNumber="11" phylum="Mollusca" rank="species" species="kameniari">Pseudoiglica kameniari</taxonomicName>
sp. n., from which it differs by its more slender, elongated shell with a more prominent umbilicus and less elongated aperture situated further to the right of the columellar axis.
<pageBreakToken id="007072652D0190891A0804B4DE63A706" pageId="3" pageNumber="12" start="start">It</pageBreakToken>
differs from syntopic
<taxonomicName id="4D797F97C4C976E2CAE4B634D4859050" lsidName="P. olsavskyi" pageId="3" pageNumber="12" rank="species" species="olsavskyi">P. olsavskyi</taxonomicName>
sp. n. by its markedly larger and more conical shell shape and proportionally larger aperture.
<taxonomicName id="85DAFD73D177A4AAE0ECFA7A73F80EB5" class="Gastropoda" family="Pomatiopsidae" genus="Pseudoiglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoiglica phonsavanica" order="Littorinimorpha" pageId="3" pageNumber="12" phylum="Mollusca" rank="species" species="phonsavanica">Pseudoiglica phonsavanica</taxonomicName>
sp. n. (Xianghouan Province) has more a robust shell with more prominent body whorl and a differently shaped aperture.
</paragraph>
</subSubSection>
<subSubSection id="BBE07D4797FAE03EED02020D57BBAA41" pageId="3" pageNumber="12" type="description">
<paragraph id="884BC980ADAF13C52E17C6F10FE5F1FE" pageId="3" pageNumber="12">Description.</paragraph>
<paragraph id="CDF4DA8E79E197DF76A75B82CEE125A0" pageId="3" pageNumber="12">The milky yellowish silky shell has six tumid convex whorls with a deep suture and a blunt apex. The surface is smooth and shiny. The shell is elongated, almost cylindrical, slightly tapering towards a blunt apex, the umbilicus is tiny, open. In frontal view, the lateral aperture protrudes against the rest of the teleoconch. The aperture is ovoid, separated from the body whorl by a gap. The peristome margin is blunt, equally thick all the way around and slightly reflexed outwards. The outer lip is sinuous in lateral view and slightly scooped forward at its lower end.</paragraph>
</subSubSection>
<subSubSection id="2B214CCCCFF694144EB0CDF4DA4DCBE7" pageId="3" pageNumber="12" type="etymology">
<paragraph id="80E8A8D90C4F2D4C900FF63FE26D48BA" pageId="3" pageNumber="12">Etymology.</paragraph>
<paragraph id="7EFEB2E5D6CE2DBB79C6C7301132A0F4" pageId="3" pageNumber="12">
See the etymology of the genus
<taxonomicName id="FAA273CBBEAFBED54ED84DC23B300CB6" class="Gastropoda" family="Pomatiopsidae" genus="Pseudoiglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoiglica" order="Littorinimorpha" pageId="3" pageNumber="12" phylum="Mollusca" rank="genus">Pseudoiglica</taxonomicName>
gen. n.
</paragraph>
</subSubSection>
<subSubSection id="DAFBE7CA895976B32DEDFA9483479B38" pageId="3" pageNumber="12" type="distribution">
<paragraph id="E45DB26EC1D0C62D9EF0F38C06BE92AC" pageId="3" pageNumber="12">Distribution.</paragraph>
<paragraph id="8086998F7C7DF9B7688C1A232F482151" pageId="3" pageNumber="12">
Only known from the type locality and nearby sites in Tham Khon
<normalizedToken id="0FEA09546D3DDF794507CA6B3931DA99" originalValue="Dôn">Don</normalizedToken>
Cave as well as in the related source of the Nam
<normalizedToken id="15517DF55AD82D1E6FEC108353CC604F" originalValue="Dôn">Don</normalizedToken>
River and from sediments in Tham Pha Soung Cave.
</paragraph>
</subSubSection>
<subSubSection id="C4750178AE4AE885AE7D9DBA6D138476" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12" type="ecology">
<paragraph id="D64FFEBC67F916FBFE43078C80570370" pageId="3" pageNumber="12">Ecology.</paragraph>
<paragraph id="AE42AFA4FFBFEAB51B80B9DDB384A384" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12">
Empty shells of the new species were extracted from the side stream sandy sediments of an underground river inside the cave Tham Khon
<normalizedToken id="43844AF4140ED012B15980344F0A021E" originalValue="Dôn">Don</normalizedToken>
about 3 km from the main entrance situated above the source of Nam
<normalizedToken id="6DCAABDB11333D50ED18B3491F1159DE" originalValue="Dôn">Don</normalizedToken>
River (Fig. 1A). The cave Tham Khon
<normalizedToken id="516C5B0620EC674EED5B2A81206BB16B" originalValue="Dôn">Don</normalizedToken>
is situated under the massif of Mount Pha Kouankaohong (Fig. 2C) north of Ban Na Village. It represents the largest explored cave system in Laos, with a length of known passages of ca. 42 km. The cave is morphologically diverse with large domes and passages modeled by phreatic corrosion representing the cave multiple genetic horizons formed during the past 11 million years (since Late Miocene-Lower Pliocene) of its natural history possibly driven by a hydrothermal, H2S speleogenesis (
<bibRefCitation id="2A9E17691C463899DFF6E5743F54F258" author="Mouret, C" journalOrPublisher="Stuttgart," pageId="23" pageNumber="32" pagination="411 - 414" title="Main 2001 to early 2005 results on the karst of Khammouane, central Laos: long caves, sloping caves, hollow stalagmites and others. In: Proceeding of the 14 - th International Congress of Speleology (Athens-Kalamos, 21 - 18 August 2005) Paper No." volume="143" year="2005">Mouret 2005</bibRefCitation>
). Despite the remarkable length of the cave system, its water passages are accessible only for a limited length near the entrance. The major part of the underground river comprises mostly unexplored submerged cave passages. The main sampling site was located at the bottom of Earthquake Dome (Fig. 2B), named after the sounds of the earthquake experienced here by the first explorers (Claude Mouret and Jean-Francois
<normalizedToken id="946BE87A67D4E1ADE15F92AA64C076B6" originalValue="Vacquié">Vacquie</normalizedToken>
pers. com.). The dome floor is covered by very large flat limestone slabs approx. 0.8-1.2 m thick fallen from the horizontally flat ceiling that reflects the horizontal beds of Carboniferous/Permian Khammouane Limestone. The fallen slabs fragmented the underground stream into several lakes and helped to create the sedimentation zones in which empty shells could be deposited during high water flows. A few shells were also found in the sand floor of the entrance passage approximately 1.5 km from the entrance close to the junction with a temporary side stream, as well as in the sand deposited directly at the cave entrance at the source of the Nam
<normalizedToken id="BA19C00422DCA827FC15BFBB97DC2CB1" originalValue="Dôn">Don</normalizedToken>
River (Fig. 1A). The character of the material deposited in the cave sediments and the freshwater shell assemblages suggested their autochthonous origin rather than the allochthonous influence of horizontal surface waters. The occurrence of tiny terrestrial gastropod shells of mainly soil and leaf litter dwelling families such as
<taxonomicName id="8803742DC214B5E760360E5A19C68FEA" family="Vertiginidae" lsidName="" pageId="3" pageNumber="12" rank="family">Vertiginidae</taxonomicName>
Fitzinger, 1833 (
<taxonomicName id="0635930439FAD92B909617B752D9935A" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="3" pageNumber="12" phylum="Mollusca" rank="genus">Hypselostoma</taxonomicName>
sp.,
<taxonomicName id="C8D670DE544C0CAD1DCE4CECBEA99892" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="4" pageNumber="13" phylum="Mollusca" rank="genus">
<pageBreakToken id="41A998476E7235E382CAF4B31CADC3D2" pageId="4" pageNumber="13" start="start">Angustopila</pageBreakToken>
</taxonomicName>
sp.,
<taxonomicName id="0A5F998388DF5FEC7B5877FD694CD4E7" class="Gastropoda" family="Gastrocoptidae" genus="Krobylos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krobylos" order="Stylommatophora" pageId="4" pageNumber="13" phylum="Mollusca" rank="genus">Krobylos</taxonomicName>
sp.,
<taxonomicName id="A5C59F91808ACDEC7160E499DA304650" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraboysidia" order="Stylommatophora" pageId="4" pageNumber="13" phylum="Mollusca" rank="genus">Paraboysidia</taxonomicName>
sp.) and
<taxonomicName id="A1BCEB8AAF8F18832B1473668DA07EA2" family="Diapheridae" lsidName="" pageId="4" pageNumber="13" rank="family">Diapheridae</taxonomicName>
Panha &amp; Naggs, 2010 (
<taxonomicName id="7467E6C8373B8E2992DF62A22E316350" class="Gastropoda" family="Diapheridae" genus="Sinoennea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinoennea" order="Stylommatophora" pageId="4" pageNumber="13" phylum="Mollusca" rank="genus">Sinoennea</taxonomicName>
sp.) in the underground river sediments suggests a stronger influence of vertically circulating surface karst waters. The presence of the same species in the sediments of nearby Tham Pha Soung Cave indicates possible communication between the two caves through phreatic waters under Ban Na polje or during the rainy seasons. Surface
<taxonomicName id="F136B5B3C47CC1E6B0906AA3CB3D71AD" lsidName="" pageId="4" pageNumber="13" rank="tribe" tribe="Triculini">Triculini</taxonomicName>
species inhabiting the Nam
<normalizedToken id="B875E0B5E21712C3451B8418B4352180" originalValue="Dôn">Don</normalizedToken>
River are larger, with a stronger periorostracum and different shell morphology. It appears that they are not penetrating deeply into the dark cave system, probably because of a lack of their main algal food. The new species probably inhabits the so far unexplored submerged cave passages of the Tham Khon
<normalizedToken id="78E0EEF0FFB2703B354B265F954CBA07" originalValue="Dôn">Don</normalizedToken>
system.
</paragraph>
<caption id="F0A59A4AB1CA44FEC75B20A395093CE0" pageId="4" pageNumber="13">
<paragraph id="8FE3680DBA7AC1648F3BAF944EC3BF2D" pageId="4" pageNumber="13">
Figure 2. Photos of sites where subterranean gastropods were found. A Khammouane: main entrance of Tham Khon
<normalizedToken id="D51E096557264DE02DFEB5600CF327B2" originalValue="Dôn">Don</normalizedToken>
Cave with source of Nam
<normalizedToken id="99B7128BD4DD529EC9D7FE1F1DFC913E" originalValue="Dôn">Don</normalizedToken>
River (2 in Fig. 1) B Earthquake Dome in cave Tham Khon
<normalizedToken id="FEABE7CF13896CCB98E0E0CC62283868" originalValue="Dôn">Don</normalizedToken>
(1 in Fig. 1) C Mount Pha Kouankaohong with entrance of Tham Khon
<normalizedToken id="E33F0A8C4AEC49F460510BF809FCA61F" originalValue="Dôn">Don</normalizedToken>
cave at its foot D Bolikhamsay, travertine cascades below the LT of
<taxonomicName id="010307B8C20921A79CDE2A9651E1CCD4" class="Gastropoda" family="Pomatiopsidae" genus="Tricula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricula viengthongensis" order="Littorinimorpha" pageId="4" pageNumber="13" phylum="Mollusca" rank="species" species="viengthongensis">Tricula viengthongensis</taxonomicName>
sp. n. (6 in Fig. 1) E Khammouane, one of the main entrances of Tham Pha Soung Cave F Tham Pha Soung Cave, sampling at Frog Lake (4 in Fig. 1). (Photos: Ondrej Kameniar,
<normalizedToken id="DDEEB3C5B0F7A1B10EEF3713B7768636" originalValue="Mário">Mario</normalizedToken>
<normalizedToken id="71888BA3AE561C7ED71A352EB4A71BA4" originalValue="Olšavský">Olsavsky</normalizedToken>
and Jozef Grego).
</paragraph>
</caption>
<caption id="0F5509259B76AEDEB1D433BCC2B768AA" pageId="4" pageNumber="13">
<paragraph id="7E578C19F1419464EF9513B279F50336" pageId="4" pageNumber="13">
Figures 3-12. Representatives of the genus
<taxonomicName id="5A24EA070EDB26C7FC0F2B7E1961046E" class="Gastropoda" family="Pomatiopsidae" genus="Pseudoiglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoiglica" order="Littorinimorpha" pageId="4" pageNumber="13" phylum="Mollusca" rank="genus">Pseudoiglica</taxonomicName>
gen. n. 3-6
<taxonomicName id="214A95BD1DB22C45053149BADA9F383B" class="Gastropoda" family="Pomatiopsidae" genus="Pseudoiglica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoiglica pseudoiglica" order="Littorinimorpha" pageId="4" pageNumber="13" phylum="Mollusca" rank="species" species="pseudoiglica">Pseudoiglica pseudoiglica</taxonomicName>
sp. n. (3-4 holotype NHMUK 20180001 5-6 paratype 1 coll. Grego F0871) 7-8
<taxonomicName id="DDA3349ECE90DD4437397DF2CC3E7439" lsidName="P. olsavskyi" pageId="4" pageNumber="13" rank="species" species="olsavskyi">P. olsavskyi</taxonomicName>
sp. n. (holotype NHMUK 20180003) 9-10
<taxonomicName id="03C9D9D5675B6DBEF77C1E95E66D9AFE" lsidName="P. kameniari" pageId="4" pageNumber="13" rank="species" species="kameniari">P. kameniari</taxonomicName>
sp. n. (holotype NHMUK 20180002) 11-12
<taxonomicName id="23AB115FDFF399A4594BE7183A202530" lsidName="P. phonsavanica" pageId="4" pageNumber="13" rank="species" species="phonsavanica">P. phonsavanica</taxonomicName>
sp. n. (holotype NHMUK 20180004).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>