246 lines
21 KiB
XML
246 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.977.56314" ID-GBIF-Dataset="03422e51-f765-4dfb-af3d-128c9c96c002" ID-PMC="PMC7596022" ID-Pensoft-Pub="1313-2970-977-41" ID-Pensoft-UUID="80EDEC82017B5A0AA6E77D7CD518E347" ID-PubMed="33177942" ID-ZooBank="D2CB068E27924007B8AFD8B65B0B7A6C" ModsDocID="1313-2970-977-41" checkinTime="1603412801257" checkinUser="pensoft" docAuthor="Edmardash, Yusuf A., El-Ghiet, Usama M. Abu, Soliman, Ahmed M., Al-Fifi, Zarrag I. A. & Gadallah, Neveen S." docDate="2020" docId="765552B32E4F5D30BD3EF3589CDA3290" docLanguage="en" docName="ZooKeys 977: 41-74" docOrigin="ZooKeys 977" docSource="http://dx.doi.org/10.3897/zookeys.977.56314" docTitle="Hecabalodes anthaxiae Wilkinson 1929" docType="treatment" docVersion="4" id="80EDEC82017B5A0AA6E77D7CD518E347" lastPageNumber="41" masterDocId="80EDEC82017B5A0AA6E77D7CD518E347" masterDocTitle="First contribution to the doryctine fauna (Hymenoptera, Braconidae, Doryctinae) of Farasan Archipelago, Saudi Arabia, with new records and the description of a new species" masterLastPageNumber="74" masterPageNumber="41" pageNumber="41" updateTime="1668169635970" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>First contribution to the doryctine fauna (Hymenoptera, Braconidae, Doryctinae) of Farasan Archipelago, Saudi Arabia, with new records and the description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Edmardash, Yusuf A.</mods:namePart>
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<mods:affiliation>Entomology Department, Faculty of Science, Cairo University, Giza, Egypt</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>El-Ghiet, Usama M. Abu</mods:namePart>
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<mods:affiliation>Biology Department, Faculty of Science, Jazan University, Saudi Arabia & Plant Protection Department, Desert Research Center, Cairo, Egypt</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0288-4199</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soliman, Ahmed M.</mods:namePart>
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<mods:affiliation>Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia & Zoology Department, Faculty of Science (Boys), Al-Azhar University, Nasr City, Cairo, Egypt</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5284-713X</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Al-Fifi, Zarrag I. A.</mods:namePart>
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<mods:affiliation>Biology Department, Faculty of Science, Jazan University, Saudi Arabia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gadallah, Neveen S.</mods:namePart>
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<mods:affiliation>Entomology Department, Faculty of Science, Cairo University, Giza, Egypt</mods:affiliation>
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<mods:nameIdentifier type="email">n_gadallah@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>977</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>74</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.977.56314</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.977.56314</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-977-41</mods:identifier>
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<mods:identifier type="ZooBank">D2CB068E27924007B8AFD8B65B0B7A6C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">80EDEC82017B5A0AA6E77D7CD518E347</mods:identifier>
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<treatment ID-GBIF-Taxon="168635187" LSID="urn:lsid:plazi:treatment:765552B32E4F5D30BD3EF3589CDA3290" httpUri="http://treatment.plazi.org/id/765552B32E4F5D30BD3EF3589CDA3290" lastPageNumber="41" pageId="0" pageNumber="41">
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<subSubSection pageId="0" pageNumber="41" type="nomenclature">
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName LSID="765552B3-2E4F-5D30-BD3E-F3589CDA3290" authority="Wilkinson, 1929" authorityName="Wilkinson" authorityYear="1929" class="Insecta" family="Braconidae" genus="Hecabalodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hecabalodes anthaxiae" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="anthaxiae">Hecabalodes anthaxiae Wilkinson, 1929</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hecabalodes anthaxiae Wilkinson, ♀: A dorsal habitus B lateral habitus C antenna (broken at tip) D head, dorsal view E head, frontal view." figureDoi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41">Figures 6A-E</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">, 7A-E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="reference_group">
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authorityName="Wilkinson" authorityYear="1929" class="Insecta" family="Braconidae" genus="Hecabalodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hecabalodes anthaxiae" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="anthaxiae">Hecabalodes anthaxiae</taxonomicName>
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Wilkinson, 1929: 106, ♀♂.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="description">
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<paragraph pageId="0" pageNumber="41">Re-description of female.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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Body length: 4.2 mm; ovipositor length: 2.35 mm; fore wing length: 2.5 mm. (we re-describe this species in full because of the short original description of
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<bibRefCitation DOI="https://doi.org/10.1017/S000748530002099X" author="Wilkinson, DS" journalOrPublisher="Bulletin of Entomological Research" pageId="0" pageNumber="41" pagination="103 - 114" refId="B57" refString="Wilkinson, DS, 1929. New parasitic Hymenoptera and notes on other species. Bulletin of Entomological Research 20 (1): 103 - 114, DOI: https://doi.org/10.1017/S000748530002099X" title="New parasitic Hymenoptera and notes on other species." url="https://doi.org/10.1017/S000748530002099X" volume="20" year="1929">Wilkinson (1929)</bibRefCitation>
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): Dark brown, except for the yellowish hue on lateral sides of T1 and T2 as well as apex of T2 (Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hecabalodes anthaxiae Wilkinson, ♀: A dorsal habitus B lateral habitus C antenna (broken at tip) D head, dorsal view E head, frontal view." figureDoi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41">6A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7E</figureCitation>
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); antenna orange, scape slightly darker (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hecabalodes anthaxiae Wilkinson, ♀: A dorsal habitus B lateral habitus C antenna (broken at tip) D head, dorsal view E head, frontal view." figureDoi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41">6C</figureCitation>
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); legs dark brown (except for the yellowish base of fore tibia, and all tarsi), telotarsi darker. Fore wing subhyaline, with distinct infuscation along marginal cell (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7D</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="41">
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<emphasis bold="true" pageId="0" pageNumber="41">Figure 6.</emphasis>
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<taxonomicName authorityName="Wilkinson" authorityYear="1929" class="Insecta" family="Braconidae" genus="Hecabalodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hecabalodes anthaxiae" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="anthaxiae">
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<emphasis italics="true" pageId="0" pageNumber="41">Hecabalodes anthaxiae</emphasis>
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</taxonomicName>
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Wilkinson, ♀:
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<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
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dorsal habitus
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<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
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lateral habitus
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<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
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antenna (broken at tip)
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<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
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head, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="41">E</emphasis>
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head, frontal view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="41">
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<emphasis italics="true" pageId="0" pageNumber="41">Head</emphasis>
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(Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hecabalodes anthaxiae Wilkinson, ♀: A dorsal habitus B lateral habitus C antenna (broken at tip) D head, dorsal view E head, frontal view." figureDoi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41">6C-E</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7A</figureCitation>
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): 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as wide as its median length, slightly wider than mesoscutum; coarsely rugose; head behind eye broadly rounded; temple 0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as eye height in dorsal view; POL 1.6
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<normalizedToken originalValue="×">x</normalizedToken>
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OD, 0.9
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<normalizedToken originalValue="×">x</normalizedToken>
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OOL; eye with few scattered short setae; malar space 0.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as eye height, 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as basal width of mandible, malar suture absent; face smooth laterally just behind eyes; face 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as wide as eye width, 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long its length combined with clypeus; hypoclypeal depression more or less quadrate, ca. as wide as its distance from eye; occipital carina complete dorsally, not meeting hypostomal carina ventrally; antenna broken; scape twice as long as its maximum width; F1 6.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as its apical width, 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as F2; ocellar triangle with base longer than lateral sides.
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<emphasis italics="true" pageId="0" pageNumber="41">Mesosoma</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7A-C</figureCitation>
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): 2.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as its height; pronotum with two sharp transverse carinae dorsally; mesoscutum gently rounded above or at the same level of pronotum when seen from lateral view, flattened on disc, densely rugose, finely alutaceous laterally; notauli indistinct; mesoscutellum slightly convex, truncate at apex, finely sculptured, with a number of thick carinae laterally; mtn 0.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as mesoscutum, with a short longitudinal median carina and 2-3 oblique submedian carinae on its depressed anterior part, convex postero-medially; propodeum finely and sparsely granulate, with two short postero-medial, parallel carinae, 0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as propodeal length, median longitudinal carina of propodeum absent; mesopleuron finely punctate, with irregular spaces in between, shiny; precoxal sulcus shallow, irregular, running ventrally along almost the entire length of mesopleuron.
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<emphasis italics="true" pageId="0" pageNumber="41">Fore wing</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7D</figureCitation>
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): 3.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as its maximum width; pterostigma 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as maximum width; vein M + CU1 slightly curved; 1-SR+M nearly straight; vein r-m absent; vein r arising at basal third of pterostigma; 2-SR 1.75
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as r, slightly longer than m-cu, 0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as 1-SR+M; 1CU1 0.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as 2CU1.
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<emphasis italics="true" pageId="0" pageNumber="41">Hind wing</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7D</figureCitation>
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): With fringe of long, fine setae along apical and anal margins; vein 1-M 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as 1-rm. Legs. Hind coxa (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7C</figureCitation>
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) 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, without distinct basoventral tubercle, finely punctate especially ventrally, with some fine whitish setae distally and laterally; hind femur 2.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; hind tarsus 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as hind tibia; hind basitarsus slightly shorter than rest of tarsomeres combined; second tarsomere 0.55
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as hind basitarsus, 2.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as telotarsus (excluding arolium); outer edge of hind tibia with long, fine whitish setae.
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<emphasis italics="true" pageId="0" pageNumber="41">Metasoma</emphasis>
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(Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hecabalodes anthaxiae Wilkinson, ♀: A dorsal habitus B lateral habitus C antenna (broken at tip) D head, dorsal view E head, frontal view." figureDoi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41">6A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Hecabalodes anthaxiae Wilkinson, ♀: A head and mesosoma, lateral view B mesosoma and metasomal T 1 C mesosoma, ventral view D fore and hind wings E metasoma, dorsal view." figureDoi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41">7E</figureCitation>
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): 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as head and mesosoma combined; T1 and basal half of T2 with distinct interrupted longitudinal striae, somewhat dotted in between; T1 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as its apical width; T2 0.9
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as its apical width, 2.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as T3; posterior half of T2 finely reticulate, T3-5 (except posterior margin of T5 smooth and shiny), finely reticulate; T6 entirely smooth and shiny. Ovipositor sheath about as long as or slightly longer than metasoma (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hecabalodes anthaxiae Wilkinson, ♀: A dorsal habitus B lateral habitus C antenna (broken at tip) D head, dorsal view E head, frontal view." figureDoi="10.3897/zookeys.977.56314.figure6" httpUri="https://binary.pensoft.net/fig/466497" pageId="0" pageNumber="41">6A, B</figureCitation>
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), and the fore wing as well.
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</paragraph>
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<caption doi="10.3897/zookeys.977.56314.figure7" httpUri="https://binary.pensoft.net/fig/466498" pageId="0" pageNumber="41" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="41">
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<emphasis bold="true" pageId="0" pageNumber="41">Figure 7.</emphasis>
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<taxonomicName authorityName="Wilkinson" authorityYear="1929" class="Insecta" family="Braconidae" genus="Hecabalodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hecabalodes anthaxiae" order="Hymenoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="anthaxiae">
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<emphasis italics="true" pageId="0" pageNumber="41">Hecabalodes anthaxiae</emphasis>
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</taxonomicName>
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Wilkinson, ♀:
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<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
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head and mesosoma, lateral view
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<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
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mesosoma and metasomal T1
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<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
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mesosoma, ventral view
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<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
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fore and hind wings
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<emphasis bold="true" pageId="0" pageNumber="41">E</emphasis>
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metasoma, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="41">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="41">
|
||
Kingdom of Saudi Arabia, 1♀, Jazan, Farasan Islands, Al-Sajid;
|
||
<geoCoordinate degrees="16" direction="north" minutes="51" orientation="latitude" precision="1" seconds="25.46" value="16.857073">16°51'25.46"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="41" direction="east" minutes="55" orientation="longitude" precision="1" seconds="58.78" value="41.932995">41°55'58.78"E</geoCoordinate>
|
||
; 10 Nov.2017; Abu El-Ghiet & El-Sheikh leg.; LT [KSMA].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="41" type="distribution">
|
||
<paragraph pageId="0" pageNumber="41">General distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="41">
|
||
Sudan (
|
||
<bibRefCitation DOI="https://doi.org/10.1017/S000748530002099X" author="Wilkinson, DS" journalOrPublisher="Bulletin of Entomological Research" pageId="0" pageNumber="41" pagination="103 - 114" refId="B57" refString="Wilkinson, DS, 1929. New parasitic Hymenoptera and notes on other species. Bulletin of Entomological Research 20 (1): 103 - 114, DOI: https://doi.org/10.1017/S000748530002099X" title="New parasitic Hymenoptera and notes on other species." url="https://doi.org/10.1017/S000748530002099X" volume="20" year="1929">Wilkinson, 1929</bibRefCitation>
|
||
), Saudi Arabia (Farasan Islands) (new record).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="41" type="remarks">
|
||
<paragraph pageId="0" pageNumber="41">Remark.</paragraph>
|
||
<paragraph pageId="0" pageNumber="41">This species has not been collected during the 90 years or more since Wilkinson described the holotype from Sudan in 1929.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |